Micronesica 35-36:456-513. 2003 The crustaceans and pycnogonids of the Mariana Islands GUSTAV PAULAY1 Marine Laboratory University of Guam Mangilao, Guam 96923 USA ROY KROPP Battelle Marine Sciences Laboratory 1529 West Sequim Bay Road Sequim WA 98382-9099 USA PETER K. L. NG Raffles Museum of Biodiversity Research Department of Biological Sciences National University of Singapore Kent Ridge, Singapore 119260 Republic of Singapore LUCIUS G. ELDREDGE Pacific Science Association Bishop Museum 1525 Bernice St. Honolulu HI 96817 USA Abstract—The crustacean and pycnogonid fauna of the Mariana Islands is reviewed, and 839 crustacean and 15 pycnogonid species are documented from the archipelago based on literature records and new collections, including 272 new records. Voucher specimens are listed for 615 and photographic records (see http://www.flmnh.ufl.edu/reefs) for 358 species. The bulk of the fauna is marine, including 12 terrestrial and 12 freshwater decapods with marine larvae. Three copepods and 6 cladocerans comprise the known freshwater fauna, and 25 peracarids are currently documented on land. Coverage reflects a taxonomically uneven effort, and is strongly biased toward macrocrustaceans, with decapods accounting for 84% and crabs for 50% of the recorded marine crustacean diversity. Introduction The Mariana Islands form a chain of 15 major volcanic and limestone islands bordering the western margin of the Pacific plate between 13 and 21oN latitude 1 Current address: Florida Museum of Natural History, Gainesville FL 32611-7800 USA, email: paulay@flmnh.ufl.edu Paulay et al.: Crustaceans and pycnogonids of the Marianas 457 (see Paulay 2003 for a more detailed description of the area). The archipelago is a classic island arc, formed by volcanic eruptions initiated by the subduction of the Pacific plate under the Philippine plate, on which the islands lie. The island group is composed of two geologically distinct chains, the volcanically-inactive and older islands of the Southern Mariana back-arc, extending from Guam to Farallon de Medinilla, and the volcanically-active, younger islands of the Northern Mariana fore-arc, extending from Anatahan to Uracas (Fig. 1). This geological separation should not be confused with the political division of the archipelago into the U.S. Territory of Guam, and the Commonwealth of the Northern Mariana Islands (CNMI) comprising all islands north of Guam. Thus, Rota, Tinian, Aguijan, Saipan and Farallon de Medinilla are part of the Southern Mariana back-arc but are under the jurisdiction of the CNMI. The Southern Marianas were initiated ~43 Ma when the Pacific plate abruptly shifted spreading from NW to WNW. The archipelago has had a complex geological history since. The southern islands are composites of submarine and subaerial volcanics and marine sedimentary formations raised by uplift. Volcanic shorelines are uncommon while karstic shores of mostly Miocene to Pleistocene limestones dominate. Limestone cliffs line large tracks of shore, and reefs are generally narrow. Large sections of especially windward shores lack reef protection, with steep fore reefs abuting eroded cliffs, often fronted by supratidal limestone benches. Other areas have narrow, largely intertidal reef flats or shallow, moderately wide (100-1000 m) fringing reefs. Deep moat and lagoonal habitats are uncommon, the only significant ones being Apra Harbor and Cocos lagoon on Guam and Tanapag-Garapan lagoon on Saipan. Only Apra Harbor has substantial lagoonal habitats >10 m deep. Mangals are consequently poorly developed, and have undergone considerable reduction since the arrival of humans (Amesbury 1999). The main remaining mangals are in inner Apra Harbor and along the SW shore of Guam. Subtidal habitats are dominated by oligotrophic reefs with interspersed sand pockets, with finer mobile sediments largely confined to small estuaries, Apra Harbor, and deep water. The fauna of soft bottoms remains poorly studied. The karstic limestones bordering and underlying reefs are riddled by caves and caverns both above and below water. Although marine caverns and crevices are fairly common, they are generally of limited horizontal extent. Large blocks fallen from cliff faces provide additional wall and crevice habitats. Rubble fields are also common, and have been the focus of considerable sampling that revealed a rich crustacean community, including many unusual, troglobitic forms living deep in rubble (e.g. Ng 2002b, Ng & Ng 2003, Ng & Takeda 2003). The Northern Marianas are presently uninhabited, active, young volcanoes. They essentially lack reefs, having rocky shore and subtidal habitats with coral communities and incipient reefs growing on igneous outcrops and rubble. Consequently there are no lagoonal habitats except for the protected crater of Maug. Although poorly known, the biota of the Northern Marianas appears to be largely a less diverse version of that of the Southern Marianas, although some 458 Micronesica 35-36, 2003 species are clearly limited to the northern islands, likely specialized to uniquely northern habitats. Figure 1. Map of the Mariana Islands. Paulay et al.: Crustaceans and pycnogonids of the Marianas 459 CARCINOLOGICAL HISTORY The first recorded collection of crustaceans from the Marianas was by Quoy & Gaimard (1824), who described the land crab Thelphusa rotunda (now Discoplax rotundum) and listed 5 additional species from Guam. In February 1900 the U.S.F.S. “Albatross” Expedition called on Guam, but only a single crab species (Ocypode ceratophthalma) was recorded from their visit (Rathbun 1907). William Edwin Safford, Assistant Botanist for the Department of Agriculture and Lieutenant U.S. Navy, was stationed on Guam between August 1899 to August 1900. He collected a number of crustaceans and wrote short notes on the crabs (Safford 1905). Alvin Seale of the U.S. War Department, spent several months on Guam in 1900, reported on the birds and fishes (Seale 1901), and deposited more than 30 lots of crustaceans in the Bishop Museum. Hans George Hornbostel, Lieutenant U.S. Naval Hospital, archaeologist, and collector for the Bishop Museum was on Guam between April 1922 and June 1928. He wrote primarily on archaeology, but also collected nearly 100 lots of crustaceans for the Bishop Museum. Naturalist Edwin H. Bryan Jr. collected extensively on Guam in 1936, his collections were also deposited at the Bishop Museum. In the wake of World War II Micronesia received considerable attention from the U.S. During 1951-1954, the U.S. Army Corps of Engineers and U.S. Geological Survey conducted field work on Guam for the “Military Geology of Guam”, headed by Kenneth O. Emery. Substantial collections were made by several visitors at this time and deposited at the USNM; these served as the basis of subsequent studies on the marine biota (e.g. Stephenson & Rees 1967). A similar effort headed by Preston Cloud Jr. on Saipan in 1949 included an intensive survey of the marine biota. Cloud (1959) recorded 125 crustaceans, most identified to species by the authorities of the time, several of whom published papers based largely on this collection (Banner 1956, Holthuis 1953, 1981). Cloud’s specimens, deposited at the USNM, have been the source of numerous Saipan records in subsequent taxonomic revisions as well (e.g. Chace & Bruce 1993, Clark & Galil 1993, Castro 2003). Carcinological studies in the Marianas gained considerable momentum with the establishment of the University of Guam Marine Laboratory (UOGML) in 1970. Lu Eldredge was involved in a number of surveys focused both on shallow and deep water crustaceans, trained several M.S. students in carcinology, distributed material to specialists, and hosted visiting carcinologists. Deep-water marine resource surveys included an assessment of deep-water shrimp resources around Guam, funded by Sea Grant between 1973-1976 (Wilder 1977), and the Resource Assessment Investigation of the Mariana Archipelago (RAIOMA) of the National Marine Fisheries Service using the NOAA ship TownsendCromwell between May 1982 and June 1984 (Polovina et al. 1985). These efforts have led to a number of publications on the systematics and fisheries potential of deep-water crustaceans (Eldredge 1976, 1980, Wilder 1977, Moffitt 1983; King 1987, Williams & Moffitt 1991, Williams & Eldredge 1994). A visit by the submersible Alvin to the hydrothermal vents of the Mariana back arc resulted in 460 Micronesica 35-36, 2003 several papers on the crustacean biota of this isolated vent system (Hessler & Martin 1989, Humes 1990, Hessler & Lonsdale 1991, Child 1989), while a survey of the much shallower, hydrothermally-active volcano at Esmerelda Bank led to the record of two unusual decapods (Türkay & Sakai 1995). Several graduate students have pursued carcinological theses at the University of Guam (Tseng 1972, Wortman 1976, Dickinson 1977, Wooster 1979, Kropp 1981, Donaldson 1981, Grimm 1983, Camacho 1997, Leberer 1997). Wooster’s study culminated in a paper on the Calcinus fauna of Guam (Wooster 1982), while Leberer went on to review the taxonomy of the island’s atyids (Leberer & Cai 2003). Roy Kropp and Jane Dominguez undertook an intensive collecting survey of the shallow water crustaceans of Guam; their large collection now at the USNM has been the basis of numerous publications (e.g. papers by Kropp and co-authors, Child 1991, Ahyong & Erdmann 2003, Tan & Ng 2003b). During the 1990s the Gustav Paulay’s laboratory has been active in marine biodiversity surveys on Guam. Material was collected during several organized surveys (see Paulay 2003), as well as incidentally by students, faculty, staff and friends of the University of Guam Marine Laboratory. Many specimens generated by these surveys have been sent to specialists for identification and study (e.g. McLay 2001, McLaughlin 2002a-b, Ng, 2002a-c). Harry Conley has contributed an especially large number of noteworthy specimens, collected from deep within rubble, a microhabitat whose exploration he pioneered. His collections have led to the discovery of numerous interesting new taxa (e.g. Kensley 2003, Ng et al. papers in this volume). The Chiba Museum undertook an expedition to the Northern Marianas in 1992 and published a comprehensive checklist of their findings in a special volume, with several papers dealing with crustaceans (e.g. Asakura et al. 1994, Hayashi et al. 1994, Takeda et al. 1994, Asakura & Tachikawa 2000). Other visiting carcinologists also published on the fauna (e.g. Belk 1973, Odinetz 1984, Weissleader et al. 1989, Foster 1990, Asami & Yamaguchi 2001, Castro 2003). A visit by the third author and Chia-Hsiang Wang in 2000 led to a review of the crabs in the University of Guam’s Invertebrate Collection, an effort that has already led to several papers that appear in this volume as well as many of the identifications listed in our checklist. A subsequent collecting visit by their team in 2001 together with ongoing studies of previously collected material (especially of the Grapsidae, Xanthidae and Parthenopidae) will undoubtedly lead to numerous studies on Guam’s crustaceans in the future. Methods This paper is based on a compilation of records from the literature and identifications made on new collections. Both efforts are ongoing and certainly incomplete. Although we strove to check all papers dealing with the Micronesian fauna as well as major taxonomic revisions, additional records of the biota Paulay et al.: Crustaceans and pycnogonids of the Marianas 461 undoubtedly appear in papers we did not consult. We did not include records listed in the voluminous gray literature of environmental surveys published on the area, as such records are usually difficult or impossible to verify. We have updated the nomenclature of literature records as much as possible; changes are given as notes in the Appendix. Numerous species listed here have not been previously recorded and comprise new additions to the Marianas’ fauna. These are marked by the name of the identifier in the Appendix and identified with a specimen voucher and/or photograph. New collections focused largely on stomatopods, decapods, and barnacles. Even among these groups efforts have been uneven, with larger species, crabs, hermit crabs, axiids, and lobsters receiving the greatest attention. Much of this material is housed at the Florida Museum of Natural History, University of Florida (UF), U.S. National Museum of Natural History, Smithsonian Institution (USNM), Zoological Reference Collection, Raffles Museum, National University of Singapore (ZRC), and the University of Guam (UGI). Most newly collected species were photo documented. Photo vouchers cited in the Appendix are available online at http://www.flmnh.ufl.edu/reefs and in the Marine Biodiversity of Guam CD-ROM co-publication. In addition to the species listed here, numerous remain to be identified and the identification of many listed species improved. Work is ongoing on these and additions and modifications will be posted on the website as they are published. Major ongoing efforts include studies on barnacles (W. Newman, A. Ross), paguroids (P. McLaughlin), galatheoids (R. Kropp), and brachyurans (B. Galil, C. McLay, P. Ng et al.). For the purposes of this checklist we define marine crustaceans as species that have at least one phase of their life cycle in the sea. These include all terrestrial and freshwater decapods in the Marianas, as all are known or presumed to have marine larvae. Truly terrestrial and freshwater crustaceans are listed below the marine taxa, and include isopods, talitrid amphipods, copepods, and cladocerans. Results and Discussion We documented 839 crustacean and 15 pycnogonid species from the Mariana Islands based on literature records and new collections supported by voucher specimens and/or photographic records (Tables 1, 2, Appendix). 483 species are based at least in part on literature records, 272 are recorded for the first time in this paper, while an additional 99 are new records from our recent biodiversity surveys that have been further documented in other papers in this volume, and in papers published recently or in press elsewhere. 358 species are photodocumented based on living or fresh specimens. 663 species are recorded from Guam, while the remainder only from the CNMI. Most of the species currently known only from the CNMI likely also occur on Guam, but remain undocumented there because of limited collecting or different 462 Micronesica 35-36, 2003 taxonomic emphasis. For example the alpheids of Saipan were reviewed in a substantial paper by Banner (1956), while those of Guam remain virtually unstudied. Table 1. Diversity of marine arthropod taxa Taxon Number of spp. Copepoda Ostracoda Amphipoda Isopoda Stomatopoda Stenopodidea Penaeoidea Caridea Palinura Astacidea Thalassinidea Anomura Brachyura Ascothoracida Cirripedia Pycnogonida 4 45 17 3 36 1 4 139 11 2 16 94 405 1 27 15 TOTAL 820 The bulk (805 species) of the crustaceans are marine, including 12 species of land and hermit crabs with marine larvae and 12 species of anadromous decapods (Table 2). Three copepods and 6 cladocerans comprise the known freshwater fauna, and 25 species of peracarids are currently documented on land. Taxonomic coverage is highly uneven, with macrocrustaceans comprising the vast majority of the records (Table 1). Thus decapods comprise 84% (674 species) and crabs 50% (405 species) of the marine crustaceans recorded. Other crustacean taxa remain poorly documented with the exception of stomatopods (36 species) and barnacles (27 species). For example only 20 peracarids and 4 copepods are currently recorded in the marine fauna. Even among decapods, coverage varies substantially among families, reflecting taxonomically focused collecting and study. Thus brachyurans have received much greater attention overall than carideans. Cryptochirids and porcellanids were intensively studied by Kropp, and are especially well known (see papers by Kropp et al.). The most species rich families are the Xanthidae (114 species), Alpheidae (61 species), Portunidae (55 species), Diogenidae (40 species), Palaemonidae (35 species), and Cryptochiridae (28 species). Even groups that have been the focus of study harbor much undocumented diversity. Thus the known paguroid diversity of Guam has more than doubled (31 vs. 69 species) in the two decades following Wooster’s (1979, 1982) thesis on the group, even though hermit crabs did not receive much specialized attention during this period. Paulay et al.: Crustaceans and pycnogonids of the Marianas 463 It is difficult to estimate the actual diversity of marine arthropods in Guam or the Marianas, as even macrocrustaceans are but modestly known. Crabs are among the best studied overall, but the rapid rate of new encounters and backlog of unidentified material implies that perhaps not more than half of the crab diversity is currently documented. Decapod diversity as a whole will certainly much more than double with additional study and the diversity of microcrustaceans cannot be reliably estimated at present. Table 2. Arthropod diversity by habitat Habitat Number of spp. Shallow (<60m) marine Deep (>60m) marine Anadromous Terrestrial with marine larvae Freshwater Terrestrial 747 49 12 12 9 25 Acknowledgements We thank the students, faculty and staff at the UOG Marine Lab and naturalists from throughout the Marianas, especially Tim Allen, Scott Bauman, Ian Elliott, Bruce Henke, Lisa Kirkendale, Chris Meyer, Rob Myers, Raphael Ritson-Williams, Teina Rongo, John Starmer, and Linda Ward for bringing crustacean specimens, records, and photos to our attention. Special thanks to Harry Conley for his indefatigable collecting over the years. We thank Shane Ahyong, Kiyo Asami, Warren Blow, Yixiong Cai, Peter Castro, Tin-Yam Chan, Bill Dall, Mark Erdmann, Charles Fransen, Bella Galil, Yoshi Hisatsune, Brian Kensley, Hung-Chang Liu, Patsy McLaughlin, Colin McLay, Junji Okuno, John Starmer, Masatsune Takeda, and Chia-Hsiang Wang for help with identifications. 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Received 3 March 2002 Museum BMNH 1999.1529 (holotype) Taxon MARINE TAXA COPEPODA: DIRIVULTIDAE Chasmatopontius thescalus Humes, 1990 Stygiopontius pectinatus Humes, 1987 Stygiopontius stabilitus Humes, 1990 COPEPODA: ERGASILIDAE Paeonodes subviridis El-Rashidy & Boxshall, 2001 OSTRACODA: PODOCOPIDA Bairdoppilata cratericola Maddocks, 1969 Bishopina timorensis (Fyan, 1916) Bosasella cf. macroloba (Hu, 1981) Bythocypris aff. prolata Maddocks, 1969 Callistocythere arcuata Bonaduce et al., 1980 1 1 1 1 1 3 3 3 G G G G G G MTV MTV MTV Hab Island 3 3 3 3 3 Notes 1 Photo 140 1, 2 2 1, 2 Refs Museum: vouchers specimens in AHF: Allan Hancock Foundation, now at Natural History Museum of Los Angeles County; AM: Australian Museum, Sidney; BMNH: The Natural History Museum (formerly British Museum of Natural History); BPBM: Bernice P. Bishop Museum, Honolulu; CBM: Natural History Museum and Institute, Chiba; KC: Roy Kropp Collection; to be deposited in an institutional collection in the future; MNHN: Muséum national d'Histoire naturelle, Paris; NNM: Nationaal Natuurhistorisch Museum, Leiden; NSMT: National Science Museum, Tokyo; SIO: Scripps Institution of Oceanography, La Jolla; SMF: Senckenberg Museum, Frankfurt; UF: Florida Museum of Natural History, University of Florida, Gainesville; UGI: University of Guam Invertebrate Collections; USNM: US National Museum of Natural History, Washington DC; USNPC: US National Parasite Collection, National Museum of Natural History, Washington; YPM: Yale University Peabody Museum of Natural History, New Haven; ZRC: National University of Singapore, UF/ZRC; specimen deposition undecided but will be UF and/or ZRC. Vouchers are from Guam unless parenthetically indicated otherwise; see under “Island” below for key to island name abbreviations. Hab: Habitat: 1: marine, 0-60m, 2: marine, 60-200m, 3: marine, >200m, FW: freshwater, T: terrestrial Island: Ag: Agrihan, Al: Alamagan, An: Anatahan, Ar: Arakane Reef, As: Asuncion, CNMI: Commonwealth of the Northern Mariana Islands, island unspecified, E: Esmeralda Bank, G: Guam, Gu: Guguan, M: Maug, MS: Mariana Islands, island unspecified, MTV: Mariana Trough vents, P: Pagan, R: Rota, S: Saipan, Sa: Sarigan, T: Tinian, U: Uracas. Refs: References listed at end of Appendix Notes: listed at end of Appendix Appendix: Checklist of the crustaceans and pycnogonids of the Marianas Callistocythere cf. keijii (Hartmann, 1978) Caudites exmouthensis Hartmann, 1978 Cletocythereis rastromarginata (Brady, 1880) Cytherelloida sp. B Cytherelloida sp. E Hemicythere sp. B Hemicytherura sp. 1 Hermanites deltoidea (Brady, 1890) Hermanites transoceanica Teeter, 1975 Keijia foveata (Hartmann, 1978) Loxoconcha heronislandensis Hartmann, 1981 Loxoconcha huahineensis Hartmann, 1984 Loxoconcha sp. 1 Loxoconcha turnerae Watson, 1988 Loxoconchella anomala (Brady, 1880) Loxoconchella cf. malaysiana Keij, 1978 Morkhovenia inconspicua (Brady, 1880) Mydinobairdia schyroconcha (Maddocks, 1969) Neonesidea cf. arenigera Maddocks, 1969 Neonesidea lentifila Hartmann, 1984 Neonesidea