CATALOG OF RECENT MOLLUSCAN TYPES IN THE
NATURAL HISTORY MUSEUM OF LOS ANGELES
COUNTY
compiled by
Lindsey T. Groves
NATURAL HISTORY MUSEUM OF LOS ANGELES COUNTY
MALACOLOGY SECTION
900 Exposition Boulevard, Los Angeles, California, 90007, USA
lgroves@nhm.org
[last updated 3 January, 2012]
Abstract. Recent molluscan types currently housed in the Malacology Section of the Natural
History Museum of Los Angeles County are listed herein. Each type lot entry includes original
species/subspecies name, original genus, author, original citation, type status, LACM number,
number of specimens and preservation, type locality or collection locality, collector [if known],
date collected [if known], and current taxonomic status if different from original description.
INTRODUCTION
This catalog is published in accordance with Recommendation 72F.4 of the International
Code of Zoological Nomenclature (Ride & others, 2000:79). This listing includes all molluscan
type material currently housed at the Natural History Museum of Los Angeles County and
supercedes the type listing of Sphon (1971). To date 1922 molluscan type lots (71,000+
specimens) are reported for 1147 species and 99 subspecies, which includes 589 holotypes, 1277
paratype lots, 10 lectotypes, 13 paralectotype lots, 29 syntype lots, 3 neotypes, and 2 neotype lots
in 550+ references.
Type categories listed include: Holotype (in red), paratype(s) (in blue), syntype(s) (in
orange), lectotype (in pink), paralectotype(s) (in aqua), and neotype and/or neotype lots (in
brown) as defined by Ride & others (2000). Because the term cotype is not recognized by the
ICZN, lots originally designated as cotype(s) (in green) by original specimen labels are now
referred to as syntype(s). Hypotypes (or figured non-type specimens) are not housed within the
type collection and are considered voucher specimens.
Species are listed alphabetically within each family regardless of subfamily and/or genus.
Classifications schemes are according to Coan & others (2000) for the bivalves, Bouchet &
others (2005) for the gastropods, Steiner & Kabat (2001) for the scaphopods, Sirenko (2006) for
the polyplacophorans, Scheltema (1998) for the aplacophorans, and Warén & Gofas (1996) for
the monoplacophorans. These arrangements are utilized for convenience as classification
schemes are subject to change.
Brief History of the Type Collection
The LACM Recent Molluscan type collection was started by George Willett (then LACM
curator of Ornithology & Mammalogy) in the late 1920’s who organized a separate sequence (ie.,
a unique lot number for each type lot) and his mollusk types became the core of the LACM
Recent type catalog. The type collection grew tremendously following the arrival in 1964 of
James H. McLean, the first full-time curator of Malacology, due to his diligent collecting and
numerous publications. Since Jim’s retirement in 2001, the type collection has experienced
sustained growth, especially the opisthobranch types, under former curator Ángel Valdés.
It is obvious from the following list that the LACM malcology type collection has greatly
benefited from its relationship with numerous professional and amateur malacologists and
conchologists of the past and present who have either donated type material and/or utilized the
world-class LACM mollusk collection for their research purposes. Notable living type species
authors include James H. McLean (Gastropoda), Eugene V. Coan (Bivalvia), Louie N.
Marincovich, (Gastropoda), Bret K. Raines (Gastropoda), Roger N. Clark (Polyplacophora),
Daniel L. Geiger (Vetigastropoda), Barry Roth (terrestrial Gastropoda), William K. Emerson
(Gastropoda), Peter Jung (Gastropoda & Bivalvia), Thomas A. Burch (Gastropoda & Bivalvia),
Hans Bertsch (Opisthobranchia), and most recently Ángel Valdés (Opisthobranchia). Prominent
molluscan type authors of the past include Twila Bratcher (Terebridae), Wendell O. Gregg
(terrestrial Gastropoda), Helen DuShane (Epitoniidae), Frank R. Bernard (Bivalvia), Dwight W.
