Marine molluscs from Cyprus: new data and checklist

Bilal Öztürk

This study has been carried out along the North Cypriot coasts on May 10th-19th 1997 and July 12th-21st 1998. As a result of the sampling in the shore region and in deep- water, depths ranging from 0 to 600 m, 304 species, for a total number of 4786 individuals, included in the classes of Polyplacophora, Gastropoda, Bivalvia and Scaphopoda have been identified. Among the 304 taxa examined, 18 species from Cyprus Island are here reported for the first time. In particular, 16 Gastropoda: Nodilittorina (Echinolittori- na) punctata (Gmelin, 1791), Rissoa splendida Eichwald, 1830, Alvania fractospira Oberling, 1970, Alvania testae (Aradas & Maggiore, 1844), Hydrobia acuta (Draparnaud, 1805), Protatlanta souleyeti (Smith E. A., 1888), Cerithiopsis scalaris Locard, 1892, ex Monterosato ms., Nassarius circumcictus (Adams A., 1852), Granulina boucheti Gofas, 1992, Gymno- bela subaraneosa (Dautzenberg & Fischer H., 1896), Teretia teres (Reeve, 1844), Chrysallida juliae (de Folin, 1872), Folin...

Bollettino Malacologico, Roma, 39 (5-8): 49-78, [2003] 2004- Società Italiana di Malacologia Marine molluscs from Cyprus: new data and checklist Bila l Öztürk, Giova nni Buzzurro & H. Avni Benli KEY WORDS: Mollusca, Recent, Checklist, Cyprus, Mediterranean Sea. ABSTRACT This study has been carried out along the North Cypriot coasts on May 10 th-19 th 1997 and July 12 th-21 st 1998. As a result of the sampling in the shore region and in deepwater, depths ranging from 0 to 600 m, 304 species, for a total number of 4786 individuals, included in the classes of Polyplacophora, Gastropoda, Bivalvia and Scaphopoda have been identified. Among the 304 taxa examined, 18 species from Cyprus Island are here reported for the first time. In particular, 16 Gastropoda: Nodilittorina (Echinolittorina) punctata (Gmelin, 1791), Rissoa splendida Eichwald, 1830, Alvania fractospira Oberling, 1970, Alvania testae (Aradas & Maggiore, 1844), Hydrobia acuta (Draparnaud, 1805), Protatlanta souleyeti (Smith E. A., 1888), Cerithiopsis scalaris Locard, 1892, ex Monterosato ms., Nassarius circumcictus (Adams A., 1852), Granulina boucheti Gofas, 1992, Gymnobela subaraneosa (Dautzenberg & Fischer H., 1896), Teretia teres (Reeve, 1844), Chrysallida juliae (de Folin, 1872), Folinella ghisottii Aartsen, 1984, Tragula fenestrata (Jeffreys, 1848), Eulimella bogii Aartsen, 1994, Turbonilla acutissima Monterosato, 1884, and 2 Bivalvia: Limaria loscombi (Sowerby G. B. I, 1823), Kelliella abyssicola (Forbes, 1844). Furthermore, six species are reported for the Eastern basin for the first time. All the species reported for the first time from Cyprus in this paper are figured and discussed. A total amount of 696 species is here reported in a newly formed checklist of Molluscs of Cyprus, considering all the previous works concerned about the malacological fauna occurring in the area under investigation and the additional species we have identified. Furthermore, the list, which is taking into account the many nomenclatural changes recently introduced in the class Mollusca, also deals with alien species. RIASSUNTO Questo studio è stato condotto lungo le coste della parte Nord dell’isola di Cipro tra il 10 e il 19 Maggio del 1997 e il 12 e il 21 Luglio del 1998. Sono stati raccolti campioni fino alla profondità massima di 600 m, individuando 4786 esemplari distribuiti tra 304 specie appartenenti alle classi Poliplacophora, Gastropoda, Bivalvia e Scaphopoda. Di queste 304 specie 18 risultano inedite per Cipro, 16 Gastropoda: Nodilittorina (Echinolittorina ) punctata (Gmelin, 1791), Rissoa splendida Eichwald, 1830, Alvania fractospira Oberling, 1970, Alvania testae (Aradas & Maggiore, 1844), Hydrobia acuta (Draparnaud, 1805), Protatlanta souleyeti (Smith E. A., 1888), Cerithiopsis scalaris Locard, 1892, ex Monterosato ms., Nassarius circumcictus (Adams A., 1852), Granulina boucheti Gofas, 1992, Gymnobela subaraneosa (Dautzenberg & Fischer H., 1896), Teretia teres (Reeve, 1844), Chrysallida juliae (de Folin, 1872), Folinella ghisottii Aartsen, 1984, Tragula fenestrata (Jeffreys, 1848), Eulimella bogii Aartsen, 1994, Turbonilla acutissima Monterosato, 1884 e 2 Bivalvia: Limaria loscombi (Sowerby G. B. I, 1823), Kelliella abyssicola (Forbes, 1844). Tra queste, 6 specie sono inedite per il bacino orientale del Mar Mediterraneo. Di tutte le specie trattate si dà anche una rappresentazione iconografica. Viene presentata inoltre una nuova lista aggiornata comprendente tutte le specie precedentemente segnalate in Letteratura e quelle ulteriormente identificate da noi, per un totale di 696 taxa. Questa lista, corredata da relative note integrative ed in accordo con gli ultimi aggiornamenti tassonomici, evidenzia anche le specie di provenienza indopacifica. B. ÖZTÜRK, Ege University, Faculty of Fisheries, Department of Hydrobiology, 35100 Bornova (Izmir), Turkey, E-mail: Ozturk@ sufak.ege.edu.tr G. BUZZURRO, Via Locatelli 44, Biassono (Mi), Italy, E-mail: giobuz@ libero.it H . A . B EN L I , D okuz Eylül Ü niversit y, I nst it ut e of Marine Science and T echnology, I nciralt ı (I zmir), T urkey, E- mail: benli@ neptune.imst.deu.edu.tr INTRODUCTION The island of Cyprus has special ecological characteristics due to its narrow sublittoral and littoral but wide bathyal zones. This island, with a coastline of 750 km in length, is an important biogeographical spot in the Levantine basin where Red Sea originated Lessepsian migrants establish. There are many studies so far carried out by different scientists on the biological diversity Cyprus’ bottoms. The first report on the Mollusca from the coasts of this island was by MONTEROSATO (1899). Thereafter, many works were carried out especially in the second half of century. Among the others, we cite T SURNAMAL (1969), DEMETROPOULOS (1969, 1971), B ARASH & LEW INSOHN (1971), D EMET R OPOULOS & H ADJ ICHR IST OPHOR OU (1976), ST AVR INOS (1984), T OR NAR IT IS (1987), B AR ASH & D ANIN (1989), B OGI et al . (1989), FISCHER (1993a, b; 1994), CECALUPO & QUADRI (1994, 1995, 1996) and HADJICHRISTOPHOROU et al. (1997). As a result of these studies, 585 Mollusca species have been reported from Cyprus (CECALUPO & QUADRI, 1996). More recently, B UZZURRO & G REPPI (1997) and SALMAN et al . (1998) have reported 42 and 10 additional species, respectively. Several checklists of the molluscan fauna of Cyprus have been published in the past by different Authors (D EMETROPOULOS, 1 9 7 1 ; B OGI et a l ., 1 9 8 9 ; F ISCHER , 1 9 9 4 and CECALUPO & QUADRI, 1996). However, some of these early reports suffered the complicated taxonomy of some groups, and several species were listed as separate taxa in spite of being synonyms according to the latest taxonomy. In this paper we present an up to date checklist of Mollusca species occurring in the Island of Cyprus. W ith this report, the total number of Mollusca occurring along the Cyprus coasts has been enriched with 16 Gastropoda and 2 Bivalvia species. MATERIAL AND METHODS Specimens and shells identified in this study were collected during the expeditions along Northern Cyprus coasts on the dates of May 10 th-19 th, 1997 and July 12 nd-21 st , 1998 by R/V “ K. Piri Reis”. Samples were taken from a depth ranging from 0 to 10 m along the shore region and from 20 to 600 m offshore (Fig.1). Sampling have been randomly selected in the coastal regions having both sandy and rocky habitat and in the substrates consisting of algae (Padina pavonica (L) Thivy, 1960, Cystoseira crinita (Desf.) Bory, 1846, Cystoseira sp., Sargassum vulgare C. Agardh, 1821, Udotea petiolata (Turra) Boerg, 1915, Caulerpa prolifera (Forskkal) Lamaroux, 1809, Caulerpa racemosa (Forskkal) J . Agardh), phanerogam (Posidonia oceanica (L.) Delile, 1813) and sponge (Sarcotragus sp.) (Table 1). Samples were collected directly by hand. A total of 70 samplings were carried out at 12 stations. Furthermore, in the open sea, trawl- > 49< B. Öztürk, G. Buzzurro & H. Benli ing and dredging have been utilized. Samples were sieved with 0.5 mm mesh size. Systematics of the identified species are according to SABELLI et al . (1990-1992). Moreover, species names reported in the past by several Authors have been reconsidered according to SABELLI et al . (1990-1992) during the preparation of this new checklist. Thus, the list was cleaned up of synonym species. The many changes occurred in the nomenclature of Mollusca after 1990 have been also considered. Furthermore, systematic information about some species was adopted by visiting the web-site Check List of European Marine Mollusca (CLEMAM: home page, http://www.mnhn.fr/base/malaco/html). The materials collected were deposited in the Depatment of Hydrobiology, Ege University (Izmir, Turkey). Fig. 1. Map of studied area with locations of shallow water stations. Fig. 1. Cartina dell’area studiata con l’indicazione delle stazioni di acque poco profonde. Table 1. Coordinates, localities, depths, dates and biotop characterizations of the samples. St. Date T1 T2 T3 T4 T5 T7 T9 T10 T11 T12 T13 T14 T17 T18 T19 T20 T21 T22 T23 T24 T25 T26 10.5.97 10.5.97 11.5.97 12.5.97 12.5.97 13.5.97 13.5.97 16.5.97 16.5.97 17.5.97 17.5.97 17.5.97 185.97 18.5.97 18.5.97 19.5.97 19.5.97 14.7.98 17.7.98 19.7.98 19.7.98 19.7.98 BOTTOM-TRAWL Coordinates or Localities Depth (m) Start / Finish 35°21’7”N-33°07’5”E /35°23’9”N-33°09’0”E 560-576 