schulzi (Hartmann, 1964) Ornatoleberis sp. 1 Paracytheridea cf. dissecta Hu, 1983 Paracytheridea tschoppi Bold, 1946 Paranesidea cf. algicola Maddocks, 1969 Paranesidea sp. A Paranesidea spongicola Maddocks, 1969 Ponticocytheres ichthyoderma (Brady, 1890) Propontocypris sp. A Propontocypris sp. B Pterobairdia maddocksae McKenzie & Keij, 1977 Taxon Museum 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 Refs Photo 3 2 1 Notes Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 G G G G G G G G G G G G G G G G G G G G G G G G G G G G G G G Hab Island Radimella sp. A Renaudcypris? natans Hartmann, 1984 Saipanetta cloudi (McKenzie, 1967) Semicytherura cf. cryptifera (Brady, 1880) Tanella cf. gracilis Kingma, 1948 Triebelina bradyi Triebel, 1948 Triebelina sertata Triebel, 1948 Triebelina sp. 1 Xestoleberis spp. AMPHIPODA: AMPHILOCHIDAE Gitanopsis sp. 1 AMPHIPODA: AMPITOIDAE Ampithoe ramondi Audouin, 1826 Ampithoe sp. 1 Ampithoe sp. 2 Ampithoe sp. 3 Grubia filosa (Savigny, 1826) Paradusa sp. 1 AMPHIPODA: COLOPHIIDAE Gammaropsis sp. 1 AMPHIPODA: GAMMARIDAE Ceradocus rubromaculatus (Stimpson, 1855) Erasmops sp. 1 Maera insignis (Chevreux, 1901) Maera sp. 1 AMPHIPODA: HYALIDAE Hyale sp. 1 Hyale sp. 2 Hyale sp. 3 AMPHIPODA: ISCHYROCERIDAE Ambicholestes squamiferus (Just, 1984) Taxon UGI (paratype) USNM 113437 (holotype, S) Museum 1 1 1 1 1 1 1 5 4 5 4 4 4 4 1 1 4 6 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 G M M M S M S M M S M Gu M S M M G G S G G G G G G Hab Island 5 4 4 4 5 4 4 68 Notes 1 Photo 4 3 3 132 3 3 3 3 3 3 Refs Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) STOMATOPODA: LYSIOSQUILLIDAE Lysiosquillina maculata (Fabricius, 1793) Lysiosquillina sulcata (Manning, 1978) Gonodactylellus espinosus (Borradaile 1898) Gonodactylellus micronesicus (Manning, 1971) Gonodactylus childi Manning, 1971 Gonodactylus chiragra (Fabricius, 1781) Gonodactylus platysoma Wood-Mason, 1895 Gonodactylus smithii Pocock, 1893 Hoplosquilla said Erdmann & Manning, 1998 STOMATOPODA: INDOSQUILLIDAE Indosquilla manihinei Ingle & Merrett, 1971 AMPHIPODA: LEUCOTHOIDAE Leucothoe sp. 1 ISOPODA: CYMOTHOIDAE Ceratothoa angulata (Richardson, 1910) Cymothoa bychowskyi Avdeev, 1979 Elthusa raynaudii (Milne-Edwards, 1840) STOMATOPODA: BATHYSQUILLIDAE Bathysquilla microps (Manning, 1961) STOMATOPODA: CORONIDIDAE Mortensenenus paulay Ahyong & Erdmann, 2003 Neocoronida martensi Manning, 1978 STOMATOPODA: EURYSQUILLIDAE Manningia zehntneri Manning, 1974 STOMATOPODA: GONODACTYLIDAE Gonodactylaceus falcatus (Forsskål, 1775) Gonodactylaceus ternatensis (de Man, 1902) Gonodactylellus erdmanni Ahyong, 2001 Taxon 9 9 9 5, 9, 10 9 9 AM P (holotype, M) AM P USNM 307137 UF 265, USNM 304400 USNM 304416 USNM 304403, USNM 304653 (R) USNM 304445 USNM 304408 USNM 304444 UF 239, USNM 304489 UF 233, USNM 304455 UF 235, USNM 304453 USNM 306080 UF 232, USNM 304425 USNM 233477, USNM 228417 (P) 8,9 USNM 228146 9 12 8, 9 9 9 9 5, 9, 10 9, 11 9 9 7 7 7 4 Refs USNPC 088995 USNPC 088996 USNPC 088997 Museum Notes RFM 226 RFM GP359-21 GP464-10 6 GP404-30 GP694-14 5 Photo Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 3 1 1 1 1 1 1 1 1 1 1 1 1 1 3 1 1 1 1 G G, S G, P G G G G, S G G G G, S G G, R G M M G G G G M Hab Island STENOPODIDEA: STENOPODIDAE Stenopus hispidus (Olivier, 1811) Raoulserenea oxyrhyncha (Borradaile, 1898) STOMATOPODA: SQUILLIDAE Leptosquilla schmeltzii (A. Milne-Edwards, 1873) Oratosquilla fabricii (Holthuis, 1941) Parvisquilla dominguez Ahyong & Erdmann, 2003 STOMATOPODA: TAKUIDAE Mesacturus dicrurus Kropp & Dominguez, 1990 STOMATOPODA: NANNOSQUILLIDAE Pullosquilla litoralis (Michel & Manning, 1971) Pullosquilla malayensis (Manning, 1968) Pullosquilla thomassini Manning, 1978 STOMATOPODA: ODONTODACTYLIDAE Odontodactylus scyllarus (Linnaeus, 1758) STOMATOPODA: PROTOSQUILLIDAE Chorisquilla gyrosa (Odhner, 1923) Chorisquilla kroppi Ahyong & Erdmann, 2003 Echinosquilla guerini (White, 1861) Haptosquilla glyptocercus (Wood-Mason, 1875) Haptosquilla tanensis (Fukuda, 1911) STOMATOPODA: PSEUDOSQUILLIDAE Pseudosquilla ciliata (Fabricius, 1787) Pseudosquilla ornata Miers, 1880 Pseudosquillana richeri (Moosa, 1991) Raoulserenea hieroglyphica (Manning, 1972) Raoulserenea komaii (Moosa, 1991) Taxon 9 9 9 9 9 9 9 5, 10 9 9 9 USNM 304435 USNM 304448 USNM 304499 (holotype) USNM 304424 UF 237, USNM 120271 Neotype: USNM 304475 USNM 304428 9, 13, 14 USNM 244236 (holotype), USNM 244238 (paratype, S) GP396-30 GP650-32 RFM 206b RK GP467-33 RK RFM 98a Photo 5, 10, 15 GP516-23 9 9 9 USNM 304446 USNM 304447 USNM 304525 (holotype) USNM 171598, BPBM 2873 9 USNM 304426 USNM 304432 USNM 304427 UF 236, USNM 304430 9 9 9 Refs USNM 304502 USNM 304529 USNM 304500 Museum Notes Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 G, S, Gu, P, M G, S, M G G G G S G G G, As, M, Ag G G G G G G G G G G Hab Island Alpheus gracilipes Stimpson, 1860 Alpheus gracilis Heller, 1862 Alpheus haani Ortmann, 1890 Alpheus idiocheles Coutière, 1905 Alpheus ladronis Banner, 1956 Alpheus deuteropus Hilgendorf, 1878 Alpheus diadema Dana, 1852 Alpheus dolerus Banner, 1956 Alpheus eulimene de Man, 1911 Alpheus frontalis H. Milne Edwards, 1837 Alpheus crockeri (Armstrong, 1941) PENAEOIDEA: PENAEIDAE Metapenaeopsis commensalis Borradaile, 1898 Penaeus monodon Fabricius, 1798 Litopenaeus stylirostris (Stimpson, 1874) Litopenaeus vannamei (Boone, 1931) CARIDEA: ALPHEIDAE Alpheopsis diabolus Banner, 1956 Alpheopsis equalis Coutière, 1896 Alpheopsis tetrarthri Banner, 1956 Alpheus bidens? (Olivier, 1811) Alpheus bradypus Coutière, 1905 Alpheus brevipes Stimpson, 1860 Alpheus bucephalus Coutière, 1905 Alpheus cf. crinitus Dana, 1852 Alpheus chamorro Banner, 1956 Alpheus coetivensis Coutière, 1908 Alpheus collumianus Stimpson, 1860 Taxon USNM 255980 USNM (holotype, S) USNM NNM D-48070 (As), UGI 6629 (CNMI) NNM D-48068 (As), UGI 6628 (CNMI) UF 829, USNM 233401 NNM-D48076 (Ag) USNM (holotype, S) USNM UF 825, NNM D-48074 (Al) USNM (holotype, S) 5, 18 18 5 5, 18, 19 18 5, 18 5, 18 18 5, 18 15, 18 18 18 5, 18 18 15 18 5, 18 5, 18 15 18 18 5, 18 USNM (holotype, S) NNM D-48087 (As) USNM (holotype, S) USNM USNM D 16 16 17 Refs UF 891 Museum GP454-17 GP758-11 GP264-14 Photo 13 12 11 9 10 8 7 7, 69 7 Notes Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 G, S Ag, S S S G, Al, S, P, M S S S G, S S As, S S As, S S Gu S S S Ag S S As, S G G G G Hab Island Alpheus sp. 1 (n. sp.) Alpheus sp. 2 (n. sp.) Alpheus sp. 3 (n. sp.) Alpheus sp. 4 (n. sp.) Arete iphianassa de Man, 1910 Athanas areteformis Coutière, 1903 Athanas djiboutiensis Coutière, 1897 Alpheus paracrinitus Miers, 1881 Alpheus parvirostris Dana, 1852 Alpheus percyi Coutière, 1908 Alpheus perplexus Banner, 1956 Alpheus rapax Fabricius, 1798 Alpheus stanleyi Coutière, 1908 Alpheus staphylianus Coutière, 1908 Alpheus sulcatus Kingsley, 1878 Alpheus superciliaris Coutière, 1905 Alpheus lutini Coutière, 1905 Alpheus malleodigitus (Bate, 1888) Alpheus microstylus (Bate, 1888) Alpheus oahuensis (Banner, 1953) Alpheus obesomanus Dana, 1852 Alpheus ovaliceps Coutière, 1905 Alpheus pachychirus Stimpson, 1860 Alpheus pacificus Dana, 1852 Alpheus leptochirus Coutière, 1905 Alpheus lottini Guérin, 1829 Taxon Photo Notes 5, 18 5, 18, 19 5, 18 18 15, 19 18 5, 18 15, 18 20 18 19 17 16 15 18 5, 15, 18 GP784-26 14 Refs 5, 18 5, 18 15 USNM (holotype, S) 18, 20 BPBM S6850 18 18 UGI 6622 (CNMI) F NNM D-48075 (Al) F UF 962 (Ag), UF 961 (As), UF F 960 (M), NNM D-48078 (As) 18 18 18 18 USNM 5, 18 15, 18 USNM 5, 18 USNM USNM USNM USNM 255979 UF 939 (Al), UF 985 (As), UF 828, UGI 6627 (CNMI), NNM-D48064 (Ag), BPBM USNM UF 826, USNM 223423 USNM Museum Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 S S S S S M, S S S G, S S S G, M S S S, Gu, Al, M S S As G, S G S CNMI Al As, M, Ag S G, Ag, Al, An, As, S Hab Island Photo 5, 10 15 GP415-18 1 1 1 1 1 5, 10 P F UF 456, BPBM UF 808 NNM D-48092 GP363-27 3 1 26 2, 22 5, 10, 24 USNM 94774 (holotype, S) 1 1 1 1 1 1 1 1 1 1 1 1 1 S Gu G, S G G MTV S G G G G G G G, As, M, Ag G, S Ag, S S M, S M, U Ag Gu, M G, S G, Ag, M, S S As, S, M G, As Hab Island FW FW FW FW FW FW 24 25 22 23 21 Notes 21 21 21 21 21 21 18 5, 15, 18 18 5, 18 5, 15, 18 15 15 18 15, 18 18 5, 15, 18 F GP830-3 F Refs UF/ZRC UF 812 UF/ZRC UF/ZRC UF/ZRC UF/ZRC USNM (holotype, S), BPBM BPBM, NNM D-48084 (Ag) USNM NNM D-48082 (As) NNM D-48085 (As) USNM NNM D-48086 (M) UF 968 (M), BPBM, NNM D-48088 (As) USNM (holotype, S), BPBM Athanas dorsalis (Stimpson, 1861) Athanas dubius Banner, 1956 Automate dolicognatha de Man, 1888 Automate johnsoni Chace, 1955 Metalpheus paragracilis (Coutière, 1897) Metalpheus rostratipes (Pocock, 1890) Racilius compressus Paulson, 1875 Synalpheus tumidomanus (Paulson, 1875) Synalpheus anceps Banner, 1956 Synalpheus charon (Heller, 1861) Synalpheus exilipes Coutière, 1905 Synalpheus paraneomeris Coutière, 1905 Synalpheus stimpsonii (de Man, 1888) CARIDEA: ATYIDAE Antecaridina lauensis Edmondson, 1935 Atyoida pilipes (Newport, 1847) Caridina brachydactyla (de Man, 1908) Caridina mertoni Roux, 1911 Caridina typus H. Milne Edwards, 1837 Halocaridinides sp. 1 (n. sp.) CARIDEA: BRESILIIDAE Chorocaris vandoverae Martin & Hessler, 1990 Discias musicus Holthuis, 1981 CARIDEA: GNATHOPHYLLIDAE Gnathophyllum americanum Guérin-Méneville, 1855 Gnathophyllum precipuum Titgen, 1989 Pycnocaris chagoae Bruce, 1972 CARIDEA: HIPPOLYTIDAE Thinora maldivensis (Borradaile, 1915) Hippolyte edmondsoni Hayashi, 1982 Museum Taxon Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) UF 819 Hippolyte sp. 1 Lysmata amboinensis (de Man, 1888) Lysmata sp. 1 Parhippolyte misticia (Clark, 1989) Saron marmoratus (Olivier, 1811) Saron neglectus de Man, 1902 Saron sp. 1 Thor amboinensis (de Man, 1888) Thor paschalis (Heller, 1862) CARIDEA: HYMENOCERIDAE Hymenocera picta Dana, 1852 CARIDEA: PALAEMONIDAE Brachycarpus biunguiculatus (Lucas, 1846) Conchodytes meleagrinae Peters, 1852 Coralliocaris graminea (Dana, 1852) Coralliocaris superba (Dana, 1852) Dactylonia ascidicola (Borradaile, 1898) Exopontonia malleatrix Bruce, 1988 Harpiliopsis beaupresii (Audouin, 1826) Harpiliopsis depressa (Stimpson, 1860) Harpiliopsis spinigera (Ortmann, 1890) Ischnopontonia lophos (Barnard, 1962) Jocaste japonica (Ortmann, 1890) Jocaste lucina (Nobili, 1901) Leander plumosus Bruce, 1993 Leander tenuicornis (Say, 1818) Macrobrachium equidens (Dana, 1852) Macrobrachium lar (Fabricius, 1798) Macrobrachium rosenbergii (de Man, 1879) Macrobrachium sp. 1 Onycocaris quadratophthalma (Balss, 1921) F F F 5, 10 134 F 15 15 F 15 15 5, 10 P F N 27, 136 16 26 15 UF 935, NNM D-48096 (As) NNM D-48095 UF 842 UF 818, NNM