Taylor (Gastropoda & Bivalvia), LeRoy Poorman (Gastropoda), George Willett (Gastropoda), S.
Stillman Berry (Gastropoda), Antonio J. Ferreira (Polyplacophora), Leo G. Hertlein
(Gastropoda), John Q. Burch (Gastropoda & Bivalvia), Herbert N. Lowe (Gastropoda), Donald
R. Shasky (Gastropoda), Archibald M. Strong (Gastropoda), Gilbert Grau (Bivalvia), and
Crawford N. & Jean M. Cate (Cypraeoidea & Mitridae) Historical paratype and syntype
material acquired through collection donations and/or purchases includes authors such as
William H. Dall, Thomas Say, Henry A. Pilsbry, Carlos de la Torre, Paul Bartsch, George B.
Sowerby III, Tom Iredale, Henry Hemphill, and Hugh C. Fulton.
Type Collection Usage and Donations
The LACM type collection is arranged numerically and housed in locked cabinets. A
complete inventory of the type collection is available in the database and reports can be generated
as with non-type specimens. A staff member must be present when type material is being
studied. Type specimens may be borrowed by bonafide researchers for a period of six months
and renewal of type loans are subject to approval by the curator. LACM type numbers may be
requested from the collection manager whether or not the specimens originated in the LACM
collection. Researchers who wish to deposit type specimens at LACM must provide identified
specimens with as complete locality data as possible upon deposition including depth or altitude,
habitat, specific locality, coordinates, collector and/or research vessel name, field number,
collection date, etc. This data can be in the form of log books, field notes, specimen labels, etc.
Type numbers will be assigned after specimens are received by the LACM Malacology section.
The donor will agree to sign and date a LACM Deed of Gift form, which gives LACM legal
ownership or title. The donor will receive a copy of this form when the accession process is
completed along with acknowledgement from the LACM President/Director if desired. The
collection manager will process the donation and file an Accession Data Sheet with the Deed of
Gift with the LACM Registrars office. All donations to LACM are tax deductible.
Natural History Museum of Los Angeles County
MALACOLOGY TYPE CATALOG
Phylum Mollusca Cuvier, 1791
CLASS APLACOPHORA von Ihering, 1876
Subclass Chaetodermorpha Pelseneer, 1906
Family Prochaetodermatidae Salvini-Plawen, 1969
californicum, Prochaetoderma Schwabl, 1963: Pacific Science 17:265-267, figs. 4-7.
LECTOTYPE: LACM 2091 (wet), designated by Scheltema (1998:44), Tax. Atlas Benthic Fauna
Santa Maria Basin & W. Santa Barbara Channel 9(2); PARALECTOTYPES: LACM 2092 (3
wet), 704 m, 4.3 mi. WSW of Pt. Fermin light, Los Angeles Co., California (33 40.2'N,
118 22.1 W), Leg.: R/V Velero IV, 24 JUN 1953
Remarks: Currently in the genus Spathoderma fide Scheltema (1998).
Family Chaetodermatidae von Ihering, 1876
elegans, Chaetoderma Scheltema, 1998: Tax. Atlas Benthic Fauna Santa Maria Basin & W.
Santa Barbara Channel 9(2):22-24, fig. 2.9.
HOLOTYPE: LACM 2748 (wet),1808 m, 14 mi. 321 deg. T to Gull Islet light, S of Santa Cruz
Id., California Channel Ids., California (33 45.85 N, 119 39.25 W), Leg.: R/V Velero IV, 18 JAN
1976.
Remarks: Spicules mounted on glass slides.
hancocki, Crystallophrisson Schwabl, 1963: Pacific Science 17:272-273, figs. 14-15.