35°22’7”N-33°02’6”E /35°22’7”N-33°04’2”E 33-35 35°20’58”N-33°23’4”E /35°20’7”N-33°22’5”E 40-46 35°20’7”N-33°28’6”E /35°20’6”N-33°27’5”E 28-38 35°22’9”N-33°41’0”E /35°22’4”N-33°39’3”E 38-45 35°19’2”N-32°54’6”E /35°17’7”N-32°54’4”E 28-45 35°12’5”N-32°52’3”E /35°11’2”N-32°51’8”E 75-85 35°40’3”N-34°39’1”E /35°39’7”N-34°37’8”E 275-337 35°40’9”N-34°35’4”E /35°40’6”N-34°35’3”E 27-45 35°26’7”N-34°10’4”E /35°25’0”N-34°08’4”E 32-95 35°25’1”N-34°11’1”E /35°26’3”N-34°11’7”E 37-69 35°21’5”N-34°10’4”E /35°21’9”N-34°08’9”E 187-155 35°13’9”N-33°56’9”E /35°15’2”N-33°57’0”E 100-145 35°13’6”N-33°56’0”E /35°15’0”N-33°55’8”E 62-70 35°12’0”N-33°55’6”E /35°13’3”N-33°55’1”E 37-38 35°11’1”N-33°56’7”E /35°10’3”N-33°55’9”E 50-100 35°11’1”N-33°58’0”E /35°10’0”N-33°57’0”E 66-150 35°19’1”N-32°49’0”E /35°18’6”N-32°49’9”E 223-227 35°47’7”N-34°43’8”E /35°47’9”N-34°44’1”E 113-114 35°06’4”N-34°00’5”E /35°07’6”N-33°59’6”E 90-89 35°09’4”N-33°59’1”E /35°10’4”N-33°58’5”E 105-131 35°11’8”N-33°56’3”E /35°10’6”N-33°57’2”E 59-70 > 50 < Biotopes Mud Hard substratum and Sarcotragus sp. Posidonia oceanica Posidonia oceanica Posidonia oceanica and Caulerpa prolifera Posidonia oceanica Muddy sand with Caulerpa racemosa Mud Posidonia oceanica Posidonia oceanica Posidonia oceanica Mud Hard substratum Muddy sand with Caulerpa racemosa Muddy-sand with Caulerpa racemosa and Udotea petiolata Mud and Sarcotragus sp. Muddy sand with Caulerpa racemosa and Udotea petiolata Mud Mud Mud Mud with shell fragments Mud Marine molluscs from Cyprus: new data and checklist St. Date D1 D2 D3 D4 D7 D8 D9 D10 D11 D13 D14 D15 D16 D17 D18 D19 D20 D21 10.5.97 12.5.97 12.5.97 15.5.97 16.5.97 16.5.97 16.5.97 16.5.97 19.5.97 19.5.97 12.7.98 14.7.98 14.7.98 16.7.98 16.7.98 17.7.98 18.7.98 19.7.98 St. Date C1 C2 C3 C4 C5 C6 C7 C8 C9 14.7.98 14.7.98 18.7.98 18.7.98 19.7.98 19.7.98 19.7.98 21.7.98 21.7.98 C10 21.7.98 C11 21.7.98 C12 13.7.98 DREDJ Depth (m) Biotopes Coordinates or Localities Start / Finish 35°29’1”N-33°58’0”E 35°22’4”N-33°39’9”E 35°20’7”N-33°28’3”E 35°37’8”N-34°21’1”E 35°35’7”N-34°27’4”E 35°34’8”N-34°25’9”E 35°34’4”N-34°26’9”E 35°34’8”N-34°28’3”E 35°10’0”N-33°59’4”E 35°08’7”N-33°57’6”E 35°30’0”N-33°09’9”E 35°10’0”N-32°50’0”E 35°19’9”N-32°50’1”E 35°43’8”N-34°37’3”E 35°33’8”N-34°13’0”E 35°24’8”N-34°10’1”E 35°08’7”N-34°00’1”E 35°09’0”N-33°57’0”E 50 35 35 35 70 35 140 210 25 92 600 69 210 250-300 32 50 120 20 Hard substratum Sandy mud with shell fragments Sandy mud with shell fragments Sand with Brachiostoma lanceolatum and Posidonia oceanica Mud with Ascidia sp. Posidonia oceanica Mud Mud Posidonia oceanica Hard substratum with Sargassum vulgare Mud Sandy mud with shell fragments Mud Mud with shell fragments Posidonia oceanica Muddy sand with Caulerpa racemosa Mud Posidonia oceanica DIVING Depth (m) Biotopes Coordinates or Localities Start / Finish Guzelyurt Bay Guzelyurt Bay Karpas Cape Famagusta Bay Karpas Cape Karpas Cape Karpas Cape Girne Malazgirt 0-10 0-10 0-10 0-10 0-10 0-10 0-10 0-10 0-10 Kormakiti Cape Famagusta Harbour Girne Harbour 0-10 0-3 0-1 Cystoseira crinita, Posidonia oceanica, rocks and sand Cystoseira crinita and sand Cystoseira crinita, Posidonia oceanica and rocks Posidonia oceanica and rocks Cystoseira sp., Posidonia oceanica and rocks Padina pavonica, Posidonia oceanica and rocks Cystoseira sp. and rocks Cystoseira crinita, Posidonia oceanica and rocks Padina pavonica, Cystoseira crinita, Posidonia oceanica and rocks Cystoseira crinita, Posidonia oceanica and sand Rocks with Cystoseira crinita Padina pavonica RESULTS During the study carried out in the Northern Cypriot coasts, 4786 specimens were collected. As a result of evaluation of this sampling, 9 Polyplacophora, 183 Gastropoda, 110 Bivalvia and 2 Scaphopoda species were identified (Table 2). Out of 304 species identified during this study, 18 species are here reported for the first time from Cyprus (Table 3). According to the literature so far analyzed, 6 of these 18 species have never been reported before for the entire Levantine basin, i.e.: Rissoa splendida Eichwald, 1830, Protatlanta souleyeti (Smith E. A., 1888), Cerithiopsis scalaris Locard, 1892, ex Monterosato ms., Granulina boucheti Gofas, 1992, Teretia teres (Reeve, 1844) and Turbonilla acutissima (Monterosato, 1884). Table 2: Taxonomical distribution of the species examined. Classis Polyplacophora Gastropoda Bivalvia Scaphopoda Total Ordo 1 12 10 23 Familia 4 59 38 2 103 Genus 6 109 83 2 200 > 51 < Species 9 183 110 2 304 B. Öztürk, G. Buzzurro & H. Benli Table 3: Molluscs species known from the Cyprus Island (examined material and literature). L: literature (Lx : Authors from the literature); FR: first record for the area; N: notes (* indopacific origin); R: findings from our material; Z : figures. SPECIES POLY PLACOPHORA Leptochitonidae Lepidopleurus cajetanus (Poli, 1791) Lepidopleurus bedullii (Dell’Angelo & Palazzi, 1986) Lepidopleurus cancellatus (Sowerby G. B. II, 1840) Ischnochitonidae Ischnochiton rissoi (Payraudeau, 1826) Callochiton septemvalvis (Montagu, 1803) Lepidochitona cinerea (Linnaeus, 1767) Lepidochitona caprearum (Scacchi, 1836) Lepidochitona monterosatoi Kaas & Van Belle, 1981 Chitonidae Chiton corallinus (Risso, 1826) Chiton olivaceus Spengler, 1797 Chiton phaseolinus Monterosato, 1879 Acanthochitonidae Acanthochitona crinita (Pennant, 1777) Acanthochitona fascicularis (Linnaeus, 1767) GASTROPODA Patellidae Patella caerulea Linnaeus, 1758 Patella rustica Linnaeus, 1758 Patella ulyssiponensis Gmelin, 1791 Lottiidae Tectura virginea (Müller O. F., 1776) Lepetidae Propilidium scabrosum Jeffreys, 1883 Addisoniidae Addisonia lateralis (Réquien, 1848) Neritidae Smaragdia souverbiana (Montrouzier, 1863) Smaragdia viridis (Linnaeus, 1758) Fissurellidae Fissurella nubecula (Linnaeus, 1758) Diodora dorsata (Monterosato, 1878) Diodora gibberula (Lamarck, 1822) Diodora graeca (Linnaeus, 1758) Diodora italica (Defrance, 1820) Emarginula octaviana Coen, 1939 Emarginula sicula Gray, 1825 Emarginella huzardii (Payraudeau, 1826) Scissurellidae Scissurella costata D’Orbigny, 1824 Anatoma crispata Fleming, 1828 Sinezona cingulata (Costa O. G., 1861) Haliotidae Haliotis tuberculata tuberculata Linnaeus, 1758 Trochidae Trochus erythraeus Brocchi, 1826 Clanculus (Clanculus) corallinus (Gmelin, 1791) Clanculus (Clanculopsis) cruciatus (Linnaeus, 1758) L N L3 L23 L3 1 R + L5 L2 L23 L5 L11 + + + + + L5 L2 L23 + + L11 L2 2 3 + L1 L1 L9 + + + L2 L4 L9 L24 L2 * + L4 L23 L4 L4 L2 L2 L2 L9 + + + + L11 L24 L11 + + L1 4 + L9 L2 L3 * + + + > 52 < Z Marine molluscs from Cyprus: new data and checklist Table 3 continued SPECIES Clanculus (Clanculopsis) jussieui (Payraudeau, 1826) Calliostoma conulus (Linnaeus, 1758) Calliostoma laugieri laugieri (Payraudeau, 1826) Calliostoma zizyphinum (Linnaeus, 1758) Calliostoma granulatum (Von Born, 1778) Gibbula ardens (Von Salis, 1793) Gibbula magus (Linnaeus, 1758) Gibbula adansonii (Payraudeau, 1826) Gibbula turbinoides (Deshayes, 1835) Gibbula fanulum (Gmelin, 1791) Gibbula guttadauri (Philippi, 1836) Gibbula leucophaea (Philippi, 1836) Gibbula philberti (Recluz, 1843) Gibbula richardi (Payraudeau, 1826) Gibbula varia (Linnaeus, 1758) Gibbula divaricata (Linnaeus, 1758) Gibbula rarilineata (Michaud, 1829) Gibbula umbilicaris nebulosa (Philippi, 1848) Osilinus articulatus Lamarck, 1822 Osilinus mutabilis (Philippi, 1846) Osilinus turbinatus (Von Born, 1778) Clelandella miliaris (Brocchi, 1814) J ujubinus exasperatus (Pennant, 1777) J ujubinus striatus striatus (Linnaeus, 1758) Colloniidae Cantrainea peloritana (Cantraine, 1835) Homalopoma carmelae Oliverio & Buzzurro, 1994 Homalopoma sanguineum (Linnaeus, 1758) Tricoliidae Tricolia pullus pullus (Linnaeus, 1758) Tricolia speciosa (Von Muehlfeldt, 1824) Tricolia tenuis (Michaud, 1829) Turbinidae Bolma rugosa (Linnaeus, 1758) Cerithiidae Cerithium alucaster (Brocchi, 1814) Cerithium haustellum Monterosato in Crema, 1903 Cerithium lividulum Risso, 1826 Cerithium nesioticum Pilsbry & Vanatta, 1906 Cerithium protractum Bivona Ant. in Bivona And., 1838 Cerithium rupestre Risso, 1826 Cerithium scabridum Philippi, 1848 Cerithium vulgatum Bruguière, 1792 Bittium jadertinum (Brusina, 1865) Bittium latreillii (Payraudeau, 1826) Bittium reticulatum (Da Costa, 1778) Bittium scabrum (Olivi, 1792) Rhinoclavis kochi (Philippi, 1848) Fossaridae Fossarus ambiguus (Linnaeus, 1758) Obtortionidae Finella pupoides Adams, A., 1860 Clathrofenella fusca (Adams, A., 1860) Potamididae L L4 L9 L2 L2 L2 L1 L3 L4 L9 L4 L9 L9 L11 L5 L2 L4 L9 L1 L1 L10 L1 L24 L2 L3 N R + + + + + + + + + + + 5 + + + + + L9 L19 L2 + L2 L2 L9 + L7 + L9 L24 L24 L11 L24 L1 L15 L1 L11 L9 L2 L23 L4 + * * + + + + + + + * L11 L23 L21 > 53 < * * + + Z B. Öztürk, G. Buzzurro & H. Benli Table 3 continued SPECIES Pirenella conica (Blainville, 1826) Siliquariidae Tenagodus obtusus (Schumacher, 1817) Turritellidae Turritella communis Risso, 1826 Turritella turbona Monterosato, 1877 Littorinidae Melarhaphe neritoides (Linnaeus, 1758) Nodilittorina (Echinolittorina) punctata (Gmelin, 1791) Cingulopsidae Eatonina fulgida (Adams, J., 1797) Rissoidae Rissoa angustior (Monterosato, 1917) Rissoa auriformis pseudomonodonta Verduin, 1983 Rissoa auriscalpium (Linnaeus, 1758) Rissoa decorata Philippi, 1846 Rissoa guerinii Récluz, 1843 Rissoa italiensis Verduin, 1985 Rissoa labiosa (Montagu, 1803) Rissoa lia (Monterosato, 1884, ex Benoit ms.) Rissoa monodonta Philippi, 1836 Rissoa rodhensis