D-48093 (S) ZRC BPBM-S 117 NNM D-48079 (Ag) NNM D-48066 NNM D-48065 P F P F P 5, 10 5, 10 P P 5, 10, 25 Refs UF 822 USNM? UF 821 UF 884 UF 807 ?UF 457 Museum Taxon TA 1999 GP410-11 GP821-37 GP653-31 RK RFM-186a RFM GP778-36 GP826-1 GP516-28 Photo 7 27 Notes Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 FW FW FW 1 1 1 1 1 1 1 1 1 1 1 G, As G G S G G M M, Gu Ag Ag Ag S G G, S G G G G M G G G G G G, S S G G S Hab Island Palaemon debilis Dana, 1852 Palaemonella rotumana (Borradaile, 1898) Palaemonella spinulata Yokoya, 1936 Periclimenaeus quadridentatus (Rathbun, 1906) Periclimenaeus tridentatus (Miers, 1884) Periclimenella spinifera (de Man, 1902) Periclimenes brevicarpalis (Schenkel, 1902) Periclimenes elegans/grandis (Paulson,1875)/(Stimpson,1860) Periclimenes imperator (Schenkel, 1880) Periclimenes soror Nobili, 1904 Periclimenes sp. 1 (n. sp. aff. holthuisi) Pliopontonia furtiva Bruce, 1973 Pontoniopsis comanthi Borradaile, 1915 Stegopontonia commensalis Nobili, 1906 Urocaridella antonbruunii/cyrtorhyncha (Bruce, 1967) / Fujino & Miyake, 1969 Vir orientalis (Dana, 1852) CARIDEA: PANDALIDAE Heterocarpus dorsalis Bate, 1888 Heterocarpus ensifer A. Milne Edwards, 1881 Heterocarpus gibbosus Bate, 1888 Heterocarpus laevigatus Bate, 1888 Heterocarpus lepidus de Man, 1917 Heterocarpus longirostris MacGilchrist, 1905 Heterocarpus sibogae de Man, 1917 Heterocarpus tricarinatus Alcock & Anderson, 1894 Plesionika edwardsii (Brandt, 1851) Plesionika ensis (A. Milne Edwards, 1881) Plesionika martia (A. Milne Edwards, 1883) Plesionika ocellus (Bate, 1888) Taxon USNM 233349 UF 813 UF 841, NNM D-48094 NNM D-48071, UF 3031 UF 838 UF 843 NNM D-48067, UGI 6064 UF 905? NNM D-48073 Museum 28 Notes 3 3 3 3 3 3 3 3 3 3 3 3 31 31, 32 31 31, 32 31 31, 33 31 31 31 31 31 31 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 MS G MS G MS MS MS MS MS MS MS MS S G G G G S G G G S Ag S S G, S G G Hab Island 1 GP518-20 GP413-25 RFM 5-98 GP739-18 GP840-17 GP741-19 GP363-37 29 GP785-10 GP513-23 Photo 5, 10, 29 F 30 F F 5, 10, 29 P O 28 5, 10 15 5, 10 10, 29 5, 10 P F Refs Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) CARIDEA: PROCESSIDAE Nikoides sibogae de Man, 1918 Nikoides steinii (Edmondson, 1935) CARIDEA: RHYNCHOCINETIDAE Cinetorhynchus concolor (Okuno, 1994) Cinetorhynchus hawaiiensis Okuno & Hoover, 1998 Cinetorhynchus hendersoni (Kemp, 1925) Cinetorhynchus hiatti (Holthuis & Hayashi, 1967) Cinetorhynchus reticulatus Okuno, 1997 Cinetorhynchus striatus (Nomura & Hayashi, 1992) PALINURA: PALINURIDAE Justitia longimanus (H. Milne Edwards, 1837) Palibythus magnificus Davie, 1990 Palinurellus wieneckii (de Man, 1881) Panulirus femoristriga (Von Martens, 1872) Panulirus longipes bispinosa Borradaile, 1899 Panulirus ornatus (Fabricius, 1798) Panulirus penicillatus (Olivier, 1791) Panulirus versicolor (Latreille, 1804) PALINURA: SCYLLARIDAE Biarctus vitiensis (Dana, 1852) Galearctus aurora Holthuis, 1981 Parribacus antarcticus (Lund, 1793) ASTACIDEA: ENOPLOMETOPIDAE Enoplometopus holthuisi Gordon, 1968 Enoplometopus occidentalis (Randall, 1840) THALASSINIDEA: AXIIDAE Allaxius clypeatus (de Man, 1887) Allaxius picteti (Zehntner, 1894) Axiopsis pica Kensley, 2003 Axiopsis sp. 1 (aff. pica Kensley, 2003) Taxon GP826-5 GP815-17 P P USNM 243494 UF 468, USNM 243502 USNM 296400 (holotype) UF 59 35 35 35 P GP818-9 GP422-7 GP816-4 C C P UF 2282 UF 2281 UF 895, BPBM UGI 2959 UGI 2961 30 Notes GP764-17 GP574-25 GP864-13 31 GP826-8 IE GP812-29 GP422-26 GP826-10 GP815-21 GP648-8 GP648-19 GP815-2 GP516-25 GP816-15 Photo P P P P P P 15 P O O O 34 O O 5, 25 5, 10 Refs UF 563 UF 1003 UF 1234 UF 894, UGI 2960 UF 705 ?UF 824 UF 903 Museum Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 2 1 2 2 1 1 1 1 1 1 1 1 1 1 1 1 1 1 G G G G G G G G G G G G G G G G, M G G G G G G G S S Hab Island P 5, 40, 41, GP575-9 42, 93 40, 42 GP811-8 UF 3117 ZRC UF 283, UF 586 (S) UF 230 UF 583, UF 600 (S) UF 581, UF 603 (S) Coenobita brevimanus Dana, 1852 Coenobita cavipes Stimpson, 1858 Coenobita perlatus H. Milne Edwards, 1837 Coenobita purpureus Stimpson 1858 Coenobita rugosus H. Milne Edwards, 1837 40 40, 42 40, 42 40, 42 GP811-3 GP809-18 5, 10, 35 GP575-4 GP275-19 UF 142, USNM 243604 UF 470, USNM 243605, BPBM 39, 40 Paraglypturus calderus Türkay & Sakai, 1995 THALASSINIDEA: CALLIANIDEIDAE Callianidea typa H. Milne Edwards, 1837 THALASSINIDEA: STRATHLAXIIDAE Neaxius acanthus (A. Milne Edwards, 1879) ANOMURA: ALBUNEIDAE Albunea speciosa Dana, 1852 ANOMURA: COENOBITIDAE Birgus latro (Linnaeus, 1767) 38 37 USNM 310677 (holotype), UF 1227 USNM 296395 (holotype) USNM 296394 (holotype) Photo 35 GP249-38 35 GP527-5 35 35, 36 35 5, 10, 35 35 GP543:12 35 USNM 243362 UF 58, USNM 296398 USNM 243378 USNM 243530 (holotype) USNM 239352 Axiopsis serratifrons (A. Milne Edwards, 1873) Bouvieraxius springeri Kensley, 1996 Manaxius pitatucensis (de Man, 1925) Marianaxius kroppi Kensley, 1996 Parascytoleptus tridens (Rathbun, 1906) Paraxiopsis bisquamosa (de Man, 1905) Paraxiopsis paulayi Kensley, 2003 Paraxiopsis plumosimanus Kensley, 2003 THALASSINIDEA: CALLIANASSIDAE Corallichirus bayeri Kensley, 2001 Refs Museum Taxon 73 32 Notes Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 T T T T T 1-2 1 1 1 1 1 1 1 1 1 1 1 1 G, S, M, As, P, Gu, T G, S, An, Gu G, An, Ag G, S, An, S, Gu, Al, P, Ag, As, M M G, S, An, Gu, Al, P, Ag, M G G, S G, P E G G G G G G S G G Hab Island BPBM-S 10588 (holotype), UF 40, 42, 43 GP864-23 AHF UF 732 CBM-ZC 4057 (holotype, P) UF 601 USNM 255985, UF 460 UF 1322 UF 1321 UF UF Calcinus guamensis Wooster, 1984 Calcinus haigae Wooster, 1984 Calcinus isabellae Poupin, 1997 Calcinus kurozumii Asakura & Tachikawa, 2000 Calcinus laevimanus (Randall, 1840) Calcinus latens (Randall, 1840) Calcinus lineapropodus Morgan & Forest, 1991 Calcinus minutus Buitendijk, 1937 Calcinus morgani Rahayu & Forest, 1999 Calcinus pulcher Forest, 1958 Calcinus revi Poupin & McLaughlin, 1998 Calcinus seurati Forest, 1951 UF 562 UF UF 430 USNM 152475, UF UF 953 UF 449 BPBM-S 10589 (holotype) UF 325 40, 42, 43, 45 42, 43 42, 43 5, 10, 40, 42, 43 M 44 40 42, 43 42, 43 44 40, 42, 43, 135 42 42 42 P M 42, 43 40, 42, 43 Notes 35 GP854-28 GP519-26 GP751-24 37 GP590-33 GP588-11 38 GP770-36 GP850-20 GP737-5 GP855-1 GP770-29 GP469-15 GP453-13 GP615-7 34 GP584-31 33 Aniculus maximus Edmondson, 1952 Aniculus retipes Lewinsohn, 1982 Aniculus ursus (Olivier, 1811) Aniculus sp. 1 Calcinus areolatus Rahayu & Forest, 1999 Calcinus argus Wooster, 1984 Calcinus elegans (H. Milne Edwards, 1836) 40 UF Photo ANOMURA: DIOGENIDAE Aniculus aniculus (Fabricius, 1787) Refs Museum Taxon Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 G, An,G, Al, Ag, M, As G G G G G G G, An, S, Gu, Al, P, Ag, As, M G, Gu, Ag, M G G P G, S, An, Al, Gu, P, Ag, M G, An, Al, Gu, M G G G, S, An, Al, M G M, A, G, An, S, Gu, P, Ag, M, U Hab Island Ciliopagurus major Forest, 1995 Ciliopagurus pacificus Forest, 1995 Ciliopagurus strigatus (Herbst, 1804) Clibanarius corallinus (H. Milne Edwards, 1848) Clibanarius eurysternus Hilgendorf, 1878 Clibanarius humilis (Dana, 1851) Clibanarius striolatus Dana, 1852 Clibanarius virescens (Krauss, 1843) Dardanus crassimanus (H. Milne Edwards, 1836) Dardanus deformis (H. Milne Edwards, 1836) Dardanus gemmatus (H. Milne Edwards, 1848) Dardanus guttatus (Oliver, 1811) Dardanus lagopodes (Forsskål, 1775) Dardanus megistos (Herbst, 1804) Dardanus scutellatus (H. Milne Edwards, 1848) Diogenes pallescens Whitelegge, 1897 Paguristes jalur Morgan, 1992 Paguristes monoporus Morgan, 1987 Pseudopaguristes janetkae McLaughlin, 2002 Pseudopaguristes sp. 2 (n. sp.) ANOMURA: GALATHEIDAE Allogalathea elegans (White, 1847) Galathea amamiensis Miyake & Baba, 1966 Munidopsis marianica Williams & Baba, 1989 ANOMURA: HIPPIDAE Hippa pacifica Dana, 1852 ANOMURA: LITHODIDAE Lithodes sp. 1 (n. sp.) Paralomis haigae Eldredge, 1976 Paralomis seagranti Eldredge, 1976 Taxon P 40 2, 55 40 C 48 48 UF 252 UF 2283 BPBM-S 8524 (holotype) BPBM-S 8526 (holotype) 46 46 42, 46 42 42 40, 42 42 42 42 42 42 42 42 42 42 M M M 47 M Refs UF 277 UF UF UF UF 326 UF 950 UF UF 141 UF UF AHF 591-01, UF USNM 270096, UF UF UF UF UF Museum GP527-27 GP254-15 GP762-27 GP413-28 GP833-13 GP517-26 GP764-31 GP681-21 GP413-38 GP770-25 GP544-30 GP848-27 GP754-9 GP771-14 GP751-27 GP753b-10 GP850-25 GP850-18 GP759-11 GP850-9 Photo Notes Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 3 3 3 1 1 1 3 2 2 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 2 2 G G G G, Ag G M MTV G G G G G G, Gu, Al G G G G G G G G G G G G G G Hab Island USNM 255986 USNM 255976 USNM 255959 USNM 255955 USNM 233491 Pylopaguropsis keijii McLaughlin & Haig, 1989 Trichopagurus trichophthalmus (Forest, 1954) ANOMURA: PARAPAGURIDAE Sympagurus dofleini (Balss, 1912) Tylaspis anomala Henderson, 1885 ANOMURA: PORCELLANIDAE Neopetrolisthes maculatus (H. Milne Edwards, 1837) Pachycheles garciaensis (Ward, 1942) Pachycheles pisoides (Heller, 1865) Pachycheles spinipes (A. Milne Edwards, 1873) Petrolisthes asiaticus (Leach, 1820) RK USNM 231413 (holotype), UF 597 USNM 231415 (paratype) UF 644 UF UF 432 UF UF UF UF 307 UF 645 AHF UF 5 USNM 259396 (holotype) ANOMURA: PAGURIDAE Catapaguroides foresti McLaughlin, 2002 Paguritta kroppi McLaughlin & Lemaitre, 1993 Pagurixus anceps (Forest, 1954) Pagurixus boninensis (Melin, 1939) Pagurixus cf. laevimanus (Ortmann, 1892) Pagurixus cf. tweediei (Forest, 1956) Pagurixus maorus (Nobili, 1906) Pagurixus sp. 2 (n. sp. fast purple) Pagurixus sp. 3 (slate) Pagurixus sp. 8 (pale) Pagurixus sp. 9 (n. sp., purple claw) Pagurus hirtimanus Miers, 1880 Pagurus? sp. 1 Porcellanopagurus sp. 1 (n. sp.) Kropp & Eldredge, MS Pylopaguropsis fimbriata McLaughlin & Haig, 1989 Museum Taxon K K 23 K 40 53 54 52 M 40 5, 10 40 M M M 51 M M M K 52 49 50 40 Refs Notes GP776-24 RK GUM218I GP845-25 GP867-5 GP847-10 41 GP751-32 GP813-3 GP820-10 GP860-8 40 GP860-21 GP846-13 GP867-2 GP859-33 GP846-18 36 GP469-25 39 Photo Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 3 3 1 1 1 1 1 1 1 1 1 1 1 1 3 1 1 1 1 G G G, S G G, M G G G G G G Gu, Ag, M, Al G, P G M G G G G G G G G G Hab Island 72 P USNM 257238 UF 477, USNM 257230 USNM 210534 KC Paratype: USNM 222328 USNM 257218 AHF USNM 222529 (holotype) USNM 189361 USNM 210539 USNM 210624 (holotype, P) USNM 210628 (holotype) USNM 257240 UF 334, USNM 257606 UF 481, ZRC USNM 282895 Petrolisthes carinipes (Heller, 1861) Petrolisthes coccineus (Owen, 1839) Petrolisthes decacanthus Ortmann, 1897 Petrolisthes donanensis Osawa, 1997 Petrolisthes eldredgei Haig & Kropp, 1988 Petrolisthes elegans Haig, 1981 Petrolisthes extremus Kropp & Haig, 