LECTOTYPE: LACM 2093 (wet), designated by Scheltema (1998:26), Tax. Atlas Benthic Fauna
Santa Maria Basin & W. Santa Barbara Channel 9(2), 816 m, 5.2 mi. ENE of Long Pt., Santa
Catalina Id., California Channel Ids., California (33 26 N, 118 15.97 W), Leg.: R/V Velero IV,
17 JUN 1954.
Remarks: Currently in the genus Chaetoderma fide Scheltema (1998).
hartmani, Crystallophrisson Schwabl, 1961: Zool. Anz. 166(7):258-277, figs. 1-10.
LECTOTYPE: LACM 2094 (wet), designated by Scheltema (1998:36), Tax. Atlas Benthic Fauna
Santa Maria Basin & W. Santa Barbara Channel 9(2), 330 m, 3.5 mi. W of Pt. Fermin light, Los
Angeles Co., California (33 42 N, 118 21.82 W), Leg.: R/V Velero IV, 27 FEB 1953;
PARALECTOTYPE: LACM 2095 (1 wet), 570 m, 7 mi. WSW of Redondo Beach pier, Los
Angeles Co., California (33 47.93 N, 118 31.27 W), Leg.: R/V Velero IV, 26 SEP 1952
Remarks: Currently in the genus Falcidens fide Scheltema (1998).
incrassatum, Crystallophrisson Schwabl, 1963: Pacific Science, 17:267, fig. 8.
LECTOTYPE: LACM 2096 (wet), designated by Scheltema (1998:42), Tax. Atlas Benthic Fauna
Santa Maria Basin & W. Santa Barbara Channel 9(2), 558 m, 19.5 mi. SW of Santa Barbara Pt.,
Santa Barbara Co., California (34 10.6 N, 120 0.7 W), Leg.: R/V Velero IV, 12 DEC 1955.
Remarks: Currently in the genus Furcillidens fide Scheltema (1998).
inflatum, Crystallophrisson Schwabl, 1963: Pacific Science, 17:276-277, figs. 21-22.
LECTOTYPE: LACM 2097 (wet), designated by Scheltema (1998:26), Tax. Atlas Benthic Fauna
Santa Maria Basin & W. Santa Barbara Channel 9(2), 759 m, 8.8 mi. WNW of Palos Verdes
light, Los Angeles Co., California (33 48 N, 118 36.05 W), Leg.: R/V Velero IV, 22 MAY 1954;
PARALECTOTYPE: LACM 2098 (1 wet), 781 m, 4.45 mi. SW of Pt. Fermin light, Los Angeles
Co., California (33 39.27 N, 118 21.48 W), Leg.: R/V Velero IV, 20 APR 1952.
Remarks: Junior synonym of Chaetoderma hancocki (Schwabl, 1963) fide Scheltema (1998).
longus, Falcidens Scheltema, 1998: Tax. Atlas Benthic Fauna Santa Maria Basin & W. Santa
Barbara Channel 9(2):38, fig. 2.17.
PARATYPE: LACM 2749 (1 wet), 92 m, Santa Maria Basin, NW of Pt. Arguello, Santa
Barbara Co., California (34 43.01 N, 120 47.39 W), Leg.: USDI Minerals Management Service,
R/V Aloha, MAY 1989
Remarks: Spicules and radula mounted on glass slides.
macracanthos, Falcidens Scheltema, 1998: Tax. Atlas Benthic Fauna Santa Maria Basin & W.
Santa Barbara Channel 9(2):40, fig. 2.18.
PARATYPE: LACM 2750 (1 wet), 145 m, Santa Maria Basin, W of Pt. Sal, Santa Barbara Co.,
California (34 55.79 N, 120 49.91 W), Leg.: USDI Minerals Management Service, R/V Aloha,
JAN 1988; PARATYPE: LACM 2751 (1 wet), 92 m, Santa Maria Basin, NW of Pt. Arguello,
Santa Barbara Co., California (34 43.01 N, 120 47.39 W), Leg.: USDI Minerals Management
Service, R/V Aloha, MAY 1988; PARATYPE: LACM 2752 (1 wet), 113 m, Santa Maria Basin,
SE of Government Pt., Santa Barbara Co., California (34 25.88 N, 120 16.31 W), Leg.: USDI
Minerals Management Service, R/V Aloha, 1988.