Verduin, 1985 Rissoa scurra (Monterosato, 1917) Rissoa similis Scacchi, 1836 Rissoa splendida Eichwald, 1830 Rissoa variabilis (Von Muehlfeldt, 1824) Rissoa ventricosa Desmarest, 1814 Rissoa violacea violacea Récluz, 1843 Alvania amatii Oliverio, 1986 Alvania aspera (Philippi, 1844) Alvania beani (Hanley in Thorpe, 1844) Alvania beniamina (Monterosato, 1884) Alvania cancellata (Da Costa, 1778) Alvania chiarellii Cecalupo & Quadri, 1995 Alvania cimex (Linnaeus, 1758) Alvania colossophilus Oberling, 1970 Alvania datchaensis Amati & Oliverio, 1987 Alvania discors (Allan, 1818) Alvania dorbignyi (Audouin, 1827) Alvania fractospira Oberling, 1970 Alvania geryonia (Nardo, 1847, ex Chiereghini ms.) Alvania cf. hallgassi Amati & Oliverio, 1985 Alvania hispidula (Monterosato, 1884) Alvania lactea (Michaud, 1832) Alvania lanciae (Calcara, 1841) Alvania lineata Risso, 1826 Alvania mamillata Risso, 1826 Alvania oranica (Pallary, 1900) Alvania paupercula (Jeffreys, 1867) Alvania rudis (Philippi, 1844) Alvania scabra (Philippi, 1844) Alvania sculptilis (Monterosato, 1877) Alvania testae (Aradas & Maggiore, 1844) Cingula trifasciata (Adams J., 1800) L L9 N R + Z L3 L2 L2 L1 FR + 6 + + 2 L11 L23 L12 L9 L23 L9 L23 L23 L9 L9 L23 L11 L9 FR L9 L2 L9 L11 L9 L9 L6 L18 L21 L1 L11 L12 L9 L11 FR L11 L11 L24 L18 L9 L9 L8 L11 L9 L23 L23 L23 FR L9 > 54 < + 7 + + + + + + + + + 3 + + + 8 + + 4 25 + + + 5 Marine molluscs from Cyprus: new data and checklist Table 3 continued SPECIES Manzonia crassa (Kanmacher, 1798) Manzonia weinkauffi jacobusi Oliverio, Amati & Nofroni, 1986 Obtusella macilenta (Monterosato, 1880) Peringiella elegans (Locard, 1892) Pusillina diversa (Nordsieck, 1972) Pusillina inconspicua (Alder, 1844) Pusillina lineolata (Michaud, 1832) Pusillina margiminia (Nordsieck, 1972) Pusillina marginata (Michaud, 1832) Pusillina munda (Monterosato, 1884) Pusillina philippi (Aradas & Maggiore, 1844) Pusillina radiata (Philippi, 1836) Setia ambigua (Brugnone, 1873) Setia fusca (Philippi, 1841) Setia slikorum (Verduin, 1984) Setia turriculata Monterosato, 1884 Rissoina bruguieri (Payraudeau, 1826) Rissoina bertholleti Issel, 1869 Adeorbidae Circulus striatus (Philippi, 1836) Anabathridae Pisinna glabrata (Von Muhelfeldt, 1824) Assimineidae Paludinella littorina (Delle Chiaje, 1828) Paludinella cf. sicana (Brugnone, 1876) Barleeidae Barleeia unifasciata (Montagu, 1803) Caecidae Caecum auriculatum de Folin, 1868 Caecum trachea (Montagu, 1803) Hydrobiidae Hydrobia acuta (Draparnaud, 1805) Tornidae Tornus subcarinatus (Montagu, 1803) Truncatellidae Truncatella subcylindrica (Linnaeus, 1767) Strombidae Strombus persicus Swainson, 1821 Aporrhaiidae Aporrhais pespelecani (Linnaeus, 1767) Crepidulidae Crepidula unguiformis Lamarck, 1822 Capulidae Capulus hungaricus (Linnaeus, 1758) Vermetidae Vermetus cristatus Biondi, 1857 Vermetus rugulosus Monterosato 1878 Vermetus triquetrus Bivona Ant., 1832 Vermetus cf. granulatus (Gravenhorst, 1831) Vermetus cf. semisurrectus Bivona Ant., 1832 Vermetus sp. Dendropoma anguliferum (Monterosato, 1884) Dendropoma petraeum (Monterosato, 1884) Petaloconchus glomeratus (Linnaeus, 1758) L L11 L23 L23 L23 L23 L23 L9 L23 L23 L23 L23 L11 L23 L23 L23 L11 L9 L11 N R + Z + 9 + * L9 L11 L11 L23 L9 L11 L11 + + FR + L9 + L2 + L9 * L2 + + 10 L2 L2 L9 L23 L2 L24 L24 L23 L11 L2 L7 > 55 < + 6 B. Öztürk, G. Buzzurro & H. Benli Table 3 continued SPECIES Serpulorbis arenaria (Linnaeus, 1767) Cypraeidae Erosaria spurca (Linnaeus, 1758) Luria lurida (Linnaeus, 1758) Lamellariidae Lamellaria latens (Müller O. F., 1776) Lamellaria perspicua (Linnaeus, 1758) Triviidae Trivia arctica (Pulteney, 1789) Trivia levantina Smriglio, Mariottini & Buzzurro, 1998 Trivia pulex (Solander in Gray, 1828) Naticidae Natica dillwynii Payraudeau, 1826 Natica hebraea (Martyn, 1784) Natica stercusmuscarum (Gmelin, 1791) Tecnonatica filosa (Philippi, 1845) Tecnonatica rizzae (Philippi, 1844) Euspira guillemini (Payraudeau, 1826) Euspira pulchella (Risso, 1826) Neverita josephinia Risso, 1826 Payraudeautia intricata (Donovan, 1804) Tonnidae Tonna galea (Linnaeus, 1758) Cassidae Galeodea echinophora (Linnaeus 1758) Galeodea rugosa (Linné, 1771) Phalium granulatum (Von Born, 1778) Phalium saburon (Bruguière, 1792) Ranellidae Ranella olearia (Linnaeus, 1758) Cymatium corrugatum corrugatum (Lamarck, 1816) Cymatium parthenopeum parthenopeum (Von Salis, 1793) Cabestana cutacea cutacea (Linnaeus, 1767) Charonia lampas lampas (Linnaeus, 1758) Charonia variegata (Lamarck, 1816) Atlantidae Atlanta peronii Lesueur, 1817 Oxygyrus keraudrenii (Lesueur, 1817) Protatlanta souleyeti (Smith E. A., 1888) Firolidae Firoloida desmarestia Lesueur, 1817 Pterotrachea hippocampus Philippi, 1836 Pterotrachea minuta Bonnevie, 1920 Triphoridae Cheirodonta pallescens (Jeffreys, 1867) Marshallora adversa (Montagu 1803) Monophorus perversus (Linnaeus, 1758) Similiphora similior (Bouchet & Guillemot, 1978) Metaxia bacillum (Issel, 1869) Metaxia metaxa (Delle Chiaje, 1828) Cerithiopsidae Cerithiopsis diadema Monterosato, 1864, ex W atson ms. Cerithiopsis fayalensis W atson, 1886 Cerithiopsis minima (Brusina, 1865) L L1 N R + Z 11 L2 L2 + L9 L3 L1 L27 L2 + L1 L7 L7 L18 L9 L18 L7 L7 L9 + + + L2 + L2 L3 L3 L2 + L9 L2 L9 L9 L3 L2 + + + 12 L24 L24 FR + + L13 L13 L13 L23 L15 L2 L23 L21 L13 L24 L23 L23 > 56 < + + * + 7 Marine molluscs from Cyprus: new data and checklist Table 3 continued SPECIES Cerithiopsis nana Jeffreys, 1867 Cerithiopsis pulvis (Issel, 1869) Cerithiopsis scalaris Locard, 1892, ex Monterosato ms. Cerithiopsis tenthrenois (Melvill, 1869) Cerithiopsis tubercularis (Montagu, 1803) Krachia tiara Monterosato, 1874, ex W atson ms. Dizoniopsis bilineata (Hörnes, 1848) Dizoniopsis coppolae (Aradas, 1870) Seila trilineata (Philippi, 1836) Janthinidae J anthina janthina (Linnaeus, 1758) J anthina nitens Menke, 1828 Cimidae Cima cylindrica (Jeffreys, 1858) Epitoniidae Epitonium algerianum (W einkauff, 1866) Epitonium clathratulum (Kanmacher, 1798) Epitonium commune (Lamarck, 1822) Epitonium turtoni (Turton, 1819) Cycloscala hyalina (Sowerby G. B. II, 1844) Gyroscala lamellosa (Lamarck, 1822) Opalia hellenica (Forbes, 1844) Eulimidae Crinophtheiros comatulicola (Graff, 1875) Ersilia mediterranea (Monterosato, 1869) Eulima bilineata Alder, 1848 Eulima glabra (Da Costa, 1778) Melanella boscii (Payraudeau, 1827) Melanella monterosatoi (Monterosato, 1890, ex De Boury ms.) Melanella polita Linnaeus, 1758) Parvioris ibizenca (Nordsieck, 1968) Nanobalcis nana (Monterosato, 1878) Sabinella piriformis Brugnone, 1873 Sticteulima lentiginosa (Adams A., 1861) Sticteulima jeffreysiana (Brusina, 1869) Vitreolina curva (Monterosato, 1874, ex Jeffreys ms.) Vitreolina devians Monterosato, 1875 Vitreolina incurva (B. D. D., 1883) Vitreolina levantina Oliverio, Buzzurro & Villa, 1994 Vitreolina perminima (Jeffreys, 1883) Vitreolina philippi (Rayneval & Ponzi, 1854) Muricidae Aspella anceps (Lamarck, 1822) Bolinus brandaris (Linnaeus, 1758) Dermomurex scalaroides (Blainville, 1829) Hadriania oretea (De Gregorio, 1885) Hexaplex trunculus (Linnaeus, 1758) Murexsul aradasii (Poirier, 1883, ex Monterosato ms.) ? Muricopsis cevikeri Houart, 2000 Muricopsis cristata (Brocchi, 1814) Ocenebra erinaceus (Linnaeus, 1758) Ocinebrina aciculata (Lamarck, 1822) Ocinebrina edwardsii (Payraudeau, 1826) Ocinebrina hispidula (Pallary, 1904) L L23 L11 FR L24 L11 L23 L11 L23 L9 N R Z + 8 * * + L9 L2 L11 L23 L3 L9 L4 L17 L2 L23 L11 L11 L25 L2 L23 L23 L11 L23 L24 L24 L24 L23 L9 L24 L11 L20 L9 L9 L4 L2 L23 L4 L2 L9 L28 L2 L4 L3 L4 L23 > 57 < + * + + + * + + * + + + + + + B. Öztürk, G. Buzzurro & H. Benli Table 3 continued SPECIES Ocinebrina hybrida (Aradas & Benoit, 1876) Typhinellus sowerbyi (Broderip, 1833) Ergalatax obscura Houart, 1996 Thais lacera (Born, 1778) Stramonita haemastoma (Linnaeus, 1766) Coralliophila meyendorffii (Calcara, 1845) Hirtomurex squamosa (Bivona And., 1838) Babelomurex babelis (Requién, 1848) Buccinidae Buccinulum corneum (Linnaeus, 1758) Chauvetia candidissima (Philippi, 1836) Chauvetia recondita (Brugnone, 1873) Engina leucozona (Philippi, 1843) Pisania striata (Gmelin, 1791) Pollia dorbignyi (Payraudeau, 1826) Pollia scabra Locard, 1886 Pollia scacchiana (Philippi, 1844) Colubraria reticulata (Blainville, 1826) Columbellidae Columbella rustica (Linnaeus, 1758) Anachis savignyi (Moazzo, 1939, ex Jousseaume ms.) Anachis troglodytes (Sowerbie G. B. I & Montrouzier, 1866) Mitrella cf. bruggeni Aartsen, Menkhorst & Gittenberg, 1984 Mitrella gervillii (Payraudeau, 1826) Mitrella minor (Scacchi, 1836) Mitrella scripta (Linnaeus, 1758) Nassariidae Nassarius corniculus (Olivi, 1792) Nassarius incrassatus (Stroem, 1768) Nassarius reticulatus (Linnaeus, 1758) Nassarius circumcictus (Adams A., 1852) Nassarius gibbosulus (Linnaeus, 1758) Nassarius mutabilis (Linnaeus, 1758) Nassarius cuvierii (Payraudeau, 1826) Nassarius louisi (Pallary, 1912) Nassarius lima (Dillwyn, 1817) Cyclope neritea (Linnaeus, 1758) Fasciolariidae Fasciolaria lignaria (Linnaeus, 1758) Fusinus sanctaeluciae (Von Salis, 1793) Fusinus rudis (Philippi, 1844) Fusinus syracusanus (Linnaeus, 1758) Costellariidae Vexillum ebenus (Lamarck, 1811) Vexillum littorale (Forbes, 1844) Vexillum savignyi (Payraudeau, 1826) Vexillum tricolor (Gmelin, 1790) Marginellidae Volvarina mitrella (Risso, 1826) Cystiscidae Gibberula miliaria (Linnaeus, 1758) Gibberula philippii (Monterosato, 1878) Granulina boucheti Gofas, 1992 