1994 Petrolisthes heterochrous Kropp, 1986 Petrolisthes lamarckii (Leach, 1820) Petrolisthes masakii Miyake, 1943 Petrolisthes mesodactylon Kropp, 1984 Petrolisthes miyakei Kropp, 1984 Petrolisthes pubescens Stimpson, 1858 Petrolisthes tomentosus (Dana, 1852) BRACHYURA: AETHRIDAE Aethra edentata Edmondson, 1951 Aethra scruposa (Linnaeus, 1764) BRACHYURA: BYTHOGRAEIDAE Austinograea williamsi Hessler & Martin, 1989 BRACHYURA: CALAPPIDAE Calappa calappa (Linnaeus, 1758) Calappa gallus (Herbst, 1803) UF 75, BPBM UF 82, USNM 277628, UGI 2580 23, 40 40, 58 58 Paratype: USNM 210633 Petrolisthes borradailei Kropp, 1984 N G 2, 59 57 58 23, 40 K 23, 40 56 K K K 40 56 23, 40 56 USNM 257207 Petrolisthes bispinosus Borradaile, 1900 Refs Museum Taxon GP856-7 GP814-26 GP780-10 GP366-7 GP751-6 GP751-5 RK GUM221G GP750-28 RK GUM208P RK GUM208M RK GUM221C RK GUM214A Photo Notes Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 3 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 G G MTV G G G G, Gu, S G, S, An G P, Gu, An, M G, M G, M, An G, P, S G G G G, S, An, Gu, P, M G G, M G, S G, S Hab Island 68, 137 N N N N K K 81, 137 97 98 K K K 98 99 K K 5, 98 K K 97 99 K 100 K 101 101 101 101 UF 81, BPBM, UGI 2575 UF/ZRC USNM UF 85 UF 3094, ZRC USNM UF 869, USNM UF 64, BPBM USNM 257551 USNM KC KC KC USNM KC USNM 257250 USNM USNM 1004935 USNM 257245 USNM 257345 USNM 257368 KC USNM 257336 USNM 268890 (holotype) USNM 257311 USNM 234266 (holotype) USNM 257478 USNM 234264 USNM 234269 (holotype) Calappa hepatica (Linnaeus, 1758) Calappa cf. matsuzawa Galil, 1997 Calappa sp. 1 (aff. sebastiani Galil, 1997) Calappa sp. 2 Calappa sp. 3 (n. sp.) Ng et al., in prep. Cycloes granulosa de Haan, 1837 BRACHYURA: CARPILIIDAE Carpilius convexus (Forsskål, 1775) Carpilius maculatus (Linnaeus, 1758) BRACHYURA: CRYPTOCHIRIDAE Cryptochirus coralliodytes Heller, 1861 Dacryomaia edmonsoni (Fize & Serène, 1956) Dacryomaia japonica (Takeda & Tamura, 1981) Dacryomaia sp. 1 (n. sp.) Kropp, MS Dacryomaia sp. 2 (n. sp.) Kropp, MS Fizesereneia heimi (Fize & Serène, 1956) Fizesereneia latisella Kropp, 1994 Fizesereneia stimpsoni (Fize & Serène, 1956) Fungicola utinomii (Fize & Serène, 1956) Hapalocarcinus marsupialis Stimpson, 1859 Hiroia krempfi (Fize & Serène, 1956) Lithoscaptus helleri (Fize & Serène, 1957) Lithoscaptus paradoxus A. Milne Edwards, 1862 Lithoscaptus prionotus Kropp, 1994 Lithoscaptus tri (Fize & Serène, 1956) Luciades agana Kropp & Manning, 1996 Neotroglocarcinus hongkongensis (Shen, 1936) Opecarcinus aurantius Kropp, 1989 Opecarcinus crescentus (Edmondson, 1925) Opecarcinus granulatus (Shen, 1936) Opecarcinus lobifrons Kropp, 1989 Refs Museum Taxon GP644-23 GP831-36 GP820-13 GP410-7 GP579-35 RK GP819-33 GP739-27 Photo Notes Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 3 1 1 1 G G G G G G G G G G, S G G G G G G G G G G G G G G, S G G G G G Hab Island Dynomene praedator Milne Edwards, 1879 Hirsutodynomene spinosa (Rathbun, 1911) Paradynomene tuberculata Sakai, 1963 Dynomene hispida Guérin-Méneville, 1832 Dynomene kroppi McLay, 2001 BRACHYURA: DROMIIDAE Cryptodromia fallax (Lamarck, 1818) Cryptodromia hilgendorfi de Man, 1888 Cryptodromia pentagonalis Hilgendorf, 1879 Cryptodromia pitiensis McLay, 2001 Cryptodromia tumida Stimpson, 1858 Cryptodromiopsis plumosa (Lewinsohn, 1984) Cryptodromiopsis unidentata (Rüppell, 1830) Dromia dormia (Linnaeus, 1763) Dromidiopsis lethrinusae (Takeda & Kurata, 1976) Takedromia cristatipes (Sakai, 1969) BRACHYURA: DYNOMENIDAE Dynomene guamensis McLay, 2001 MNHN-B26476 (holotype), UF 401 ZRC 2000.0743 ZRC 2000.0745, UF 313 ZRC 2000.2122, UF 21 61, 63 61, 63 61, 63 62 61 GP532-31 GP819-25 GP832-18 RK GUM 221F 61 MNHN-B26474 (holotype) ZRC 2000.2121 ZRC 2000.0755 60, 61, 62 42 61 61 61 61 61 60, 61 GP380-32 60 61 61 Notes ZRC 2000.0751 USNM 128571, ZRC 2000.2119 ZRC 2000.2115 MNHN-B27303 (holotype) ZRC 2000.2113 ZRC 2000.2112 UGI 6020 5, 10, 62 GP845-4 UF 1953 Photo 101 101 K 102 98 98 98 USNM 234275 (holotype) USNM 234278 (holotype) USNM USNM 257314 USNM 257285 USNM 257512 USNM 257277 Opecarcinus pholeter Kropp, 1989 Opecarcinus sierra Kropp, 1989 Pelycomaia minuta (Edmondson, 1933) Pseudocryptochirus viridis Hiro, 1938 Pseudohapalocarcinus ransoni Fize & Serène, 1956 Utinomiella dimorpha (Henderson, 1906) Xynomaia sheni (Fize & Serène, 1956) BRACHYURA: DAIRIDAE Daira perlata (Herbst, 1790) Refs Museum Taxon Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 G G, As G M G G G, M G G G G G G S G G G, S, Gu, P, Al G G G G G G G Hab Island Grapsus sp. 1 (aff. longitarsis Dana, 1851) UF 134 UF 92 ZRC UF 93 P 62 62 62 L 62 5, 62 UF 111 62 62 K N 128 62 K 62 92 92 92, 133 62, 92, 93 92 94 UF 113, USNM 154148, BPBM USNM, ZRC UF/ZRC USNM, UGI, UF 2842 UF 282, BPBM, ZRC UF 293, USNM 154842 UF 868, USNM, UGI Epixanthus corrosus A. Milne Edwards, 1873 Eriphia scabricula Dana, 1852 Eriphia sebana (Shaw & Nodder, 1803) Globopilumnus sp. 1 Hypothalassia armata (de Haan, 1835) Lydia annulipes (H. Milne Edwards, 1834) Ozius rugulosus Stimpson, 1858 Ozius tuberculosus H. Milne Edwards, 1834 62 5, 10 K Refs UF 114 UF 145, BPBM, ZRC UF 147, USNM 181977, ZRC UF 144, USNM 155049, ZRC UF 89, ZRC ZRC USNM UF 70, USNM, ZRC BRACHYURA: ERIPHIIDAE Dacryopilumnus eremita Nobili, 1906 Dacryopilumnus rathbunae Balss, 1932 Epixanthoides anomalus Balss, 1935 BRACHYURA: GECARCINIDAE Cardisoma carnifex (Herbst, 1796) Discoplax hirtipes (Dana, 1851) Discoplax longipes A. Milne Edwards, 1867 Discoplax rotunda (Quoy & Gaimard, 1824) Epigrapsus notatus (Heller, 1865) Epigrapsus sp. 1 (n. sp.) Ng, 2003 BRACHYURA: GRAPSIDAE Geograpsus crinipes (Dana, 1851) Geograpsus grayi (H. Milne Edwards, 1853) Geograpsus stormi de Man, 1895 Grapsus albolineatus Lamarck, 1818 Grapsus longitarsis Dana, 1851 Grapsus tenuicrustatus (Herbst, 1783) Museum Taxon Notes GP414-11 GP338-25 GP809-30 GP811-20 GP810-7 GP791-29 GP331-32 GP809-1 GP275-29 64 GP333-34 PKLN GP403-2 GP357-32 GP829-20 RK GP414-4 GP463-37 GP786-17 RK-GUM 208G Photo Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 T T T T T 1 1 1 1 1 3 1 1 1 1 1 1 G, Gu, M An, Gu, U G, M G G, An, Ag G, S, An, Gu, Sa, P, Ag, M, Al G G G G G, M G G M G, Al G G G G, An, Ag G G, An, Gu, Ag, As M G, S G Hab Island Intesius sp. 1 (n. sp.) Crosnier & Ng, 2003 Parapilumnus oryctus Ng, 2003 BRACHYURA: HYMENOSOMATIDAE Elamena sp. 1 BRACHYURA: LEUCOSIIDAE Alox rugosum (Stimpson, 1858) Drachiella angulata (Ihle, 1918) Ebalia(?) sp. 1 Ebaliopsis erosa (A. Milne Edwards, 1874) Heteronucia venusta Nobili, 1906 Leucosia sp. 1 Myra fugax (Fabricius, 1798) New genus? sp. 1 Nucia speciosa Dana, 1852 Nucia? sp. 1 Nucia? sp. 2 143 83 T P B N B B B P B 5, 10 N N UF 628 UF 729 UF 1019 UF 728 UF 621 UF 610 UF 67 UF 2937 UF 1021 UF 670 UF 285 UF/ZRC 82 UF 2098 (holotype), ZRC 2002.17 BPBM ZRC 2002.174 (holotype) 64 66 142 64, 65 BPBM-S 8595 (holotype), ZRC, UGI 6767 BPBM S10479, UGI 6764 ZRC BPBM (paratype) Homola orientalis Henderson, 1888 Lamoha personata (Guinot & Richer de Forges, 1981) Lamoha williamsi (Takeda, 1980) BRACHYURA: GONEPLACIDAE Conleyus defodio Ng & Ng, 2003 P P P 62 127 62 UF 101 UF 103 UF 95, UF 591 (S) UF 123, UF 590 (S) UF 129 UF 130, UF 584 (S), BPBM Metopograpsus latifrons (White, 1874) Metopograpsus oceanicus (Jacquinot, 1852) Metopograpsus thukuhar (Owen, 1839) Pachygrapsus minutus A. Milne Edwards, 1873 Pachygrapsus planifrons de Man, 1888 Pachygrapsus plicatus (H. Milne Edwards, 1837) BRACHYURA: HOMOLIDAE Homola dickinsoni Eldredge, 1980 Refs Museum Taxon GP597-26 GP527-21 GP818-20 GP408-24 GP537-1 GP408-22 GP533-12 GP596-23 GP391-9 GP469-33 GP694-21 GP410-2 GP409-34 GP526-28 GP409-7 GP871-20 Photo 71 Notes Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 3 1 1 3 3 3 3 1 1 1 1 1 1 G G G G G G G G G, S G G G G G G G G S G G G G, S G, S, M, Al G G, S, P Hab Island Micippa parca Alcock, 1895 Micippa platipes Rüppell, 1830 Perinia tumida Dana, 1852 Pseudomicippe eldredgei Griffin & Tranter, 1986 Rochinia decipiata Williams & Eldredge, 1994 Schizophroida hilensis (Rathbun, 1906) Schizophrys aspera (H. Milne-Edwards, 1834) Simocarcinus obtusirostris (Miers, 1879) Tiarinia cornigera (Latreille, 1825) Tiarinia spinigera Stimpson, 1907 Tylocarcinus styx (Herbst, 1803) Xenocarcinus depressus Miers, 1874 UF 659 UF 666, UGI UF 665 AHF (holotype), UF 425, ZRC USNM 251434 (holotype) UF 667, ZRC, UGI UF 287, UGI UF 688, UGI UF 312, UGI UF 629 UF 690 USNM 250884 (holotype, S) USNM 282898, UF 2918 UF 855, ZRC, UGI 2945 UF 15, UGI UF 857 ZRC UF 394, UGI 2652 UF 655 UF 850 UF 289, UGI UF 79, USNM 134298, UGI 2646 USNM 266801 (paratype) USNM 255984 (holotype), UF 530 UF 856 Oreotlos lagarodes Tan & Ng, 1995 Oreotlos pala Tan & Ng, 1995 "Randallia" serenei Richer de Forges, 1983 BRACHYURA: MAJIDAE Achaeus lacertosus? Stimpson, 1907 Achaeus laevioculis (de Man, 1905) Bothromaia griffini Williams & Moffitt, 1991 Camposcia retusa Latreille, 1829 Criocarcinus superciliosus (Linnaeus, 1767) Cyclax suborbicularis (Stimpson, 1858) Cyrtomaia intermedia Sakai, 1938 Dumea sp. 1 (aff. taiwanicus (Loh & Wu, 1998)) Huenia proteus? de Haan, 1839 Hyastenus convexus Miers, 1884 Hyastenus hilgendorfi de Man, 1887 Hyastenus planasius (Adams & White, 1848) Menaethius monoceros (Latreille, 1825) Museum Taxon 5, 10 5, 10 5, 10, 62 130 71 N N 5, 10 N 62 5, 10 N N N 67 P N N 70 N N N N N 5, 10, 62 N 69 69 Refs Notes 44 GP540-6 GP826-31 GP405-3 43 GP837-22 GP537-17 GP414-23 GP576-10 GP814-13 GP357-12 GP590-8 GP367-13 GP692-22 GP463-1 GP823-28 GP583-6 GP733-24 Photo Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 3 1 1 1 1 1 1 1 1 1 3 1 1 1 3 1 1 1 1 1 1 3 1 1 S G, S S, M G G G G G, S G Ag S G G G S, E G G G, As G G G G G G G, S, Gu G G G Hab Island UF 2102, ZRC 2002.215 UF 419 Rhinolambrus rudis (Rathbun, 1916) Thyrolambrus efflorescens (Alcock, 1895) 73 73 UF 2099, ZRC 2002.201 UF 2101 (holotype), ZRC 2002.204 UF 9, ZRC 2002.206 UF 6, USNM Furtipodia petrosa (Klunzinger, 1906) Lambrachaeus ramifer Alcock, 1895 Neikolambrus polemistes Tan & Ng, 2003 Parthenopoides cariei Bouvier, 1914 T R UF 517 UF 330 GP414-12 GP540-29 GP597-22 GP408-31 GP412-34 GP412-25 GP409-17 GP336-3 GP336-17 GP412-13 Photo 73 73, 74 73 144 73 73, 74 RK GUM221D GP250-35 GP836-8 GP540-8 73 5, 10, 73, GP836-3 74 73, 74 73 T T P K N K N 5, 62 62 Refs UF 118, BPBM, UGI 6737 UF 2846 UF 98, BPBM, ZRC UF 97 UF 987, BPBM UF 115, BPBM, USNM 168462, ZRC UF 96 UF 119 UF/ZRC Museum UF/ZRC ZRC 2002.211 (holotype), UF 2103 UF 8, ZRC 2002.213 UF/ZRC ZRC 2002.214 (holotype) UF 37 Daldorfia leprosa (Nobili, 1905) Furtipodia gemma Tan & Ng, 2003 Certolambrus pugilator (A. Milne Edwards, 