Remarks: Radulae from paratypes LACM 2751 and 2752 mounted on glass slides.
marinelli, Crystallophrisson Schwabl, 1963: Pacific Science, 277-278, figs. 23-24.
LECTOTYPE: LACM 2099 (wet), designated by Scheltema (1998:28), Tax. Atlas Benthic
Fauna Santa Maria Basin & W. Santa Barbara Channel 9(2), 29 m, 0.75 mi. SE of Pt. Vicente
light, Los Angeles Co., California (33 44 N, 118 34.97 W), Leg.: R/V Velero IV, 28 OCT 1953.
Remarks: Currently in the genus Chaetoderma fide Scheltema (1998).
pacificum, Crystallophrisson nitidulum Schwabl, 1963: Pacific Science, 17:273-274, figs.
16-18.
LECTOTYPE: LACM 2100 (wet), designated by Scheltema (1998:30), Tax. Atlas Benthic
Fauna Santa Maria Basin & W. Santa Barbara Channel 9(2), 283 m, 7.0 mi. SW of Huntington
Beach pier, Los Angeles Co., California (33 34 N, 118 6.0 W), Leg.: R/V Velero IV, 3 MAR
1954; PARALECTOTYPE: LACM 2102 (1 wet), 51 m, 1.9 mi. W of Salta Verde Pt., Santa
Catalina Id., California Channel Ids., California (33 29.12 N, 118 27.42 W), Leg.: R/V Velero
IV, 31 OCT 1952; PARALECTOTYPE: LACM 2103 (1 wet), 39.3 m, 4 mi. SE of Pt. Fermin
light, Los Angeles Co., California (33 39.32 N, 118 10.3 W), Leg.: R/V Velero IV, 12 OCT
1953; PARALECTOTYPE: LACM 2104 (1 wet), 270 m, 2.35 mi. S of Newport Beach pier,
Orange Co., California (33 33 N, 117 53.93 W), Leg.: R/V Velero IV, 14 MAY 1954.
Remarks: Elevated to full species in the genus Chaetoderma by Scheltema (1998).
riedli, Crystalloprisson Schwabl, 1963: Pacific Science, 17:274-276, figs. 19-20.
LECTOTYPE: LACM 2105 (wet), designated by Scheltema (1998:30), Tax. Atlas Benthic
Fauna Santa Maria Basin & W. Santa Barbara Channel 9(2); PARALECTOTYPE: LACM 2106
(1 wet), 570 m, 7 mi. W of Redondo Beach pier, Los Angeles Co., California (33 47.93 N,
118 31.27 W), Leg.: R/V Velero IV, 26 SEP 1952.
Remarks: Junior synonym of Chaetoderma nanulum Heath, 1911 fide Scheltema (1998).
Subclass Noemeniomorpha Pelseneer, 1096
Family Gymnomeniidae Odhner, 1906
mariensis, ?Genitoconia Scheltema, 1998: Tax. Atlas Benthic Fauna Santa Maria Basin & W.
Santa Barbara Channel 9(2):15, fig. 2.3.
PARATYPE: LACM 2747 (1 wet), 145 m, Santa Maria Basin, W of Pt. Sal, Santa Barbara Co.,
California (34 55.79 N, 120 49.91 W), Leg.: USDI Minerals Management Service
minuta, ?Gymnomenia Scheltema, 1998: Tax. Atlas Benthic Fauna Santa Maria Basin & W.
Santa Barbara Channel 9(2):12-13, fig. 2.2.
PARATYPE: LACM 2745 (1 wet), 410 m, Santa Maria Basin, NW of Pt. Arguello, Santa
Barbara Co., California (34 41.4 N, 120 57.9 W); PARATYPE: LACM 2746 (1 wet), 410 m,
Santa Maria Basin, W of Pt. Sal, Santa Barbara Co., California (34 53.68 N, 120 29.12 W),
Leg.: USDI Minerals Management Service, R/V Aloha, OCT 1988.