Granulina marginata (Bivona, 1832) L L11 L4 L24 L28 L2 L4 L9 L3 N Z * * + L2 L18 L23 L4 L2 L7 L9 L9 L3 L1 L24 L23 L23 L9 L11 L2 R + + + + + * * + + + + L2 L7 L2 FR L2 L2 L9 L9 L9 L2 + + + + + + 9 + L2 L2 L9 L2 + + + 13 L2 L9 L9 L3 + + L4 + L2 L11 FR L23 + + + > 58 < + 14 10 Marine molluscs from Cyprus: new data and checklist Table 3 continued SPECIES L Granulina occulta (Monterosato, 1869) L11 Mitridae Mitra cornicula (Linnaeus, 1758) L2 Mitra zonata Marryat, 1818 L9 Cancellariidae Cancellaria cancellata (Linnaeus, 1767) L4 Turridae Haedropleura septangularis (Montagu, 1803) L18 Conidae Conus mediterraneus Hwass in Bruguière, 1792 L1 Bela cf. cycladensis (Reeve, 1845) L24 Bela laevigata (Philippi, 1836) L15 Bela menkhorsti Aartsen, 1988 L15 Bela nebula (Montagu, 1803) L3 Bela ornata (Locard, 1897) L23 Bela taprurensis (Pallary, 1904) L11 Clathromangelia granum (Philippi, 1844) L9 Gymnobela subaraneosa (Dautzenberg & Fischer H. , 1896) FR Mangelia attenuata (Montagu, 1803) L3 Mangelia barashi (Aartsen & Fehr de W al, 1978) L11 Mangelia bertrandii (Payraudeau, 1826) L11 Mangelia costulata (Blainville, 1829) L3 Mangelia fieldeni (Aartsen & Fehr de W al, 1978, ex Mont. ms.) L11 Mangelia cf. pallary (Nordsieck, 1977) L11 Mangelia sandrii (Brusina, 1865) L23 Mangelia secreta (Aartsen & Fehr de W al, 1978, ex Mont. ms.) L23 Mangelia sicula Reeve, 1846 L23 Mangelia taeniata (Deshayes, 1835) L3 Mangelia unifasciata (Deshayes, 1835) L7 Mangelia vauquelini (Payraudeau, 1826) L9 Taranis moerchi (Malm, 1863) L24 Mitrolumna olivoidea (Cantraine, 1835) L3 Raphitoma alternans (Monterosato, 1884) L24 Raphitoma cf. atropurpurea (Locard & Caziot, 1900, ex Mont. ms.) L24 Raphitoma bicolor (Risso, 1826) L7 Raphitoma concinna (Scacchi, 1836) L9 Raphitoma cordieri (Payraudeau, 1826) L24 Raphitoma echinata (Brocchi, 1814) L9 Raphitoma laviae (Philippi, 1844) L23 Raphitoma linearis (Montagu, 1803) L9 Raphitoma papillosa (Pallary, 1904) L24 Raphitoma philberti (Michaud, 1829) L11 Raphitoma pruinosa (Pallary, 1906) L24 Raphitoma pupoides (Monterosato, 1884) L23 Leufroya leufroyi (Michaud, 1828) L11 Comarmondia gracilis (Montagu, 1803) L13 Teretia teres (Reeve, 1844) FR Drilliidae Crassopleura incrassata (Dujardin, 1837) L9 Mathildidae Mathilda gemmulata Semper, 1865 L9 Rissoellidae Rissoella inflata Locard, 1892 L11 Omalogyridae > 59 < N R Z + + + + + 15 + + + 16 11 + + + + + + + + + + 12 B. Öztürk, G. Buzzurro & H. Benli Table 3 continued SPECIES Omalogyra atomus (Philippi, 1841) Ammonicera fischeriana (Monterosato, 1869) Pyramidellidae Chrysallida clathrata (Jeffreys, 1848) Chrysallida decorata (Philippi, 1849) Chrysallida decussata (Montagu, 1803) Chrysallida emaciata (Brusina, 1866) Chrysallida indistincta (Montagu, 1808 ) Chrysallida intermixta (Monterosato, 1884) Chrysallida jeffreysiana (Monterosato, 1884, ex Seguenza G. ms.) Chrysallida juliae (de Folin, 1872) Chrysallida limitum (Brusina, 1876) Chrysallida obtusa (Brown T., 1827) Chrysallida sp. B Chrysallida suturalis (Philippi, 1844) Chrysallida terebellum (Philippi, 1844) Clathrella clathrata (Philippi, 1844) Euparthenia bulinea (Lowe, 1841) Euparthenia humboldti (Risso, 1826) Folinella excavata (Philippi, 1836) Folinella ghisottii Aartsen, 1984 Odostomella doliolum (Philippi, 1844) Tragula fenestrata (Jeffreys, 1848) Adelactaeon amoenus (Adams A., 1851) Eulimella acicula (Philippi, 1836) Eulimella bogii Aartsen, 1984 Eulimella cerullii (Cossmann, 1915) Eulimella ventricosa (Forbes, 1844) “Syrnola” fasciata Jickeli, 1882 Odostomia carrozzai Aartsen, 1987 Odostomia erjaveciana Brusina, 1869 Odostomia imponderabilior Oberling, 1970 Odostomia lukisii Jeffreys, 1859 Odostomia nofronii Buzzurro, 2002 Odostomia plicata (Montagu, 1803) Odostomia scalaris Mac Gillivray, 1843 Odostomia striolata Forbes & Hanley, 1850 Odostomia turriculata Monterosato, 1869 Odostomia turrita Hanley, 1844 Odostomia unidentata (Montagu, 1803) Megastomia conoidea (Brocchi, 1814) Megastomia sicula Philippi, 1851 Liostomia clavula (Lovén, 1846) Ondina crystallina Locard, 1892 Ondina vitrea (Brusina, 1866) Ondina warreni (Thompson, 1845) Turbonilla acuta (Donovan, 1804) Turbonilla acutissima Monterosato, 1884 Turbonilla delicata Monterosato, 1874 Turbonilla edgari (Melvill, 1896) Turbonilla gradata B. D. D., 1883 Turbonilla lactea (Linnaeus, 1758) Turbonilla pumila Seguenza, G., 1876 Turbonilla pusilla (Philippi, 1844) L L11 L11 L9 L24 L24 L24 L11 L11 L11 FR L23 L11 L14 L23 L14 L18 L9 L11 L9 FR L9 FR L23 L23 FR L15 L24 L22 L11 L23 L23 L11 L29 L11 L23 L23 L23 L11 L11 L9 L23 L11 L23 L23 L11 L24 FR L10 L23 L23 L2 L23 L10 > 60 < N R Z + * 17 + + 13 + + + + 14 + 15 + + 16 * * + + 18 18 + + + * 17 Marine molluscs from Cyprus: new data and checklist Table 3 continued SPECIES Turbonilla cf. rosewateri Corgan & Aartsen, 1993 Turbonilla rufa (Philippi, 1836) Turbonilla sinuosa (Jeffreys, 1884) Turbonilla striatula (Linnaeus, 1758) Turbonilla cf. syrtensis Aartsen, 1981 Anisocyclidae Anisocycla folini (Fischer P. in de Folin, 1869) Anisocycla nitidissima (Montagu, 1803) Anisocycla pointeli (de Folin, 1867) Acteonidae Acteon tornatilis (Linnaeus, 1758) Crenilabium exile (Jeffreys, 1870, ex Forbes ms.) Retusidae Pyrunculus fourierii (Audouin, 1826) Retusa leptoeneilema (Brusina, 1866) Retusa mammillata (Philippi, 1836) Retusa truncatula (Bruguière, 1792) Ventomnestia girardi (Audouin, 1826) Cylichnina crebrisculpta Monterosato, 1884 Cylichnina laevisculpta (Granata-Grillo, 1877) Cylichnina multiquadrata (Oberling, 1970) Cylichnina umbilicata (Montagu, 1803) Volvulella acuminata (Bruguière, 1792) Ringiculidae Ringicula auriculata (Ménard de la Groye, 1811) Ringicula conformis Monterosato, 1877 Bullidae Bulla striata Bruguière, 1792 Haminoeidae Haminoea hydatis (Linnaeus, 1758) Haminoea navicula (Da Costa, 1778) Haminoea orbignyana (Férrussac, 1822) Atys jeffreysi (W einkauff, 1868) Weinkauffia turgidula (Forbes, 1844) Smaragdinellidae Smaragdinella algirae (Adams A. in Sowerby G. B. II, 1850) Philinidae Philine aperta (Linnaeus, 1767) Philine catena (Montagu, 1803) Philine punctata (Adams J., 1800) Aglajidae Philinopsis depicta (Renier, 1804) Akeridae Akera bullata Müller O. F., 1776 Cylichnidae Cylichna crossei B. D. D., 1886 Cylichna cylindracea (Pennant, 1777) Acteocina mucronata (Philippi, 1849) Roxania utriculus (Brocchi, 1814) Scaphander lignarius (Linnaeus, 1758) Gastropteridae Gastropteron meckeli Kosse, 1813 Cavolinidae Cavolinia gibbosa gibbosa (D’ Orbigny, 1835, ex Rang ms.) L L11 L9 L15 L10 L11 N L11 L24 L11 19 L2 L18 L23 L11 L15 L3 L23 L23 L23 L24 L2 L9 R + + + 20 * + * 21 L9 L4 + L2 + L2 L9 L11 L23 L11 + + L23 L2 L11 L10 + L10 L4 L10 L9 L17 L10 L25 L3 L13 > 61 < + * + Z B. Öztürk, G. Buzzurro & H. Benli Table 3 continued SPECIES Cavolinia tridentata (Niebuhr, 1775, ex Forskâl ms.) Clio cuspidata (Bosc, 1802) Clio pyramidata lanceolata (Leseur, 1813) Creseis acicula Rang, 1828 Hyalocylis striata (Rang, 1828) Styliola subula (Quoy & Gaimard, 1827) Limacinidae Limacina trochiformis (D’ Orbigny, 1836) Limacina inflata (D’ Orbigny, 1836) Oxynoidae Oxynoe olivacea Rafinesque, 1814 Lobiger serradifalci (Calcara, 1840) Volvatellidae Ascobulla fragilis (Jeffreys, 1856) Elyssidae Elysia timida (Risso, 1818) Thuridilla hopei (Vérany, 1853) Umbraculidae Umbraculum umbraculum (Röding, 1798) Tylodinidae Tylodina perversa (Gmelin, 1791) Pleurobranchidae Pleurobranchus membranaceus (Montagu, 1815) Berthella ocellata (Delle Chiaje, 1830) Berthellina citrina (Rueppell & Leuckart, 1830) Berthellina engeli Gardiner, 1936 Pleurobranchaea meckelii Meckel in Leue, 1813 Pleurobranchus testudinarius Cantraine, 1835 Aplysidae Aplysia depilans Gmelin, 1791 Aplysia parvula Guilding in Mörch, 1863 Aplysia punctata (Cuvier, 1803) Aplysia fasciata Poiret, 1789 Dolabriferidae Petalifera gravieri (Vayssière, 1906) Notarchus punctatus Philippi, 1836 Aegiretidae Aegires punctilucens (D’ Orbigny, 1837) Triophidae Kaloplocamus ramosus (Cantraine, 1835) Chromodorididae Hypselodoris tricolor (Cantraine, 1835) Hypselodoris villafranca (Risso, 1818) Discodorididae Discodoris atromaculata (Bergh, 1880) Dendrodorididae Dendrodoris grandiflora (Rapp, 1827) Doriopsilla areolata Bergh, 1880 Tethyidae Tethys fimbria Linnaeus, 1767 Zephyrinidae J anolus cristatus (Delle Chiaje, 1841) Flabellinidae L L9 L9 L9 L13 L13 L10 L11 L11 N R + + L4 L25 L24 L10 L4 L2 + L23 L9 L10 L11 L10 L2 L10 L2 L10 L11 L4 L23 L10 L10 L10 L4 L4 L2 L4 L10 L10 L4 > 62 < + Z Marine molluscs from Cyprus: new data and checklist Table 3 continued SPECIES Flabellina affinis (Gmelin, 1791) Siphonariidae Siphonaria pectinata (Linnaeus, 1758) Williamia gussonii (Costa O. G., 1829) Trimusculidae Trimusculus mammillaris (Linnaeus, 1758) Melampidae Auriculinella erosa (Jeffreys, 1830) Ovatella firminii (Payraudeau, 1826) Ovatella denticulata (Montagu, 1803) Ovatella myosotis (Draparnaud, 1801) BIVALVIA Solemyidae Solemya togata (Poli, 1791) Nuculidae Nucula nitidosa W inckworth, 1931 Nucula nucleus (Linnaeus, 1758) Nucula sulcata Bronn, 1831 Nuculoma tenuis (Montagu, 1808) Nuculanidae Lembulus pellus (Linnaeus, 1758) Saccella commutata (Philippi, 1844) Arcidae Arca noae Linnaeus, 1758 Arca tetragona Poli, 1795 Barbatia barbata (Linnaeus, 1758) Acar plicata (Dillwyn, 1817) Anadara corbuloides (Monterosato, 1878) Anadara polii (Mayer, 1868) Bathyarca pectunculoides (Scacchi, 1834) Striarca lactea (Linnaeus, 1758) Glycymerididae Glycymeris bimaculata (Poli, 