1873) Daldorfia horrida (Linnaeus, 1758) Uca crassipes (Adams & White, 1848) Uca vocans (Linnaeus, 1758) Ocypodidae sp. 2 BRACHYURA: PALICIDAE Crossotonotus spinipes (de Man, 1888) Exopalicus maculatus (Edmondson, 1930) BRACHYURA: PARTHENOPIDAE Aulacolambrus hoplonotus (Adams & White, 1848) Aulacolambrus hystricosus Tan & Ng, 2003 BRACHYURA: OCYPODIDAE Macrophthalmus convexus Stimpson, 1858 Macrophthalmus latreillei (Desmarest, 1822) Macrophthalmus verreauxi H. Milne Edwards, 1848 Macrophthalmus sp. 1 Ocypode ceratophthalma (Pallas, 1772) Ocypode cordimana Desmarest, 1825 Taxon 72 Notes Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 G G G G G G G G G G, S G G G G G G G G G G G G, S, Ag G, P Hab Island BRACHYURA: PILUMNIDAE Actumnus sp. 1 (n. sp.) Ng, in prep. Caecopilumnus hirsutus (Borradaile, 1903) Echinoecus pentagonus (A. Milne Edwards, 1879) Glabropilumnus dispar (Dana, 1852) Glabropilumnus laevimanus (Dana, 1852) Lophoplax? sp. 1 Nanopilumnus coralliophilus (Takeda & Miyake, 1969) Permanotus purpurea (Gordon, 1934) Pilumnus cursor A. Milne Edwards, 1873 Pilumnus cf. oahuensis Edmondson, 1931 Pilumnus cf. scabriusculus Adams & White, 1849 Pilumnus longicornis Hilgendorf, 1898 Pilumnus serenei Ng, 1988 Pilumnus taeniola Rathbun, 1906 Pilumnus vespertilio (Fabricius, 1798) Pilumnus sp. 1 (n. sp.) Ng, in prep. Pseudolitochira integer (Miers, 1884) Tanaocheles stenochilus Kropp, 1984 Tiaramedon spinosum (Miers, 1879) Viaderiana aranea (Tesch, 1918) Viaderiana quadrispinosa (Zehntner, 1894) Viaderiana sp. 4 Viaderiana typica? Ward, 1942 Viaderiana? sp. 5 BRACHYURA: PINNOTHERIDAE Aphanodactylus sp. 1 (terebellid commensal) BRACHYURA: PLAGUSIIDAE Percnon abbreviatum (Dana, 1851) Percnon affine (H. Milne Edwards, 1853) Percnon guinotae Crosnier, 1965 Taxon N P P P UF 106 UF 88 UF 104, ZRC N P N N N N 62 75 5, 10 N N N N N K N 5, 10 89, 131 R 5, 10 N N N N Refs UF/ZRC UF 80, UGI UF 310 UF/ZRC UF/ZRC USNM 210636 (holotype) USNM, UF/ZRC UF 723 UF/ZRC UF 724 UF 787 UF 725 USNM, UF/ZRC UF/ZRC UF 482 UF 358 UF/ZRC USNM 154036 UF 715 UF 716 UF 785 Museum 66 Notes GP808-21 GP512-24 GP512-10 GP786-13 GP814-17 GP837-29 GP528-2 GP827-20 GP466-7 GP466-22 GP466-26 GP393-31 66 45 50 GP783-22 66 GP592-21 ?GP532-19 GP858-15 GP534-2 GP580-3 Photo Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 G G G G G G G G G G M G S G G G G G G G S G G S G G G G Hab Island 76 UF 2091 (holotype), ZRC 2002.173 UF 167, UF 588 (S) UF 2588 UF 411 UF 152 UF/ZRC UF 54 UF 182, USNM UF 1915 UF 158 UF 151 UF 192 UF 179 UF 3149 UF/ZRC UF 472 UF 2071, UF 972 (As) UF UF 163 UF 204 UF 169, BPBM UF/ZRC UGI 6729 UF 241 Caphyra rotundifrons (A. Milne Edwards, 1869) Caphyra sp. 1 (on Asterospicularia randalli) Caphyra unidentata? Lenz, 1910 Carupa ohashii Takeda, 1993 Carupa sp. 2 (n. sp.) Ng & Takeda, in prep. Carupa tenuipes Dana, 1851 Catoptrus nitidus A. Milne Edwards, 1870 Charybdis erythrodactyla (Lamarck, 1818) Charybdis hawaiensis Edmondson, 1954 Charybdis hellerii (A. Milne Edwards, 1867) Charybdis obtusifrons Leene, 1936 Charybdis orientalis Dana, 1852 Charybdis paucidentata (A. Milne Edwards, 1861) Charybdis sp. 1 (n. sp. aff. omanensis Leene, 1934) Coelocarcinus foliatus Edmondson, 1930 Laleonectes nipponensis (Sakai, 1938) Libystes nitidus? A. Milne Edwards, 1867 Libystes villosus Rathbun, 1924 Lissocarcinus laevis Miers, 1886 Lissocarcinus orbicularis Dana, 1852 Lupocyclus quinquedentatus Rathbun, 1906 Podophthalmus vigil (Weber, 1795) Portunus alcocki (Nobili, 1905) 62?, 129 125 62?, 129 ZRC UF 87, BPBM ZRC Plagusia immaculata Lamarck, 1818 Plagusia speciosa Dana, 1851 Plagusia squamosa (Herbst, 1790) BRACHYURA: PORTUNIDAE Atoportunus gustavi Ng & Takeda, 2003 5, 10 N S P T P P 62 N N P N P N 77 P P P P 5, 10 P P N 62 BPBM 1671 Percnon planissimum (Herbst, 1804) Refs Museum Taxon Notes GP422-1 GP617-15 GP592-12 GP411-23 GP410-14 GP533-7 GP693-10 GP598-16 GP459-6 GP397-25 GP828-24 GP749-12 GP597-18 GP513-19 GP834-34 GP818-32 GP767-4 GP694-24 GP571-4 GP790-23 GP832-12 GP583-36 65 GP512-22 65 GP424-7 Photo Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 1 2 1 1 1 1 1 1 1 1 1 1 1 1 1 1 G, S G G G G G G G, Ag G G G G G G G G, As G G G G, S G G G G G, An, P, Ag, M G, ?Ag G G, ?Ag Hab Island Thalamita woodmasoni Alcock, 1899 Thalamita auauensis Rathbun, 1906 Thalamita cf. cerasma Wee & Ng, 1995 Thalamita coeruleipes Jacquinot, 1846 Thalamita corrugata Stephenson & Rees 1961 Thalamita crenata H. Milne Edwards, 1834 Thalamita dakini Montgomery, 1931 Thalamita exetastica Alcock, 1899 Thalamita gatavakensis Nobili, 1906 Thalamita integra Dana, 1852 Thalamita medipacifica Edmondson, 1954 Thalamita mitsiensis Crosnier, 1962 Thalamita picta Stimpson, 1958 Thalamita pilumnoides Borradaile, 1903 Thalamita prymna (Herbst, 1803) Thalamita pseudopelsarti Crosnier, 2002 Thalamita quadrilobata Miers, 1884 Thalamita sima H. Milne Edwards, 1834 Thalamita spinifera Borradaile, 1902 Thalamita spinimera Stephenson & Rees, 1967 UF 209 UF 166, BPBM UF 171 UF 183 UF 188 USNM UF 165 Portunus convexus de Haan, 1833 Portunus granulatus (H. Milne Edwards, 1834) Portunus longispinosus Rathbun, 1906 Portunus macrophthalmus Rathbun, 1906 Portunus petreus (Alcock, 1899) Portunus sanguinolentus (Herbst, 1783) Portunus tuberculosus (A. Milne Edwards, 1861) Scylla serrata (Forsskål, 1775) Thalamita admete (Herbst, 1803) Refs P P N N N 78 N 79 UF 164, USNM 149616, BPBM 5, 10, 62, 78, 141 USNM 78 UF 218 P UF 149, USNM 78 UF 180 P UF 205, BPBM 78 USNM 78 UF 206 N 80 UF 200, UF 587 (S), USNM 78 UF 197 N UF 190 N UF 177, UGI 5, 10, 62 5,10 UF 221, BPBM 78 UF 156, UGI, ZRC 126 UF 201 N 141 UF 208, USNM 255982 N USNM 112418 (holotype), 78 UF 173 5, 10 Museum Taxon GP813-19 GP576-36 GP576-19 GP382-3 GP544-13 GP540-12 GP749-24 GP412-30 GP467-24 GP850-30 GP356-25 GP409-28 GP414-36 GP391-17 GP810-28 GP847-19 GP516-30 GP387-19 GP580-16 Photo Notes Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 S G G G G G G G G G, S, Gu, M, U G G G G G G G S G G G G, S, Al S G G G G G G Hab Island P N N N 5, 10 N N N N P 124 N 85 85 85 85 85 85 85 85, 90 85 UF 90, ZRC UF/ZRC UF 91, ZRC UF 108, ZRC UF 99, ZRC UF 107, ZRC ZRC UF 128 UF/ZRC? UF, ZRC UF 125 SMF 26927 USNM (S) USNM, SMF 26282 UF 911, SMF 26929 USNM (S) USNM 285980, SMF 26284 BPBM 1629, SMF 25171 SMF 26285 N 62 UF 302 UF 294, BPBM, ZRC UF 1013 67 USNM 250886 BRACHYURA: RANINIDAE Raninidae sp. 1 BRACHYURA: SESARMIDAE Chiromantes obtusifrons (Dana, 1852) Clistocoeloma sp. 1 Labuanium rotundatum (Hess, 1865) Metasesarma rousseauxi H. Milne Edwards, 1853 Neosarmatium fourmanoiri Serène, 1973 Neosarmatium meinerti (de Man, 1887) Neosarmatium trispinosum Davie, 1994 Parasesarma pictum (De Haan, 1835) Sesarma s.l. sp. 2 Sesarmoides guamensis Ng, 2002e Stelgistra stormi de Man, 1895 BRACHYURA: TRAPEZIIDAE Quadrella sp. Tetralia cinctipes Paulson, 1875 Tetralia fulva Serène, 1984 Tetralia rubridactyla Garth, 1971 Tetralia vanninii Galil & Clark, 1988 Tetraloides heterodactyla (Heller, 1861) Tetraloides nigrifrons (Dana, 1852) Trapezia cymodoce (Herbst, 1799) Trapezia digitalis Latreille, 1828 5, 10 UF 170, USNM 255987 Thalamitoides quadridens A. Milne Edwards, 1869 BRACHYURA: POUPINIIDAE Poupinia hirsuta Guinot, 1991 BRACHYURA: PSEUDOZIIDAE Euryozius? sp. 1 Pseudozius caystrus (Adams & White, 1848) Refs Museum Taxon Notes GP856-27 PKLN HCL GP409-22 GP758-17 GP423-8 GP758-28 GP656-11 GP738-9 GP810-12 GP850-26 67 GP836-23 67 GP361-14 Photo Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 T T 1 1 1 1 1 1 1 1 1 1 3 1 G G S G, S G S G, S G G, M G G G G, S G G G G G G G P G G, Sa, Gu, M, U Ar G, S Hab Island UF 1977, BPBM 1616 UF 386 UF 758 USNM 154036, UF 2991 UF 585, BPBM, UGI 2593 1 1 1 1 FW FW 1 1 FW 1 62 P P N 95 P N N P 96 5, 10 GP858-23 N 5, 10 46 K GP782-12 5, 10 47 5, 10, 137 GP366-27 GP738-18 PKLN GP733-22 GP738-2 GP738-30 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 G, S G G, S G S G, S M G G G G G G G G E G, S G, S, As, M G, M G, M G, S G, S G, S G S, Ag G, S G G Hab Island UF 133 UF/ZRC UF/ZRC UF 112, ZRC UGI 2621 UF 126 UF/ZRC UGI 2626 SMF 22945 Notes 1 GP656-26 GP644-31 GP290-15 GP656-20 GP656-2 Photo 85 Trapezia speciosa Dana, 1852 BRACHYURA: VARUNIDAE Cyclograpsus integer H. Milne Edwards, 1837 Cyclograpsus sp. 1 Cyclograpsus sp. 2 Helice subquadrata (Dana, 1851) Orcovita mollitia Ng, Guinot, Iliffe, 1996 Ptychognathus sp. 1 Ptychognathus sp. 2 (cf. barbatus (A. Milne Edwards, 1873)) Scutumara sp. 1 Varuna litterata (Fabricius, 1798) Xenograpsus novaeinsularis Takeda & Kurata, 1977 BRACHYURA: XANTHIDAE Actaea polyacantha (Heller, 1861) Actaea spinosissima Borradaile, 1902 Actaeodes hirsutissimus (Rüppell, 1830) Actaeodes tomentosus (H. Milne Edwards, 1834) Atergatis dilatatus de Haan, 1835 Atergatis floridus (Linnaeus, 1767) 85 85 85 85 85, 91 85 85 85 85 85, 91 85, 91 BPBM-S 11880, SMF 26289 ZRC SMF 26286 MNHN-B 26132 USNM, SMF 25201, UGI 2607 BPBM S1629, USNM SMF 26290 (holotype) USNM 260835 (S) BPBM-S 11879, USNM USNM 286485, MNHN-B 9714 MNHN B-9681 (holotype), USNM 285933 USNM Trapezia ferruginea Latreille, 1828 Trapezia flavopunctata Eydoux & Souleyet, 1842 Trapezia formosa Smith, 1869 Trapezia garthi Galil, 1984 Trapezia guttata Rüppell, 1830 Trapezia lutea Castro, 1997 Trapezia neglecta Castro, 2003 Trapezia plana Ward, 1941 Trapezia rufopunctata (Herbst, 1799) Trapezia septata Dana, 1852 Trapezia serenei Odinetz, 1984 Refs Museum Taxon Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) USNM 154154 UF 757 UF 589 (S), UGI UF 871 UF 3560, USNM UF 862, BPBM Etisus electra (Herbst, 1801) Etisus frontalis (Dana, 1852) Etisus laevimanus Randall, 1840 Etisus splendidus Rathbun, 1906 Etisus utilis Jacquinot, 1854 Euxanthus exsculptus (Herbst, 1790) Forestia depressa (White, 1847) Gaillardiellus orientalis (Odhner, 1925) Gaillardiellus superciliaris (Odhner, 1925) Hypocolpus abbotti (Rathbun, 1894) UF 726 UF 861 P P N P P N P N 5, 10 UF 391 5, 10, 62 5, 10 UF 788 5, 10 UF 863 P UF 711 5, 10, 62 62 5, 10 UF 76, USNM N UF 804 N UF 341, UGI N UF 740, USNM 154153, BPBM P 5, 10 N P P K K 62 N 5, 10 P UF 874 UF 706 UF 714 UF 272 UF 3147 UF 875, UGI 6753 UF 880 UF 797 Atergatis granulatus de Man, 1889 Atergatis latissimus (H. Milne Edwards, 1834) Atergatis nitidus A. Milne Edwards, 1865 Atergatis subdentatus de Haan, 1835 Atergatopsis granulatus A. Milne Edwards, 1861 Atergatopsis lucasi Montrouzier, 1865 Atergatopsis signatus (Adams & White, 1848) Banareia cf. armata A. Milne Edwards, 1869 Chlorodiella barbata (Borradaile, 1900) Chlorodiella cytherea (Dana, 1852) Chlorodiella laevissima (Dana, 1852) Cymo andreossyi (Audouin, 1826) Cymo