CLASS MONOPLACOPHORA Odhner in Wenz, 1940
Order Tryblidioidea Lemche, 1957
Family Neopilinidae Knight & Yochelson, 1958
hyalina, Vema (Laevipilina) McLean, 1979: Nat. His. Mus. L.A. Co. Cont. Sci. 307:11-18, fig.
9.
HOLOTYPE: LACM 1787 (wet), PARATYPES: LACM 1788 (1 wet, 1 dry), 373-384 m, Santa
Rosa-Cortes Ridge, Los Angeles Co., California (32 59 N, 119 32.8 W), Leg.: H.A. Lowenstam
& P.I. LaFollette, 14 FEB 1977.
veleronis, Neopilina (Neopilina) Menzies & Layton, 1962: Ann. & Mag. Nat. Hist., Ser. 13,
5:402-405, pls. 7-10.
HOLOTYPE: LACM 2720 (wet); PARATYPE: LACM 2721 (1 wet), 2730-2769 m, 30.75 mi.
268.5 deg.T from Isla Natividad Light, Pacific Coast Baja California Sur, México (27 52 N,
115 44.9 W), Leg.: R/V Velero IV, 31 DEC 1960.
Remarks: Holotype ex AHF 60. Currently in the genus Veleropilina fide Warén & Gofas (1996).
Holotype and paratype radulae mounted on SEM stubs. Shells completely recrystallized
(paratype destroyed during body extraction) fide Warén & Gofas (1996).
Family Vemidae Clark & Menzies, 1959
levinae, Vema Warén & Gofas, 1996: Zool. Scripta 25(3):226-232, figs. 1A,13A-G, 14A-G,
15B.
HOLOTYPE: LACM 2790 (dry), 1058 m, close to peak of submarine volcano, East Pacific Rise,
East Pacific (12 56'N, 103 29'W), Leg.: L. Levin, R/V Alvin, 13 JUN 1984.
CLASS POLYPLACOPHORA Gray, 1821
Subclass Loricata Schumacher, 1817
Order Lepidopleurida Thiele, 1910
Suborder Lepidopleurina Thiele, 1910
Family Leptochitonidae Dall, 1889
bartletti, Lepidopleurus Ferreira, 1986c: The Nautilus 100(3):99-101.
PARATYPES: LACM 2123 (2 wet), 3982 m, Venezuela Basin, NE of Bonaire, Netherlands
Antilles (15 08 N, 69 12 W), Leg.: R/V Bartlett, 28 OCT 1981.
Remarks: Currently in the genus Ferreiraella fide Kaas & Van Belle (1998). Emended to
Lepidopleurus bartlettae Ferreira, 1986 by Kaas & Van Belle (1987).
scrippsianus, Lepidopleurus Ferreira, 1980: The Veliger 23(1):55-60.
PARATYPE: LACM 1700 (1 wet), 2507-2891 m, SW of Cabo San Lucas, Golfo de California,
Baja California Sur, México (22 30.8 N, 110 03.8 W), Leg.: C.L. Hubbs, 29-30 JUL 1965.
Remarks: Currently in the genus Ferreiraella fide Kaas & Van Belle (1998).
xylophagus, Xylochiton Gowlett-Holmes & Jones, 1992: Jour. Malac. Soc. Australia 13:38-43.
PARATYPE: LACM 2280 (1 wet), 1075-1100 m, NE of White Id., South Auckland, North Id.,
New Zealand (37 23.7 S, 177 38.1 E), Leg.: R/V Kalinovo.
Family Ferreiraellidae Dell’Angelo & Palazzi, 1991
caribbensis, Ferreiraella, Sirenko, 1988: Zool. Zhurl. 67(12):1778-1781.
PARATYPE: LACM 1972 (1 wet), 6740-6780 m, Cayman Trench, S of Santiago de Cuba Prov.,
Cuba (19°39'N, 76°37'W), Leg.: R/V Kurchatov, 24 MAR 1973.