1795) Glycymeris glycymeris (Linnaeus, 1758) Glycymeris insubrica (Brocchi, 1814) Mytilidae Mytilus galloprovincialis Lamarck, 1819 Brachidontes pharaonis (Fischer P., 1870) Mytilaster minimus (Poli, 1795) Crenella arenaria Monterosato, 1875, ex Martin H. ms. Gregariella petagnae (Scacchi, 1832) Modiolarca subpicta (Cantraine, 1835) Musculus costulatus (Risso, 1826) Lithophaga lithophaga (Linnaeus, 1758) Modiolus barbatus (Linnaeus, 1758) Modiolus adriaticus (Lamarck, 1819) Modiolula phaseolina (Philippi, 1844) Pinnidae Pinna nobilis Linnaeus, 1758 Pteriidae Pteria hirundo (Linnaeus, 1758) Pinctada radiata (Leach, 1814) Malleidae Malleus regula (Forskâl, 1775) L L4 N L9 L11 R + L9 L18 L11 L23 L2 + L9 + L11 L3 L4 L11 + + + L3 L3 + + L1 L9 L1 L18 L9 L3 L24 L3 + + L9 L3 L3 + + + L3 L9 L11 L11 L9 L4 L4 L3 L3 L3 L20 + * L3 + + + + + + + + + + + + L4 L3 * + + L3 * 22 + > 63 < Z B. Öztürk, G. Buzzurro & H. Benli Table 3 continued SPECIES Malleus sp. Pectinidae Pecten jacobaeus (Linnaeus, 1758) Aequipecten opercularis (Linnaeus, 1758) Pseudamussium septemradiatum (Müller O. F., 1776) Pseudamussium clavatum (Poli, 1795) Similpecten similis (Laskey, 1811) Palliolum incomparabile (Risso, 1826) Crassadoma multistriata (Poli, 1795) Mimachlamys varia (Linnaeus, 1758) Manupecten pesfelis (Linnaeus, 1758) Flexopecten flexuosus (Poli, 1795) Flexopecten glaber (Linnaeus, 1758) Lissopecten hyalinus (Poli, 1795) Spondylidae Spondylus gaederopus Linnaeus, 1758 Anomiidae Anomia ephippium Linnaeus, 1758 Limidae Lima lima (Linnaeus, 1758) Limaria hians (Gmelin, 1791) Limaria loscombi (Sowerby G. B. I, 1823) Limaria tuberculata (Olivi, 1792) Limatula gwyni (Sykes, 1903) Limatula subauriculata (Montagu, 1808) Ostreidae Ostrea edulis Linnaeus, 1758 Ostreola stentina (Payraudeau, 1826) Gryphaeidae Neopycnodonte cochlear (Poli, 1795) Lucinidae Ctena decussata (Costa O. G., 1829) Loripes lucinalis (Lamarck, 1818) Megaxinus unguiculinus Pallary, 1904 Divaricella angulifera Von Martens, 1880 Lucinella divaricata (Linnaeus, 1758) Anadontia fragilis (Philippi, 1836) Myrtea spinifera (Montagu, 1803) Thyasiridae Thyasira biplicata (Philippi,1836) Thyasira planata (Jeffreys, 1882) Axinulus croulinensis (Jeffreys, 1847) Axinulus cycladius (W ood S., 1848) Leptaxinus cf. subovatus (Jeffreys, 1881) Ungulinidae Diplodonta apicalis Philippi, 1836 Chamidae Chama gryphoides Linnaeus, 1758 Pseudochama corbierei (Jonas, 1846) Pseudochama gryphina (Lamarck, 1819) Kelliidae Bornia sebetia (Costa O. G., 1829) Montacutidae Tellimya ferruginosa (Montagu, 1808) L L23 N * R L3 L7 L3 L9 L13 L9 L9 L3 L3 L3 L9 L3 + + L3 + L3 + L3 L3 FR L23 L9 L18 + + + L3 L24 + + + + + + L9 L4 L3 L12 L16 L9 L9 L3 + + + + + L3 L25 L25 L9 L24 + L23 L3 L3 L9 L9 L25 > 64 < Z + * + 18 Marine molluscs from Cyprus: new data and checklist Table 3 continued SPECIES Carditidae Cardita calyculata (Linnaeus, 1758) Glans trapezia (Linnaeus, 1767) Glans elegans (Réquien, 1848) Pteromeris jozinae Aartsen, 1984 Pteromeris minuta (Scacchi, 1836) Cardites antiquatus (Linnaeus, 1758) Astartidae Astarte fusca (Poli, 1791) Astarte sulcata (Da Costa, 1778) Cardiidae Acanthocardia aculeata (Linnaeus, 1758) Acanthocardia echinata (Linnaeus, 1758) Acanthocardia paucicostata (Sowerby G. B. II, 1834) Acanthocardia tuberculata (Linnaeus, 1758) Parvicardium exiguum (Gmelin, 1791) Parvicardium ovale (Sowerby G. B. II, 1840) Parvicardium scabrum (Philippi, 1844) Parvicardium scriptum (B. D. D., 1892) Parvicardium trapezium Cecalupo & Quadri, 1996 Plagiocardium papillosum (Poli, 1791) Laevicardium crassum (Gmelin, 1791) Laevicardium oblongum (Gmelin, 1791) Cerastoderma edule (Linnaeus, 1758) Cerastoderma glaucum (Poiret, 1789) Fulvia fragilis (Forskâl in Niebuhr, 1775) Mactridae Mactra stultorum (Linnaeus, 1758) Mesodesmatidae Donacilla cornea (Poli, 1791) Solenidae Solen marginatus Pulteney, 1799 Pharidae Ensis arcuatus (Jeffreys, 1865) Ensis ensis (Linnaeus, 1758) Ensis minor (Chenu, 1843) Phaxas adriaticus (Coen, 1933) Phaxas pellucidus (Pennant, 1777) Tellinidae Tellina compressa Brocchi, 1814 Tellina distorta Poli, 1791 Tellina donacina Linnaeus, 1758 Tellina fabula Gmelin, 1791 Tellina incarnata Linnaeus, 1758 Tellina nitida Poli, 1791 Tellina pygmaea Lovén, 1846 Tellina planata Linnaeus, 1758 Tellina pulchella Lamarck, 1818 Tellina serrata Brocchi, 1814 Tellina tenuis Da Costa, 1778 Arcopagia balaustina (Linnaeus, 1758) Macoma cf. balthica (Linnaeus, 1758) Gastrana fragilis (Linnaeus, 1758) Donacidae L N R L3 L3 L4 L11 L23 L3 + + + L3 L9 + L4 L3 L1 L1 L4 L3 L18 L11 L23 L3 L9 L3 L3 L3 L16 + + + + + + + + + * L3 + L7 + L3 + L18 L3 L9 L25 L25 L23 L25 L3 L9 L3 L3 L25 L7 L3 L13 L9 L3 L24 L3 > 65 < + + + + + + + + + 23 + Z B. Öztürk, G. Buzzurro & H. Benli Table 3 continued SPECIES Donax semistriatus Poli, 1795 Donax venustus Poli, 1795 Donax trunculus Linnaeus, 1758 Psammobiidae Gari depressa (Pennant, 1777) Semelidae Abra nitida (Müller O. F., 1776) Abra prismatica (Montagu, 1808) Abra segmentum (Récluz, 1843) Abra tenuis (Montagu, 1803) Abra alba (W ood W ., 1802) Abra longicallus (Scacchi, 1834) Solecurtidae Solecurtus strigilatus (Linnaeus, 1758) Azorinus chamasolen (Da Costa, 1778) Kelliellidae Kelliella abyssicola (Forbes, 1844) Glossidae Glossus humanus (Linnaeus, 1758) Veneridae Venus verrucosa Linnaeus, 1758 Venus casina Linnaeus, 1758 Globivenus effosa (Philippi, 1836, ex Bivona ms.) Chamelea gallina (Linnaeus, 1758) Clausinella fasciata (Da Costa, 1778) Timoclea ovata (Pennant, 1777) Gouldia minima (Montagu, 1803) Dosinia lupinus (Linnaeus, 1758) Dosinia exoleta (Linnaeus, 1758) Pitar rudis (Poli, 1795) Callista chione (Linnaeus, 1758) Ruditapes decussatus (Linnaeus, 1758) Irus irus (Linnaeus, 1758) Paphia aurea (Gmelin, 1791) Paphia lucens (Locard, 1886) Paphia rhomboides (Pennant, 1777) Venerupis senegalensis (Gmelin, 1791) Petricolidae Petricola lithophaga (Retzius, 1786) Petricola substriata (Montagu, 1808) Petricola lajonkairii (Payraudeau, 1826) Mysia undata (Pennant, 1777) Corbulidae Corbula gibba (Olivi, 1792) Gastrochaenidae Gastrochaena dubia (Pennant, 1777) Hiatellidae Hiatella arctica (Linnaeus, 1767) Pholadidae Pholas dactylus Linnaeus, 1758 Barnea candida (Linnaeus, 1758) Teredinidae Teredo navalis Linnaeus, 1758 Teredo utriculus Gmelin, 1791 L L3 L3 L3 N L7 R + + L18 L25 L25 L3 L9 L25 + + L3 L3 + + FR + L3 + L3 L9 L25 L3 L9 L3 L3 L3 L9 L9 L3 L1 L3 L1 L1 L9 L3 + + 24 + + + + + + + + + + + + + + L3 L9 L9 L9 + + L7 + L7 + L9 + L4 L3 + + L9 L3 > 66 < Z + 19 Marine molluscs from Cyprus: new data and checklist Table 3 continued SPECIES Lyrodus pedicellatus (Quatrefages, 1849) Nototeredo norvegica (Spengler, 1792) Thraciidae Thracia papyracea (Poli, 1791) Thracia pubescens (Pulteney, 1799) Clavagellidae Clavagella melitensis Broderip, 1835 Pandoridae Pandora inaequivalvis (Linnaeus, 1758) Lyonsiidae Lyonsia norwegica (Gmelin, 1791) Cuspidariidae Cuspidaria cuspidata (Olivi, 1792) Cuspidaria rostrata (Spengler, 1793) Cardiomya costellata (Deshayes, 1835) L L13 L18 N R L9 L3 L9 L9 L25 SCAPHOPODA Dentaliidae Antalis dentalis (Linnaeus, 1758) Antalis entalis (Linnaeus, 1758) Antalis panorma (Chenu, 1842) Antalis rossati (Caprotti, 1966) Antalis vulgaris (Da Costa, 1778) Fustiariidae Fustiaria rubescens (Deshayes, 1826) Gadilidae Dischides politus (W ood S., 1842) Entalinidae Entalina tetragona (Brocchi, 1814) L3 L3 L3 + + + L3 L23 L4 L13 L2 + L2 + L18 L13 CEPHALOPODA Sepiidae Sepia elegans Blainville 1827 Sepia officinalis Linnaeus, 1758 Sepia orbignyana Férrussac in D’ Orbigny, 1826 Sepiolidae Sepiola robusta Naef, 1912 Sepiola rondeleti Leach, 1817 Sepiola steenstrupiana Levy, 1912 Sepietta oweniana (D’ Orbigny, 1840) Sepietta neglecta Naef, 1916 Loliginidae Loligo forbesii Steenstrup, 1856 Loligo vulgaris Lamarck, 1798 Alloteuthis media (Linnaeus, 1758) Ommastrephidae Illex coindetii (Vérany 1839) Todaropsis eblanae (Ball, 1841) Todarodes sagittatus (Lamarck, 1798) Octopodidae Octopus macropus Risso, 1826 Octopus salutii Vérany, 1839 Octopus vulgaris Cuvier, 1798 Scaeurgus unicirrhus (Delle Chiaje in Férrussac & D’ Orbigny, 1840) L2 L2 L26 L26 L2 L26 L26 L26 L26 L2 L26 L4 L26 L2 L2 L26 L2 L26 > 67 < Z B. Öztürk, G. Buzzurro & H. Benli Table 3 continued SPECIES Eledone moschata (Lamarck, 1798) Argonautidae Argonauta argo Linnaeus, 1758 L L2 N R Z L2 Literatur : L1 : MONTEROSATO, 1899; L2 : D EMETROPOULOS, 1969; L3 : D EMETROPOULOS, 1971; L4 : D EMETROPOULOS & H AD 1976; L5 : B AR ASH & D ANIN , 1977; L6 : V ERDUIN , 1984; L7 : STAVRINOS, 1984; L8 : V ERDUIN , 1986; L9 : T ORNARITIS, 1987; L10: B ARASH & DANIN, 1989; L11: B OGI et al., 1989; L12: AARTSEN & K INZELBACH, 1990; L13: B ARASH & DANIN, 1992; L14: LINDEN & EIKENBOOM, 1992; L15: FISCHER , 1993a; L16: FISCHER , 1993b; L17: CECALUPO & QUADRI, 1994; L18: FISCHER , 1994; L19: OLIVERIO & B UZZURRO, 1994; L20: OLIVERIO et al., 1994; L21: CECALUPO & QUADRI, 1995; L22: NOFRONI & T RINGALI, 1995; L23: CECALUPO & QUADRI, 1996; L24: B UZZURRO & GREPPI, 1997; L25: HADJICHRISTOPHOROU et al., 1997; L26: SALMAN et al., 1998; L27: SMRIGLIO et al., 1998; L28: HOUART , 2001, L29: B UZZURRO, 2002. J ICHRISTOPHOROU , NOTES (1997). 12: B EU (1998) has demonstrated that the Mediterranean form related to Charonia tritonis represents a distinct species instead of a subspecies. 13: Vexillum hypatiae (Pallary, 1912), reported by CECALUPO & QUADRI (1996) is not listed according to T RINGALI & V ILLA (1991). 