quadrilobatus Miers, 1884 Domecia glabra Alcock, 1899 Domecia hispida Eydoux & Souleyet, 1842 Etisus molokaiensis (Rathbun, 1906) Etisus anaglyptus H. Milne Edwards, 1834 Etisus bifrontalis (Edmondson, 1935) Etisus demani Odhner, 1925 Etisus dentatus (Herbst, 1785) Refs Museum Taxon GP828-18 GP414-17 GP319-32 GP597-33 GP808-9 RFM145N GP827-25 GP656-6 GP859-21 GP819-16 GP598-6 GP598-31 GP367-18 Photo 49 48 Notes Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 G, S G G G G G P, M G S G G G G G G G G G S G, S, P, M S G, S G G, S, M As S G G G G Hab Island USNM USNM Jonesius triunguiculatus (Borradaile, 1902) Juxtaxanthias lividus (Lamarck, 1818) Juxtaxanthias tetraodon (Heller, 1865) Kraussia rugulosa (Krauss, 1843) Lachnopodus bidentatus (A. Milne Edwards, 1867) Lachnopodus rodgersi Stimpson, 1858 Lachnopodus subacutus (Stimpson, 1858) Leptodius exaratus (H. Milne Edwards, 1834) Leptodius nudipes (Dana, 1852) Leptodius sanguineus (H. Milne Edwards, 1834) USNM 154732 USNM 154139 UF 568, USNM UF 718 USNM 155973 UF 777, USNM 93940 UF/ZRC UF 779 UF 896 UF 805 UF 803, USNM 286496 BPBM S1615 Liomera semigranosa de Man, 1888 Liomera stimpsoni (A. Milne Edwards, 1865) Liomera striolata (Odhner, 1925) Liomera tristis (Dana, 1852) Liomera? sp. 1 Liomera sp. 2 (n. sp.? aff. tristis (Dana, 1852)) Lophozozymus bertonciniae Guinot & Richer de Forges, 1981 Lophozozymus edwardsi (Odhner, 1925) Lophozozymus incisus (H. Milne Edwards, 1834) Lophozozymus pictor (Fabricius, 1798) Lophozozymus pulchellus A. Milne Edwards, 1867 USNM 154716 Liocarpilodes armiger pacificus Balss, 1938 Liocarpilodes harmsi Balss, 1934 Liocarpilodes integerrimus (Dana, 1852) Liomera bella (Dana, 1852) Liomera cinctimana (White, 1847) Liomera monticulosa (A. Milne Edwards, 1873) Liomera rugata (H. Milne Edwards, 1834) UF 860 USNM, UGI UF 423, USNM UF 292, USNM 134558, UGI UF 886 USNM 134551 Museum Taxon Photo 5, 10 N K 5, 10 N N N N N K 5, 10, 62 5, 10 62 5, 10 5, 10 K 5, 10 62 54 53 55 51 52 Notes GP357-3 56 GP865-5 GP615-9 GP422-29 GP519-18 GP544-10 GP456-26 RK-GUM208F GP832-24 84, 85 K 86 5, 10, 62 GP814-28 K 62 N GP751-21 K K 62 GP836-31 Refs Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 2 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 S G G G, S G G G G G G S, Ag G G G G, S, Al G M G G G G, Aan, Gu, P, Ag, M, Al G, S Gu, M G, S G, S G S G, P, M Hab Island Palapedia sp. 1 Palapedia sp. 2 Paractaea rufopunctata complex (H. Milne Edwards, 1834) Paramedaeus noelensis (Ward, 1934) Paramedaeus simplex (A. Milne Edwards, 1873) Paraxanthias notatus (Dana, 1852) Paraxanthias parvus (Borradaile, 1900) Phymodius nitidus (Dana, 1852) Phymodius ungulatus (H. Milne Edwards, 1834) Pilodius areolatus (H. Milne Edwards, 1834) Pilodius granulatus Stimpson, 1859 Pilodius melanospinis (Rathbun, 1911) Pilodius paumotensis Rathbun, 1907 Pilodius pilumnoides (White, 1848) Pilodius pubescens Dana, 1852 Pilodius pugil Dana, 1852 Pilodius scabriculus Dana, 1852 Pilodius serenei Miyake & Takeda, 1968 UF 770, BPBM UF 383 UF/ZRC UF 398 Lybia tesselata (Latreille, 1812) Lybia sp. 1 Macromedaeus crassimanus (A. Milne Edwards, 1867) Macromedaeus nudipes (A. Milne Edwards, 1867) Medaeus? ornatus Dana, 1852 Neoliomera richtersoides? Sakai, 1969 Neoliomera sp. 1 (pink-red) Neoliomera sp. 2 (orange) Neoliomera sp. 3 (flesh) Neoliomera striata Buitendijk, 1941 New genus (aff. Paramedaeus) sp. 1 (n. sp.) Ng, in prep. Olenothus uogi Ng, 2003 UF 780 UF 881, USNM 154954 UF 36, USNM UF 712 UF 764 USNM 134593 UF/ZRC UF 495 UF 29 UF 2010 UF 420 UF 500, USNM, ZRC UF 42 UF 2096 (holotype), ZRC 2002.171 UF 396 UF 461 UF 3 UF 756 UF 380 UF 416, USNM 154846 USNM Museum Taxon P P N N N 5, 10, 62 5, 10 5, 10 5, 10 K N N 62, 88 5, 10, 88 5, 10, 88 5, 10, 88 88 N 5, 10, 62 P N 62 5, 10 P K K N K N 87 Refs GP273-22 GP463-18 GP851-16 GP786-23 GP847-13 GP457-5 GP836-15 GP751-16 GP832-22 GP576-30 GP402-32 GP829-28 GP829-18 GP829-34 GP538-24 GP858-5 GP467-18 GP851-10 Photo 61 58 59 60 57 Notes Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 G G G G G G, S, M G, S S G, S G G G G, M G, S S S S G G, S, M G G G, M S G G G G G G G Hab Island 1? 106, 110 106 106 105, 106 106, 107 UF 336 UF 223 USNM, SIO UF 264, SIO GP842-13 GP401-4 GP596-24 GP463-23 GP456-19 GP466-21 GP533-8 GP821-2 63 1 1 1 1 1 UF 717 UF 792 UF 763 UF/ZRC UF 721 UF 864 UF 775 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 G G G G S G G G G G, S G G, S M M, U S G, M G M, U G G G M G, S G M G, S G Hab Island 103 Notes ?GP859-24 62 GP766-11 GP534-16 ?GP791-11 Photo USNM 228258 (holotype) Refs UF 865, USNM USNM, UGI 2608 UF/ZRC UF 767, USNM UF 774 UF 776 UF 783 UF 765 Pilodius sp. 1 Pilodius spinipes Heller, 1861 Platypodia anaglypta (Heller, 1861) Platypodia semigranosa (Heller, 1861) Polydectus cupulifer (Latreille, 1812) Psaumis cavipes (Dana, 1852) Pseudoliomera granosimana (A. Milne Edwards, 1865) Pseudoliomera helleri (A. Milne Edwards, 1865) Pseudoliomera lata (Borradaile, 1902) Pseudoliomera remota (Rathbun, 1907) Pseudoliomera ruppellioides (Odhner, 1925) Pseudoliomera speciosa (Dana, 1852) Tweedieia laysani (Rathbun, 1906) Xanthias aff. cherbonnieri Guinot, 1964 Xanthias canaliculatus Rathbun, 1906 Xanthias gilbertensis (Balss, 1938) Xanthias lamarcki (H. Milne Edwards, 1834) Xanthias latifrons (de Man, 1888) Xanthias lividus (Lamarck, 1818) Xanthias punctatus (H. Milne Edwards, 1834) Zosimus aeneus (Linnaeus, 1758) ASCOTHORACIDA: SYNAGOGIDAE Sessilogoga elongata Grygier, 1990 CIRRIPEDIA: ARCHAEOBALANIDAE Conopea folliculus Hiro, 1937 CIRRIPEDIA: BALANIDAE Balanus amphitrite Darwin, 1854 Balanus eburneus Gould, 1841 CIRRIPEDIA: CHTHAMALIDAE Chthamalus malayensis Pilsbry, 1916 Chthamalus proteus Dando & Southward, 1980 N K N 5, 10 P 5, 10 62 62 5, 10 62 N 62 N N N 62 5, 10 N 62 5, 10 K Museum Taxon Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) USNM 237820 (holotype) UF 222 Pseudoeuraphia montgomeryi (Foster, 1990) Nesochthamalus intertextus (Darwin, 1854) CIRRIPEDIA: CRYPTOPHIALIDAE Cryptophialus coronophorus Smyth, 1986 CIRRIPEDIA: PYRGOMATIDAE Cantellius euspinulosum (Broch, 1931) Cantellius pallidus (Broch, 1931) Cantellius septimus (Hiro, 1938) Neotrevathana elongata (Hiro, 1931) Savignium crenatum (Sowerby, 1823) Trevathana dentata (Darwin, 1854) Trevathana orientale (Ren, 1986) Trevathana paulayi Asami & Yamaguchi, 2001 Wanella milleporae (Darwin, 1854) CIRRIPEDIA: SCALPELLIDAE Lithotrya nicobarica Reinhardt, 1850 CIRRIPEDIA: TETRACLITIDAE Tetraclitella divisa (Nilsson-Cantell, 1921) CIRRIPEDIA: NEOVERRUCIDAE Neoverruca brachylepadoformis Newman, 1989 CIRRIPEDIA: PACHYLASMATIDAE Tetrapachylasma ornatum Jones, 2000 CIRRIPEDIA: LITHOGLYPTIDAE Lithoglyptes mitis Tomlinson, 1969 Cryptophialus cf. zulloi Tomlinson, 1973 USNM Euraphia hembeli Conrad, 1837 106 105, 106 SIO USNM NSMT-Cr13681 (holotype) 106 106 106 106 106 106, 111 106 106, 111 106 GP263-2 GP529-20 GP263-6 GP263-9 106, 109 GP768-2 YPM 9303 (holotype, S), SIO, UF 480 Photo 2, 104 106, 113, 114 106, 112, 113 106, 113, 114 106, 107, 108 105, 106 107, 108 Refs USNM USNM 222986 (holotype) Museum Taxon Notes Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) 1 1 1 1 1 1 1 1 1 1 1 1 3 1 1 1 1 1 1 G G G S G G G G G G G G, S MTV G G G G G G Hab Island 1 FW FW FW 115 115 115, 116, 117 115, 116, 117 115 115 115, 117 115 115 2, 118 115, 117 115 115 115 115 120 139 139 USNM USNM USNM 195374 (holotype) USNM 195376 (holotype) USNM USNM USNM USNM USNM 234586 (holotype) USNM 234505 (holotype) USNM USNM USNM USNM USNM Anoplodactylus chamorrus Child, 1983 Anoplodactylus glandulifer Stock, 1954 Anoplodactylus longiceps Stock, 1951 Anoplodactylus pycnosoma (Helfer, 1938) Anoplodactylus tenuicorpus Child, 1991 Eurycyde sertula Child, 1991 Sericosura cochleifovea Child, 1989 Tanystylum rehderi Child, 1970 PYCNOGONIDA: CALLIPALLENIDAE Callipallene novaezealandiae (Thomson, 1884) Seguapallene micronesica Child, 1983 PYCNOGONIDA: ENDEIDIDAE Endeis mollis (Carpenter, 1904) PYCNOGONIDA: NYMPHONIDAE Nymphon diabolis Child, 1988 FRESHWATER TAXA COPEPODA: PHYLLOGNATHOPODIDAE Phylloganthopus viguieri (Maupas, 1892) COPEPODA: CYCLOPIDAE Bryocyclops anninae (Menzel, 1926) Mesocyclops leuckarti (Claus, 1857) 1 1 1 1 1 1 1 1 3 1 1 1 1 1 1 1 1 G G G, Ag G G G G G G G G G MTV G G G G G G G G Hab Island 105, 106 Notes USNM Photo 105, 106 GP420-21 106 USNM Tesseropora pacifica Pilsbry, 1928 Tesseropora sp. 1 CIRRIPEDIA: VERRUCIDAE Verruca cookei Pilsbry, 1928 PYCNOGONIDA: AMMOTHEIDAE Achelia assimilis (Haswell, 1884) Ammothella sp. indet. Ammothella tippula Child, 1983 Refs Museum Taxon Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) Sphaerillo sp. 3 Sphaerillo sp. 4 ISOPODA: LIGIIDAE Ligia sp. 1 (aff. yamanishii) Nunomura, 1990 ISOPODA: ONISCIDAE Oniscidae sp. 1 ISOPODA: PHILOSCIIDAE Burmoniscus bicolor Nunomura, 2001 Philosciidae sp. 1 Setaphora sp. 1 CLADOCERA: SIDIDAE Diaphanosoma sarsi Richard, 1894 s. l. Latonopsis australis Sars, 1888 CLADOCERA: MACROTHRICIDAE Macrothrix laticornis (Jurine, 1820) CLADOCERA: MOINIDAE Moina australiensis Sars, 1896 Moina macrocopa macrocopa (Straus, 1820) Moina weismanni Ishikawa, 1896 TERRESTRIAL TAXA AMPHIPODA: TALITRIDAE Floresorchestia anomala (Chevreux, 1901) Floresorchestia sp. 1 ISOPODA: ARMADILLIDAE Cubaris sp. 1 Hybodillo sp. 1 Sphaerillo sp. 1 Sphaerillo sp. 2 Taxon USNM 141128 USNM 141128 Museum T T T T T 123 122 123 123 T T T T 122 123 123 123 123 T T 121 121 T T FW FW FW 119 119 119 123 123 FW S Ag As, Ag, P, Al, Gu, Sa P Ag S P, An, Gu P M, As, Ag, Sa, An Gu P, An Ag, Al Sa G G G G G G Hab Island 119 Notes FW FW Photo 138 119, 138 Refs Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) T T T T T T T T T 123 123 122 122 123 123 123 123 123 Ag, Al, An, P M, Ag, Gu M, An M Al As M, Al S S M, As M, Ag, Al, Sa Ag References: Literature records listed by numeric code, while identifier of species not previously documented from the Marianas listed by alphabetic code as follows. B) W. Blow, C) T.-Y. Chan, D) W. Dall, F) C. Fransen, G) B. Galil, K) R. Kropp, L) H.-C. Liu, M) P. McLaughlin, N) P. Ng, O) J. Okuno, P) G. Paulay, R) P. Castro, S) J. Starmer, T) M. Takeda. 