14: for the claimed synonymy reported by CECALUPO & QUADRI (1996) with G ranulina clandestina (Brocchi, 1814) see G OFAS (1992). 15: in the past it has been identified by many Authors as Clathromangelia quadrillum (Dujardin, 1837), but this is a fossil species (OLIVERIO, 1995). 16: Mangelia fieldeni (Aartsen & Fehr de W al, 1978) = M. angelinae (Cecalupo & Quadri, 1996): see B UZZURRO & G REPPI (1997). 17: Chrysallida decorata (Philippi, 1849) is the correct name for the taxon commonly reported by several Authors as Chrysallida maiae (Hornung & Mermod, 1924) (B UZZUR R O & G R EPPI , unpublished data). 18: the genus has been assigned following SCHANDER & SUNDBERG (1997). 19: probably the record referred to Anisocycla nitidissima (Montagu, 1803). 20: reported by FISCHER (1994) from shallow water of the littoral zone, this is the only record of this deep-sea species, probably it is a case of misidentification with Caecilianella acicula Müller, 1774, a terrestrial species commonly found in our samples: see G AGLINI (1982). 21: for the putative finding of Cylichnina cf. mongii (Audouin, 1827) reported by CECALUPO & QUADRI (1996) see B UZZURRO & G REPPI (1997). 22: the name of this species comes from the Latin word regula(ae) = a ruler [and not from regulus(-i) = a small king!]. 23: for the presence of Macoma balthica (Linnaeus, 1758) in the Eastern Mediterranean Sea we refer to B UZZURRO (unpublished data). 24: European Authors traditionally refer decussatus to Tapes, whereas they currently use Ruditapes for the J apanese carpet shell (R. philippinarum (Adams & Reeve, 1850), which has been imported for acquaculture since a number of years. Both species PI 1: Lepidopleurus boettgeri Sulc, 1934 for the claimed synonymy with L . bedulli (Dell’Angelo & Palazzi, 1986), reported by CECALUPO & Q UADR I (1996), see D ELL’ANGELO & SMR IGLIO (1999). 2: Acanthochitona aeneus (Risso, 1826), reported by BOGI et al. (1989) is hard to consider, since the species belonging to this genus are only two according to D ELL’ANGELO & SMR IGLIO (1999) and are already present in the checklist, so we prefer not to report it. 3: Acanthochitona crinita oblonga Leloup, 1981 reported by CECALUPO & QUADRI (1996) has been excluded: see DELL’ANGELO & SMRIGLIO (1999). 4: this is the correct name considering that Haliotis lamellosa Lamarck, 1822 is a synonym of H. tuberculata : see G EIGER (1998). 5: several Authors have reported this species, while others have identified it as Gibbula umbilicaris (Linnaeus, 1758). Anatomical studies are needed to definitively solved this taxonomical problem. Due to the lack of data, we are conservatively in line with the interpretation of MONTEROSATO (1899: 398), who wrote: “Forme peu commune, voisine du G. umbilicaris Linné, dont elle diffère par sa forme conique, par ses taches suturales blanches et son ombilic plus étroit”. 6: following R EID (1996) we use Melarhaphe for neritoides (Linnaeus, 1758), the type and only living member of this genus. 7 : B OG I et a l. (1 9 8 9 ) reported a record of this species by T ORNARITIS (1987), who has nevertheless not reported it in his work. 8: the origin of Alvania dorbigny (Audouin, 1827) is still debated by nowadays Authors: we actually follow the opinion of MIENIS (1985) considering this species as non lessepsian. 9: G IANNUZZI-SAVELLI et al . (1997: 15) ascribed this species to the genus Rudolphosetia Monterosato, 1917: we follow PONDER (1985: 34) in considering Rudolphosetia as a junior synonym of Setia (H. & A. Adams, 1852). 1 0 : according t o A B B O T T (1 9 7 4 : 1 3 9 ) Calypt raeidae de Blainville, 1824 is a junior synonym of Crepulidae Fleming, 1822. 11: Erronea caurica (Linnaeus, 1758) reported by G IANNUZZI SAVELLI et al. (1997) has been excluded: see B UZZURRO & G REP- > 68 < Marine molluscs from Cyprus: new data and checklist 2 3 4 5 6 9 8 7 11 10 15 16 17 12 18 13 14 19 Figs 2-19 . Gastropods and bivalves from Cyprus. 2 : Nodilittorina (Echinolittorina) punctata (Gmelin, 1791) mm 6.6: St. C7. 3 : Rissoa splendida Eichwald, 1830 mm 4.8: St. C8. 4 : Alvania fractospira Oberling, 1970 mm 2.5: St. C10. 5 : Alvania testae (Aradas & Maggiore, 1844) mm 2.1: St. D10. 6 : Hydrobia acuta (Draparnaud, 1805) mm 3.2: St. C11. 7 : Protatlanta souleyeti (Smith E. A., 1888) mm 2.1: St. D17. 8 : Cerithiopsis scalaris Locard, 1892, ex Monterosato ms. mm 4.2: St. C8. 9 : Nassarius circumcictus (Adams A., 1852) mm 15: St. C10. 10 : Granulina boucheti Gofas, 1992 mm 2.2: St. C8. 11 : Gymnobela subaraneosa (Dautzenberg & Fischer H., 1896) mm 5.4: St. D17. 12 : Teretia teres (Reeve, 1844) mm 4.3: St. D17. 13 : Chrysallida juliae (de Folin, 1872) mm 1.9: St. C8. 14 : Folinella ghisottii Aartsen, 1984 mm 2.5: St. C8. 15 : Tragula fenestrata (Jeffreys, 1848) mm 3.1: St. C8. 16 : Eulimella bogii Aartsen, 1994 mm 1.9: St. C8. 17 : Turbonilla acutissima Monterosato, 1884 mm 5.3: St. D21. 18 : Limaria loscombi (Sowerby G. B. I, 1823) mm 8.9: St. C10. 19 : Kelliella abyssicola (Forbes, 1844) mm 1.6: St. D16 > 69 < B. Öztürk, G. Buzzurro & H. Benli belong to Ruditapes, as proposed by FISCHER -PIETTE & MÉTIVER (1971) and MATSUKUMA (1986); in particular, the last author excludes Ruditapes from Tapes. 25: Alvania sp., under study. New records of the molluscs fauna of Cyprus LITTORINIDAE Nodilittorina (Echinolittorina) punctata (Gmelin, 1791) (Fig. 2) Turbo punctatus Gmelin, 1791: 3597 Littorina syriaca Philippi, 1847: 165 (23), pl. 3, figs 21-22 Turbo siculus Chiereghin in N ARDO, 1847: 78, sp. 20, figs 909910 T urbo siculus B rugnone in MONT ER OSAT O , 1872: 33 (nomen nudum) Littorina punctata , Gmelin in MONTEROSATO, 1872: 33 Littorina (Melaraphe) punctata (Gmelin, 1791) in PRIOLO, 1953: 124 L ittorina punctata (Gmelin, 1791) = T ricolia draparnauldii Audouin, 1826 = Littorina pulchella Dunker, 1853 = Littorina syriaca Philippi, 1847 = Littorina guttata Reeve, 1858 = Littorina perplexa Turton, 1932 in R OSEW ATER , 1970: 474 Nodilittorina (Echinolittorina) punctata (Gmelin, 1791) in R EID, 1996: 99. Remarks: The allocation in the genus Nodilittorina von Martens, 1897 recently proposed by R EID (1996) for this species is surprising because Nodilittorina , type species L. pyramidalis Quoy & Gaimard, 1833, which is characterised by a strong ribbed sculpture with nodules, includes mainly Indo-pacific species with very different shell. The modern systematics of the family Littorinidae is primarily based on anatomical characters of the penis, pallial oviduct, radula, sperm nurse cells or spawn. See R EID (1986, 1989, 1996). Material: St. C7, C12, 19 specimens, Supralittoral Zone, rocky habitats. Distribution: Discontinuous distribution in Mediterranean basin, Portugal and African coasts of Atlantic Ocean. RISSOIDAE Rissoa splendida Eichwald, 1830 (Fig. 3) Rissoa splendida Eichwald, 1830: 219 Rissoa violaestoma Krynicky, 1837 fide CLEMAM Remarks: Rissoa splendida Eichwald, 1830 figured by G IAN NUZZI -SAVELLI et al. , 1997: 74, figs 203-204. Material: St. C8, 1 specimen, Cystoseira sp., 2-3 m and St. C10, 1 shell, grit near Posidonia oceanica , 4 m. Distribution: Aegean Sea, Central Mediterranean Sea, Adriatic Sea, the Marmara Sea and Black Sea. This species is here reported for the first time from the Levantine Basin. Alvania fractospira Oberling, 1970 (Fig. 4) Alvania fractospira Oberling, 1970: 3 Alvania monterosatoi sensu B OGI et al ., 1989: 194, fig. 7 (not FIS- CHER P., 1877) Remarks: Alvania fractospira Oberling, 1970 in MOOLENBEEK et al ., 1991: 113, fig. 16 (the lectotype is selected) and in OLIVERIO, 1993: 33 (restored as a valid name) . Material: St. C1, 3 specimens, Posidonia oceanica , 5 m; St. C8, 2 shells in grit near Posidonia oceanica , 4 m and St. C10, 2 specimens, Cystoseira sp., 1-2 m. Distribution: Eastern Mediterranean Sea and Adriatic Sea (G IANNUZZI-SAVELLI et al ., 1997). Alvania testae (Aradas & Maggiore, 1844) (Fig. 5) Rissoa reticulata Philippi, 1836: 156, tav. 10, fig. 14 (not T. reticulata Montagu, 1803) Rissoa testae Aradas & Maggiore, 1844: 135 Rissoa abyssicola Forbes in FORBES & HANLEY , 1850: 86 Rissoa partschii Hörnes, 1857: 573 (fossil) Rissoa (Alvania) testae (Aradas & Maggiore, 1844) in ARADAS & B ENOIT , 1870a: 199 Rissoa abyssicola var. conformis Jeffreys, 1870: 77 Rissoa (Actonia) testae Aradas & Maggiore in CARUS, 1889: 336 Actonia testae (Aradas & Maggiore, 1844) in MONTER OSATO , 1890: 147 Alvania testae (Aradas & Maggiore, 1844) in LOCARD, 1892: 161 Alvania (Actonia) testae (Aradas & Maggiore) in T HIELE, 1931: 163 Turbona testae (Aradas & Maggiore, 1844) in N ORDSIECK , 1972: 186 Remarks: Alvania testae (Aradas & Maggiore, 1844) in B OUCHET & W AR ÉN , 1993: 628, figs 1386-1387, 1400-1405 and in OLIVERIO et al ., 1992: 251 (A. testae complex is here analyzed and discussed). Material: St. D10 (35°34’8”N-34°28’3”E), 3 shells, muddy habitats, 210 m, dredged. Distribution: Eastern Mediterranean Sea (Aegean Sea), W estern Mediterranean Sea and coastal line of the Eastern Atlantic Ocean from Morocco to Norway. HYDROBIIDAE Hydrobia acuta (Draparnaud, 1805) (Fig. 6) Cyclostoma acutum Draparnaud, 1805: 40, pl. I, fig. 23 Paludestrina acuta Draparnaud in MONTEROSATO, 1878a: 86 Paludestrina glyca Servain, 1880: 151 Hydrobia acuta (Draparnaud, 1805) in G IUSTI & PEZZOLI (1984: 125, tav. 1) Remarks: Hydrobia acuta (Draparnaud, 1805) figured by G IUSTI et al. , 1995: 116, 117, figs 39-42, 50-52. After B OETERS (1984) that designated a lectotype for this species from two syntypes, G IUSTI et al . (1998) demonstrated that they to belong to different species and designate a neotype “in line with the earlier and more widely accepted usage of the name” (1998: 2, figs 3-6). Material: St. C11, 5 shells, grit, 1 m. Distribution: W hole Mediterranean basin (B ARASH & D ANIN, 1992). ATLANTIDAE > 70 < Marine molluscs from Cyprus: new data and checklist Protatlanta souleyeti (Smith E. A., 1888) (Fig. 7) Atlanta souleyeti Smith E. A., 1888: 1-51, pl. X II, fig. 7a, b Protatlanta souleyeti (Smith E. A., 1888) = Atlanta lamanonii Souleyet, 1852 = Protatlanta sculpta Issel, 1911 = Protatlanta sculpta var. mediterranea Issel, 1911 = Protatlanta mediterranea (Issel) Di Geronimo, 1970 in VATOVA, 1975: 1 Remarks: Protatlanta souleyeti (Smith E. A., 1888) figured by G IANNUZZI-SAVELLI et al ., 1997: 248, figs 921-923. Material: St. D17 (35°43’8”N-34°37’3”E), 1 shell, 250-300 m dredged. Distribution: It has a cosmopolitan distribution due to its pelagic life style, similar to other members of the family. Southern Aegean Sea, Central Mediterranean and W estern Mediterranean Sea. It is here reported for the first time in the Levantine basin. CERITHIOPSIDAE Cerithiopsis scalaris Locard, 1892, ex Monterosato ms. (Fig. 8) Cerithiopsis scalaris Monterosato, 1878a = C. corona var. scalaris Monterosato: 99 (nomen nudum) Cerithiopsis scalaris Monterosato, 1878 in LOCARD, 1892: 118 Cerithiopsis scalaris Coen, 1937: 31, tav. 3, fig. 26 (nomen nudum) Cerithiopsis scalaris (Monterosato, 1877) in NORDSIECK , 1976: 7; 6, fig. 10 Cerithiopsis scalaris Monterosato, 1877 in PIANI, 1980: 136 Cerithiopsis sca la ris Mont erosat o, 1 8 7 8 ( nomen nudum) in G AGLINI, 1992: 148, fig. 139 Cerithiopsis sca la ris Locard, 1 8 9 2 , ex Mont erosat o ms. in T RINGALI, 1999: 129, 130: figs 1-3 Remarks: Cerithiopsis scalaris Locard, 1892 figured by G IAN NUZZI -SAVELLI et al ., 1999: 41, figs 60- 63. Material: St. C8, 2 shells, grit near Posidonia oceanica , 4 m. Distribution: Central Mediterranean Sea. It is here reported from the Eastern Mediterranean Sea for the first time. NASSARIIDAE Nassarius circumcictus (Adams A., 1852) (Fig. 9) Nassa circumcincta Adams A., 1852: 102. Arcularia circumcincta Adams A. in PALLARY , 1912: 96, pl.X V (I), figg. 14-17 Nassarius (Plicarcularia) circumcictus (A. Adams, 1851) in PIANI, 1980: 149 Na ssa rius (Plica rcula ria ) circumcictus (Adams A., 1 8 5 2 ) = Naytiopsis granum flammulata Nordsieck, 1972 in NOFRONI, 1986: 13 Remarks: Arcularia circumcincta (Adams A., 1851) in SABELLI & SPADA, 1977: 3, figs 5a-5b and Nassarius circumcictus (Adams A., 1851) in OLIVERIO & T RINGALI, 1992: 159, figs 4-8. Material: St. C10, 2 specimens, sandy habitat, 2 m. Distribution: Eastern Mediterranean Sea. CYSTISCIDAE Granulina boucheti Gofas, 1992 (Fig. 10) Granulina boucheti Gofas, 1992: 10, fig. 9; 11, fig. 10; 24, fig. 26. Material: St. C8, 2 specimens, alive collected in brown algae, 1 m. Distribution: Sicily, France, Corse, Tunisie and Algerie. It is here reported from the Levantine basin for the first time. CONIDAE Gymnobela subaraneosa (Dautzenberg & Fischer H., 1896) (Fig. 11) Pleurotoma subaraneosa Dautzenberg & Fischer H., 1896: 422, pl. 16, figs 11-12 G ymnobela subaraneosa (Dautzenberg & Fischer H., 1896) in B OUCHET & W ARÉN, 1980: 58, figs 114-115 Pleurotomella subaraneosa (Dautzenberg & Fischer H., 1896) in POWELL, 1966: 131. Material: St. D17 (35°43’8”N-34°37’3”E), 1 shell, muddy bottoms, 250-300 m, dredged. Distribution: Mediterranean Sea and North Eastern Atlantic Ocean. R eported once from the Eastern Mediterranean Sea (J ANSSEN, 1989). Teretia teres (Reeve, 1844) (Fig. 12) Pleurotoma teres Reeve, 1844: 190 Philbertia anceps (Eichwald, 1830) (fossil): 225 Pleurotoma boreale Phil., ms. in LOVÉN, 1846: 14 Pleurotoma fusiforme Réquien, 1848: 101 Pleurotoma anceps in LOCARD, 1866: 109 Raphitoma barbierii Brusina, 1866: 33 Pleurotoma teres Forbes in MONTEROSATO, 1870: 45 Defrancia (Pleurotoma) teres Forbes in MONTEROSATO, 1872: 51 Pleurotoma (Defrancia) anceps Eichwald in MONTEROSATO, 1875: 44 Clathurella anceps V. Crs. in CARUS, 1889: 427 Teretia teres Forbes in MONTEROSATO, 1890: 187 Teretia anceps (Eichwald, 1830) in PIANI, 1980: 157 Teretia teres (Reeve, 1844) in SABELLI et al., 1990: 45 Remarks: Teretia teres (Forbes, 1844) figured by B OUCHET & W ARÉN, 1980: 81, fig. 168 and GIRIBET & PEÑAS, 1997: 75, fig. 69. Material: St. D17 (35°43’8”N-34°37’3”E), 1 shell, muddy bottoms, 250-300 m, dredged. Distribution: Coastline from Gibraltar Strait to Northern Norway, W estern Mediterranean Sea, Adriatic Sea and Aegean Sea. It is here reported from the Levantine basin for the first time. PYRAMIDELLIDAE Chrysallida juliae (de Folin, 1872) (Fig. 13) Truncatella juliae de Folin, 1872: 49, pl. II, fig. 4 Odostomia (Pyrgulina) nanodea Monterosato, 1878a: 93 Parthenina tenuistriata Milachewitch, 1909: 314 Chrysallida sarsi Nordsieck, 1972: 98, pl. II, fig. 4 in PEÑAS et al ., 1996: 24 Chrysallida juliae (de Folin, 1872) in AARTSEN, 1977: 61, fig. 13. Remarks: Chrysallida juliae (de Folin, 1872) in MICALI et al . (1993: 150): it is there designated the lectotype. Regarding all > 71 < B. Öztürk, G. Buzzurro & H. Benli the synonyms listed above we are waiting for the revision of these taxa accomplished by I. Nofroni (personal communication, manuscript in preparation). Material: St. C8, 3 shells, grit near Posidonia oceanica , 4 m. Distribution: Mediterranean Sea and Atlantic Ocean including the Canary Islands. Folinella ghisottii Aartsen, 1984 (Fig. 14) Odostomia intermedia Brusina, 1869: 237 (non Deshayes, 1861) Odostomia (Pyrgulina) intermedia B rusina in MONT ER OSAT O , 1878b: 317 Folinella ghisottii Aartsen, 1984: 137. Material: St. C8, 1 shell, grit near Posidonia oceanica , 4 m. Distribution: Mediterranean Sea and Atlantic Ocean including the Cape Verde Islands. T ragula fenestrata (Jeffreys, 1848) (Fig. 15) Odostomia fenestrata Jeffreys, 1848: 345, pl. 6, fig. 17 Chemnitzia fenestrata Jeffreys in FORBES & HANLEY , 1853: 249 Odostomia (Pyrgulina) fenestrata , Forbes in MONTEROSATO, 1877: 35 Tragula fenestrata (Jeffreys, 1848) in MONTEROSATO, 1884: 86 Odostomia (Tragula) fenestrata Forbes in PALLARY , 1912: 132, pl. X VI (II), fig. 42 Turbonilla fenestrata (Forbes) in B ABIO & T HIRIOT -QUIEVREUX , 1974: 544, pl. VI, H, L Tragula fenestrata (Forbes, 1848) in G ROSSU, 1986: 24 Chrysallida fenestrata (Forbes in J effreys, 1848) in AARTSEN , 1981: 80, pl. 1, fig. 1 Chrysallida fenestrata (Jeffreys, 1848) in LINDEN & EIKENBOOM, 1992: 45 Remarks: C. fenestrata (Jeffreys, 1848) figured by PEÑAS et al ., 1996: 27, fig. 54. Material: St. C8, 1 shell, grit near Posidonia oceanica , 4 m. Distribution: Mediterranean Sea and Atlantic Ocean including Corimba. Eulimella bogii Aartsen, 1994 (Fig. 16) Eulimella bogii Aartsen, 1994: 89-90, fig. 5. Material: St. C8, 1 shell, near Posidonia oceanica , 4 m. Distribution: Mediterranean Sea including the Eastern basin (NOFRONI & T RINGALI, 1995). LIMIDAE Limaria loscombi (Sowerby G. B. I, 1823) (Fig. 18) Lima loscombi Sowerby G. B. I, 1823: 17, pl. 113, fig. 4 Limea loscombei Sowerby in ARADAS & B ENOIT , 1870b: 94 Lima (Limatella) loscombei Leach in Sowerby in ANDERSON, 1964: 144, figs 27a-c L i ma ( L i mea ) l oscombi i Sowerby G . B ., 1 8 2 3 in A LT E N A R EGTEREN et al ., 1969: 24 (fossil) Lima (Mantellum) loscombi Sowerby = Lima bullata Turton = Lima clausa Danilo & Sandri in MUSETTI, 1969: 52 Limaria (Limatulella) loscombi (Sowerby G. B., 1830) in J ANSSEN, 1972: 16, 74, figs 4-5 (fossil) Limea (Limea) loscombi MacGillivray, 1843 in PIANI, 1980: 187 Limea loscombi (Sowerby G. B. I, 1823) in SABELLI et al., 1990: 85 Limea (Limatulella) loscombi (Sowerby G. B. I, 1823) in ZENETOS, 1996: 88 Limaria loscombii (Sowerby G. B. I, 1823) in SALAS, 1996: 57. Material: St. C10, 1 specimen, muddy–sandy habitats, 1.5 - 2 m. Distribution: This species is common in muddy and muddysandy bottoms down to a depth of 3000 m (PARENZAN, 1974), distributed in the Aegean Sea, Adriatic Sea, W estern Mediterranean Sea and the coastline of the Eastern Atlantic Ocean from Senegal and to Norway (J EFFREYS, 1879; T EBBLE, 1976; B ARASH & DANIN, 1992). KELLIELLIDAE Kelliella abyssicola (Forbes, 1844) (Fig. 19) Kellia abyssicola Forbes, 1844: 122 Venus miliaris Philippi, 1844: 36, pl. 14, fig. 15 Kelliella abyssicola Sars in MONTEROSATO, 1870: 39 Kelliella miliaris Philippi in SARS, 1878: 65 Kelliella miliaris Philippi in MONTEROSATO, 1880: 59 Kellyella miliaris (Phil.) in P AR ENZAN , 1974: 154, 265, fig. 158 Kelliella abyssicola (Forbes, 1844) in SABELLI et al ., 1990: 95. Material: St. D16 (35°19’9”N-32°50’1”E), 1 specimen, muddy habitats, 210 m, dredged. Distribution: Aegean Sea, Taranto Gulf and Northern Norway coasts (P AR ENZAN , 1974; P OPPE & G OT O , 1993; Z ENET OS, 1996). Already reported from the Eastern Mediterranean Sea by J ANSSEN (1989). This species shows a vertical distribution from the continental shelf down to the abyssal zone. DISCUSSION T urbonilla acutissima Monterosato, 1884 (Fig. 17) Turbonilla acutissima Monterosato, 1884: 92 Remarks: Turbonilla acutissima Monterosato, 1884 figured by AARTSEN, 1981: 85, pl. 4, fig. 24 and PEÑAS et al., 1996: 69, fig. 185. Material: St. D21(35°09’0”N-33°57’0”E), 1 shell, Posidonia oceanica , 20 m, dredge. Distribution: Mediterranean Sea (POPPE & G OTO, 1990). There