1) Humes 1990, 2) Hessler & Lonsdale 1991, 3) Weissleader et al. 1989, 4) Komai 1994, 5) Cloud 1959, 6) Just 1998, 7) Williams et al. 2000, 8) Manning et al. 1990, 9) Ahyong & Erdmann 2003, 10) Holthuis 1953, 11) Edmondson 1921, 12) Manning 1978, 13) Kropp & Dominguez 1990, 14) Hamano 1994, 15) Hayashi et al. 1994, 16) Eldredge 1994, 17) Vic Camacho, Univ. of Guam, pers. comm. 2000, 18) Banner 1956, 19) Kropp 1987, 20) Banner & Banner 1985, 21) Leberer & Cai 2003, 22) Martin & Hessler 1990, 23) Kropp 1983[1984], 24) Holthuis 1981, 25) Chace 1997, 26) Camacho 1997, 27) Grimm 1983, 28) Best & Davidson 1981, 29) Chace & Bruce 1993, 30) Bruce & Coombes 1995, 31) King 1987, 32) Wilder 1977, 33) Moffitt 1983, 34) Okuno 1994, 35) Kensley 2003, 36) Kensley 1996, 37) Kensley 2001, 38) Türkay & Sakai 1995, 39) Kensley & Heard 1991, 40) Asakura et al. 1994, 41) Amesbury 1980, 42) Wooster 1979, 43) Wooster 1982, 44) Asakura & Tachikawa 2000, 45) Asakura 1987, 46) Forest 1995, 47) McLaughlin 2002a, 48) Eldredge 1976, 49) McLaughlin 2002b, 50) McLaughlin & Lemaitre 1993, 51) McLaughlin & Haig 1984, 52) McLaughlin & Haig 1989, 53) Dunn et al. 1980, 54) Lemaitre 1998, 55) Williams & Baba 1990, 56) Haig & Kropp 1987, 57) Kropp & Haig 1994, 58) Kropp 1986, 59) Hessler & Martin 1989, 60) McLay 1993, 61) McLay 2001, 62) Takeda et al. 1994, 63) McLay 1999, 64) Eldredge 1980, 65) Guinot & T 123 T Hab Island T Notes 123 Photo Setaphora sp. 3 ISOPODA: PORCELLIONIDAE Porcellionides pruinosus (Brandt, 1833) ISOPODA: SCYPHACIDAE Alloniscus sp. 1 (aff. boninensis) Nunomura, 1984 Alloniscus sp. 2 Alloniscus saipanensis Nunomura, 2001 Alloniscus allspachi Nunomura, 2001 ISOPODA: TRACHELIPIDAE Nagurus sp. 1 Nagurus sp. 2 Nagurus sp. 3 Nagurus sp. 4 ISOPODA: TRICHONISCIDAE Trichoniscidae sp. 1 Refs 123 Museum Setaphora sp. 2 Taxon Appendix: Checklist of the crustaceans and pycnogonids of the Marianas / (continued) Richer de Forges 1995, 66) Ng & Wang 2001, 67) Williams & Moffitt 1991, 68) Galil 1997, 69) Tan & Ng 1995, 70) Guinot & Richer de Forges 1985, 71) Williams & Eldredge 1994, 72) Ng & Takeda 1998, 73) Tan & Ng 2003a, 74) Tan & Ng 2003b, 75) Stevcic et al. 1988, 76) Ng & Takeda 2003, 77) Ng 2002a, 78) Stephenson & Rees 1967, 79) Dickinson 1977, 80) Wee & Ng 1995, 81) Vermeij 1978, 82) Ng & Ng 2003, 83) Ng 2002b, 84) Galil & Takeda 1986, 85) Castro 2003, 86) Takeda 1976, 87) Ng 2002c, 88) Clark & Galil 1993, 89) Kropp 1984, 90) Galil & Clark 1990, 91) Odinetz 1984, 92) Ng & Guinot 2001, 93) Quoy & Gaimard 1824, 94) Ng 2003, 95) Ng et al. 1996, 96) Türkay & Sakai 1995, 97) Kropp 1988a, 98) Kropp 1990, 99) Kropp 1994, 100) Kropp & Manning 1996, 101) Kropp 1989, 102) Kropp 1988b, 103) Grygier 1990, 104) Newman & Hessler 1989, 105) Foster 1990, 106) Paulay & Ross 2003, 107) Southward et al. 1998, 108) Foster & Newman 1987, 109) Jones 2000, 110) Hiro 1937, 111) Asami & Yamaguchi 2001, 112) Smyth 1986, 113) Smyth 1989, 114) Smyth 1990, 115) Child 1991, 116) Child 1983, 117) Stock 1994, 118) Child 1989, 119) Belk 1973, 120) Kikuchi 1994, 121) Morino 1994, 122) Nunomura 2001, 123) Nunomura 1994, 124) Ng 2002d, 125) Ng et al 2000, 126) Crosnier 2002, 127) Edmondson 1959, 128) Koh & Ng 2000, 129) Schubart & Ng 2000, 130) Griffin & Tranter 1986, 131) Ng & Clark 2000, 132) McKenzie 1967, 133) Ng 1998a, 134) Fransen 2002, 135) Moreira & Nelson 1990, 136) Nelson & Kropp 1985, 137) Yamaguchi 1975, 138) Korovchinsky 2001, 139) Watkins & Belk 1975, 140) ElRashidy & Boxshall 2001, 141) Edmondson 1954, 142) Takeda 1980, 143) Crosnier & Ng 2003, 144) Ng & McLay 2003. Notes: 1) as H. poorei Cronin in Weissleader et al. (1989), a nomen nudum (T. M. Cronin pers. comm. 2001) 2) as L. henryi Cronin in Weissleader et al. (1989), a nomen nudum (T. M. Cronin pers. comm. 2001) 3) as O. rossi Cronin in Weissleader et al. (1989), a nomen nudum (T. M. Cronin pers. comm. 2001) 4) as T. weissleaderae Cronin in Weissleader et al. (1989), a nomen nudum (T. M. Cronin pers. comm. 2001) 5) as Gonodactylus f. in Holthuis (1953) and Cloud (1959), see Ahyong & Erdmann (2003) for generic reassignment 6) as G. chiragra var. acutus in Edmondson (1921), see Ahyong & Erdmann. (2003) for reidentification 7) introduced to Guam in fish ponds 8) Original spelling as A. diabilus in Banner (1956) as a result of typographical error, see Chace (1988) 9) as A. edmondsoni in Banner (1956), see Banner & Banner (1983) for synonymy 10) as A. c. inermis, A. c. medius, and A. c. probabilis in Banner (1956), see Banner & Banner (1982) for synonymy 11) as A. tuthilli in Banner (1956), see Banner & Banner (1982) for synonymy 12) as A. gracilis var. simplex in Banner (1956), see Banner & Banner (1982) for synonymy 13) as Thunor i. in Banner (1956), see Banner & Banner (1982) for generic reassignment 14) as A. ventrosus in Banner (1956) and Cloud (1959), see Banner & Banner (1982) for synonymy 15) as A. phyrgianus in Banner (1956) and Cloud (1959), see Banner & Banner (1982) for synonymy 16) as A. p. bengalensis in Banner (1956) and Cloud (1959), see Banner & Banner (1982) for synonymy 17) misspelled as A. pearcyi in Hayashi et al. (1994) 18) as Thunor sp. 1 (n. sp.) in Banner (1956), see Banner & Banner (1982) for generic reassignment 19) as A. cloudi in Banner (1956), see Chace (1988) for synonymy 20) as A. dubius in Banner (1956), see Banner & Banner (1973) for synonymy 21) as A. gardineri in Cloud (1959) and as A. johnsoni in Banner (1956), see Banner & Banner (1973) for synonymy 22) as Alpheus p. in Cloud (1959), see Banner & Banner (1982) for generic reassignment 23) as A. nanus in Banner (1956) and Cloud (1959), see Banner & Banner (1982) for synonymy 24) as A. c. obscurus in Banner (1956), see Banner & Banner (1975) for synonymy 25) as A. coutierei exilipes in Banner (1956), see Banner & Banner (1975) for synonymy 26) as D. n. sp. in Cloud (1959), described in Holthuis (1981) 27) as M. latimanus in Camacho 1997, reidentified by Cai & Ng (unpublished data) 28) as P. vestigialis in Holthuis (1953) and Cloud (1959), see Chace & Bruce (1993) for generic reassignment 29) as Periclimenes s. in Holthuis (1953) and Cloud (1959), see Duris & Bruce (1995) for generic reassignment 30) as Nikoides nanus in Holthuis (1953) and Cloud (1959), see Chace & Bruce (1993) for synonymy 31) This is a third color form of Axiopsis on Guam that likely represents yet another sibling species in the A. serratifrons-complex, so separated by Paulay 32) as Axius a. in Holthuis (1953) and Cloud (1959), see Kensley (2003) for generic reassignment 33) erroneously recorded in Wooster (1979); Wooster’s Aniculus corresponds to A. ursus, checked and corrected by Paulay 34) as A. aniculus in Wooster (1979), checked and corrected by Paulay 35) as Calcinus imperialis in Wooster (1982), see Poupin (1997) for reidentification 36) as Pagurus laevimanus in Holthuis (1953) and Cloud (1959); see McLaughlin & Haig (1984) for generic reassignment. Identification of Cloud’s specimens need to be verified, as Pagurixus is a diverse genus in the Marianas; see comment 40. McLaughlin’s current concept of this species is known on Guam, see photo and specimen vouchers. 37) as Calcinus sp. 1 in Wooster (1982), checked and corrected by Paulay 38) as C. gaimardi in all refs. cited, separated out by Rahayu & Forest (1999); Guam material reidentified by McLaughlin 39) as Orthopagurus harmsi in Wooster (1979), see McLaughlin & Lemaitre (1993) for description of new species 40) as Pagurixus laevimanus in McLaughlin & Haig (1984), reidentified by McLaughlin 41) as Parapagurus d. in Dunn et al. (1980), see Lemaitre (1989) for generic reassignment; associated with the deep water shell-forming anemone Stylobates aeneus 42) as Cryptodormia canaliculata in Takeda et al. (1994), see McLay (1993) for synonymy 43) as Trigonothir o. in Holthuis (1953) and Cloud (1959), see Griffin & Tranter (1986) for synonymy 44) identification of Cloud’s specimens not checked; potentially Tiarinia cornigera 45) as Litocheira aranea in Holthuis (1953), Cloud (1959), generic reassignment by Ng (1987) 46) as Actaea h. in Cloud (1959), see Serène (1984) for generic reassignment 47) likely corresponds to Atergatis subdentatus recorded here 48) may be juvenile of A. granulatus 49) as Actaea s. in Holthuis (1953) and Cloud (1959), see Serène (1984) for generic reassignment 50) as Heteropilumnus i. in Holthuis (1953), Cloud (1959), generic reassignment by Ng (1987) View publication stats 51) as L. a. in Holthuis (1953) and Cloud (1959), subspecific identification by Kropp (unpublished data) 52) as Pilodius h. in Takeda et al. (1994), see Clark & Galil, 1993 for generic reassignment 53) as Carpilodes m. in Holthuis (1953) and Cloud (1959), see Serène (1984) for generic reassignment 54) as Carpilodes s. in Holthuis (1953) and Cloud (1959), see Serène (1984) for generic reassignment 55) as Carpilodes b. in Holthuis (1953) and Cloud (1959), see Serène (1984) for generic reassignment 56) as Carpilodes t. in Holthuis (1953) and Cloud (1959), see Serène (1984) for generic reassignment 57) spelling corrected from P. parrus in Cloud (1959) 58) as Chlorodopsis p. in Holthuis (1953) and Cloud (1959), see Clark & Galil, 1993 for generic reassignment 59) as Chlorodopsis melanodactyla in Holthuis (1953) and Cloud (1959), see Clark & Galil, 1993 for synonymy and generic reassignment 60) as Chlorodopsis p. in Holthuis (1953) and Cloud (1959), see Clark & Galil, 1993 for generic reassignment 61) considered a syn. of P. granulatus by Clark & Galil (1993), but not Ng 62) as Actaea cavipes in Holthuis (1953) and Cloud (1959), see Serène (1984) for generic reassignment 63) as Actaea l. in Holthuis (1953) and Cloud (1959), see Serène (1984) for generic reassignment 64) as Thelphusa r. in Quoy & Gaimard (1924) and as Cardisoma r. in Takeda et al. (1994), see Ng & Guinot (2001) for generic reassignment 65) Takeda et al. (1994) record P. tuberculata, now considered a species complex of P. immaculata and P. squamosa (Ng et al. 2000) 66) The Eumedonidae has been synonymised with the Pilumnidae by Ng & Clark (2002) and now regarded only as a subfamily within it. 67) Ng & Liao (2002) recently recognised the Pseudoziidae as a distinct family and formally transferred Euryozius to this taxon as well. 68) as Saipanella cloudi in McKenzie (1967), generic reassignment by McKenzie (1970) 69) as Penaeus s. in Eldredge (1994), generic reassignment by Pérez Farfante & Kensley (1997) 70) as Thor maldivensis in Cloud (1959) and Holthuis (1953), generic reassignment by Bruce (1997, 1998) 71) describe as Hypsophrys williamsi, generic reassignment by Ng (1998b) 72) This species is common in Pleistocene deposits in Apra Harbor (Kesling 1958), but was never seen by us alive. However a single, unlabeled specimen (UF 2846) of this species was found in the UOGML teaching collection. This specimen likely came from Guam, but additional specimens are needed to ascertain this. The crustacean fauna of Apra harbor is still poorly sampled. 73) as Pagurus latro in Quoy & Gaimard (1824), now in monotypic Birgus (Amesbury 1980)