has been a significant increase in the number of molluscan species reported from Cyprus since the first study carried out by MONTEROSATO (1899). In fact, in his report, an initial amount of 29 species was listed. But the number has grown up to 696 to date. Particularly, the studies of B OGI et al . (1989), F ISCHER (1994), CECALUPO & Q UADR I (1996) and B UZZURRO & G REPPI (1997) made important contribution to such increase. B OGI et al . (1989), reported 221 species in total, 89 as new records from this island. Then, FISCHER (1994) gave > 72 < Marine molluscs from Cyprus: new data and checklist a list of 123 fossils and 434 living species for Cyprus. This Author, following SABELLI et al . (1990-1992), has reconsidered t he syst emat ic posit ions of several species report ed by D EMET R OPOULOS (1 9 6 9 , 1 9 7 1 ), D EMET R OPOULOS & H AD J ICHRISTOPHOROU (1976), STAVRINOS (1984) and T ORNARITIS (1987). Since there are many different views concerning classification, at the mean time FISCHER (1994) followed his personal conception regarding some cases of synonymy, changing the taxonomical rank of some species. For example, while Emarginula cancellata Philippi, 1836 and Emarginula sicula Gray J . E., 1825 are two synonyms according to SABELLI et al . (1990), on the contrary FISCHER (1994: 87) reported these two taxa as distinct species. In our opinion the sole valid species name is E. sicula . Other controversial cases of synonymy that we had to face making the new checklist are discussed below. B OGI et al . (1989) reported Pirenella conica (Blainville, 1826) and Pirenella cailliaudi (Potiez & Michaud, 1838) as different species, while the latter should be considered just as junior synonym of the first one according to SABELLI et al . (1990-1992). In like man- ner, Lima lima (Linnaeus, 1758) and Lima inflata (Chemnitz, 1784) should be thought as synonyms, taking as valid species name L . lima . Similarly, MI CALI et a l . (1 9 9 3 ) considered Chrysallida cf. delpretei (Sulliotti, 1889) as junior synonym of Chrysallyda indistincta (Montagu, 1808): both taxa are listed as separate species in B OGI et al. (1989). An another example of systematic confusion is the case of Retusa semisulcata (Philippi, 1836) and Retusa truncatula (Brugière, 1792), both listed in CECALUPO & QUADRI (1996), but re-discussed as synonyms by T RINGALI (1995). Of a total of 3 8 9 species listed by CECALUPO & Q UAD R I (1996), 134 species were new ones for Cyprus. In their papers, B UZZURRO & G REPPI (1997) and SALMAN et al. (1998) have reported 42 and 10 additional new species, respectively, for the Cyprus coasts. W hen the lists of B OGI et al . (1989) and CECALUPO & QUADRI (1996) are compared, 6 and 24 species, respectively, given as new records had already been reported previously (Table 4). Additionally, Argonauta argo Linnaeus, 1758 reported by SALMAN et al . (1998) as a new record, was Table 4. Species previously listed by B OGI et al. (1989) (* ) and CECALUPO & QUADRI (1996) (* * ) as new records and the literature relevant to these species. Literature giving former records B ARASH & DANIN (1977: 7) B ARASH & DANIN (1977: 9) B ARASH & DANIN (1977: 12) FISCHER (1993 A: 38) FISCHER (1993 A: 39) AARTSEN & K INZELBACH (1990: 109) AARTSEN & K INZELBACH (1990: 109) VERDUIN (1986: 30) VERDUIN (1984: 49) FISCHER (1993 A: 39) STAVRINOS (1984: 21) OLIVERIO et al . (1994: 214) T ORNARITIS (1987: 103) FISCHER (1993 A: 40) FISCHER (1993 A: 40) DEMETROPOULOS (1971: 20) DEMETROPOULOS (1971: 16) LINDEN & EIKENBOOM (1992: 18) B ARASH & DANIN (1989: 248) B ARASH & DANIN (1989: 248) FISCHER (1993 A: 40) FISCHER (1993 A: 40) DEMETROPOULOS (1969: 13) B ARASH & DANIN (1989: 250) T ORNARITIS (1987: 114) B ARASH & DANIN (1989: 253) DEMETROPOULOS & HADJICHRISTOPHOROU (1971: 79) FISCHER (1994: 93) DEMETROPOULOS (1971: 21) FISCHER (1994: 93) FISCHER (1994: 93) Species Ischnochiton rissoi (Payraudeau, 1826) * Lepidochitona corrugata (Reeve, 1848) * Chiton corallinus (Risso, 1826)* * Gibbula umbilicaris (Linnaeus, 1758)* * Cerithium scabridum Philippi, 1848 * * Rissoa auriformis pseudomonodonta (Verduin, 1983)* * Alvania datchaensis Amati & Oliverio, 1987 * * Alvania mamillata Risso, 1826 * * Alvania beniamina (Monterosato, 1884) * * Strombus persicus Swainson, 1821 * * Petaloconchus glomeratus (Linnaeus, 1758) * * Vitreolina levantina Oliverio, Buzzurro & Villa, 1994 * * Vexillum littorale (Forbes, 1844) * Bela laevigata (Philippi, 1836) * * Bela menkhorsti Aartsen, 1988 * * Bela nebula (Montagu, 1803) * Mitrolumna olivoidea (Cantraine, 1835)* Chrysallida terebellum (Philippi, 1844)* * Turbonilla delicata (Monterosato, 1874) * * Turbonilla striatula (Linnaeus, 1758)* * Retusa mamillata (Philippi, 1836) * * Retusa semisulcata (Philippi, 1836) * * Cylichnina umbilicata (Montagu, 1803) * Cylichna crossei (B.D.D., 1886) * * Cavolinia tridentata (Niebuhr, 1775 ex Forskäl, ms.) * * Styliola subula (Quoy & Gaimard, 1827) * * Modiolarca subpicta (Cantraine, 1835)* Lucinella divaricata (Linnaeus, 1758) * * Cerastoderma edule (Linnaeus, 1758) * * Abra nitida (Müller O. F., 1776) * * Gastrochaena dubia (Pennant, 1777) * * > 73 < B. Öztürk, G. Buzzurro & H. Benli previously reported by D EMETROPOULOS (1969). Besides that, Ana chis troglodytes (Souverbie & Mont rouzier, 1 8 6 6 ) and Anachis savignyi Moazzo, 1939 ex J ousseaume ms. reported as synonyms by CECALUPO & QUADRI (1996), are considered as two distinct species according to T RINGALI & VILLA (1995). Furthermore, although CECALUPO & QUADRI (1996) discussed that some of the species which had been reported as synonyms by SABELLI et al . (1990) can be accepted as valid taxa , species such as Ema rginula crebrisculpta (Coen, 1 9 3 9 ), Ca lliostoma alexandrinum Pallary, 1911, Muricopsis inermis (Philippi, 1836), Mangelia derelicta R eeve, 1846, Raphitoma intermedia Nordsieck, 1977 and Raphitoma lineolata (B.D.D., 1883) are not valid species according to the CLEMAM database. Some of the species names found in literature referred to the Island of Cyprus have been omitted from the checklist because fossils: Alvania monterosatoi (Fischer P., 1876), Granulina clandestina (Brocchi, 1814), Clathromangelia quadrillum (Dujardin, 1837), Raphitoma histrix Bellardi, 1847, Odostomia bismichaelis Sacco, 1892 and Odostomia cf. clavulina Fischer P., 1877. Also Hydrobia ulvae (Pennant, 1777), most probably confused with other Hydrobia sp., has been deleted since we reasonably think it is not a Mediterranean species. Other cases of shell misidentification reported by T ORNARITIS (1987) are: Cerithiopsis tubercularis (Montagu, 1803) which is Dizoniopsis coppolae (Aradas, 1870) (: 58, fig. 80b), Fusinus pulchellus (Philippi, 1844) is just a juvenile form of F. syracusanus (Linnaeus, 1758) (: 98, fig. 146), Mathilda cochlaeformis Brugnone, 1873 which is M. gemmulata Semper, 1865 (:54, fig. 72); Mytilus edulis Linnaeus, 1758 which is M. galloprovincialis Lamarck, 1819, since the former is not an east ern medit erranean species. Hinnites distorta , to allocate within the genus Crassadoma according to W ALLER (1993), has been reported only once by D EMETROPOULOS (1971), most probably it has been misidentified with C. multistriata . W e remove this taxon from the proposed checklist of molluscs of Cyprus, since this species is mainly distributed in the Atlantic Ocean and very seldom reported from the W estern Mediterranean basin. FISCHER (1994) has reported over 90 Indo-Pacific species in the Levantine basin, and 8 of them have also been recorded from the Cyprus coasts. Today, the number of the known Indo-Pacific species occurring in the Island of Cyprus increased up to 32 (Table 3). Summing up all the data coming from this investigation and those of the past, a total of 696 Molluscs species are presently known from Cyprus. ACKNOWLEDGEMENTS Sincere thanks are due to: Dr. P. Mariottini (Università di Roma Tre, Roma, Italy) for critical reading of the manuscript, Dr. J . M. Poutiers (MNHN) for helpful suggestions, Dr. V. Aysel (University of Canakkale, Turkey) for identifying the sampled algae, Dr. R. La Perna (Università di Bari, Italy) for helpful advises on Bivalvia Systematics, Dr. M. Bodon (Università di Siena, Italy), Dr. S. Chiarelli (Porto Garibaldi, Italy), Dr. S. Gofas (University of Malaga, Spain), Dr. A. W . J anssen (Gozo, Malta), Dr. G. Manganelli (Università di Siena, Italy), Dr. H. Mienis (University of J erusalem, Israel), Mr. G. Nardi (Ghedi, Italy), Dr. M. Oliverio (Università “La Sapienza”, Roma, Italy), Mr. P. Russo (Venezia, Italy), Dr. M. Streitz (Valbonne, France) and Dr. A. Zenetos (National Centre for Marine Research, Hellenikon, Greece) for generously providing references. W e are very indebted to Mr. A. Cecalupo (Buccinasco, Italy) for his cooperation in dealing with the many synonyms treated in the present paper and in supplying references. W e are really grateful to Dr. J . J . Van Aartsen (Dieren, The Netherland) and Dr. P. Micali (Fano, Italy) for G. subaraneosa and T . acutissima identification, respectively. W e are deeply indebted to Dr. S. Chiarelli (Porto Garibaldi, Italy) and Dr. E. Greppi (Castellanza, Italy) for having taken the pictures utilized in the present paper. REFERENCES AARTSEN J . J . VAN, 1977. European Pyramidellidae: I Chrysallida . Conchiglie, Milano, 13 (3-4): 49-64. AARTSEN J . J . VAN , 1981. 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