Order Chitonida Thiele, 1910
Superfamily CHITONOIDEA Rafinesque, 1815
Family Ischnochitonidae Dall, 1889
allynsmithi, Lepidozona (Lepidozona) Ferreira, 1974: The Veliger 17(2):173-175.
PARATYPE: LACM 1727 (1 dry), 2-3 m, Caleta El Candelero, W side of Isla Espíritu Santo,
Golfo de California, Baja California Sur, México (24 30 N, 110 23 W), Leg.: A.J. Ferreira, 18
OCT 1970.
attuensis, Lepidozona (Tripoplax) Clark, 2000a: Nemouria 42:9-11, figs. 19-21.
HOLOTYPE: LACM 2801 (wet), intertidal, Murder Pt., Massacre Bay, Pacific side of Near Ids.,
Aleutian Ids., Alaska (52°47.6'N, 173°11.0'E), Leg.: D.R. Lindberg, 20 JUL 1979.
baxteri, Lepidozona (Tripoplax) Clark, 2000a: Nemouria 42:6-9, figs. 8-11.
HOLOTYPE: LACM 2796 (wet); PARATYPES: LACM 2797 (10 wet), 0-1 m, Eider Pt., W side
entrance to Unalaska Bay, Bering Sea side of Unalaska Id., Aleutian Ids., Alaska (53°57.6'N, 16°
35.50'W), Leg.: R. Baxter, JUN 1993.
beringiana, Lepidozona (Tripoplax) Clark, 2000a: Nemouria 42:3-6, figs. 5-7.
HOLOTYPE: LACM 2798 (dry); PARATYPE: LACM 2799 (1 dry), 121 m, S of
Semisopochnoi Id., Rat Ids., Aleutian Ids., Alaska (51°53.34'N, 179°45.58'E), Leg.: R. Baxter, 7
SEP 1986; PARATYPE: LACM 2800 (1 dry), 241 m, W of Amatignak Id., Andreanof Ids.,
Aleutian Ids., Alaska (51°14.46'N, 179°11.84'W), Leg.: R. Baxter, 17 AUG 1986.
Remarks: Paratype LACM 2799 consists of disarticulated valves only.
calypso, Tripoplax Clark, 2008: Amer. Malac. Bull. 25(1/2):78-81, figs. 2-7.
HOLOTYPE: LACM 2882 (on SEM stub), 650-700 m, Monterey Submarine Canyon, Monterey
Co., California (36º45.163'N, 122º3.447'W), 28 SEP 1995
Remarks: Radula, girdle, and intermediate valve fragment on SEM stub, head and tail valves
and intermediate valve fragments unmounted, body in ETOH.
carolianus, Ischnochiton (Ischnochiton) Ferreira, 1984: The Veliger 26(3):179-181.
PARATYPE: LACM 2032 (1 dry), 100 m, 3 mi. S of Punta San Antonio, Bahía San Carlos, near
Guaymas, Sonora, México, Leg.: F. & R. Poorman, NOV 1980.
catalinae, Ischnochiton Willett, 1940: Bull. So. Calif. Acad. Sci. 39(3):185-186, pl. 31, fig. 2.
HOLOTYPE: LACM 1063 (dry); PARATYPES: LACM 1000 (2 dry); LACM 2146 (7 dry), 55
m, off Whites Landing, Santa Catalina Id., California Channel Ids., California, Leg.: G. Willett.
Remarks: Junior synonym of Lepidozona willetti (Berry, 1917) fide Kaas & Van Belle (1998).
Paratypes ex UCLA 22320.
clarionensis, Lepidozona (Lepidozona) Ferreira, 1983b: The Veliger 25(4):314-316.
PARATYPE: LACM 1821 (1 wet), 10 m, Bahía Sulphur, Isla Clarión, Islas Revillagigedo,
México, Leg.: A.J. Ferreira & J. Hewitt, 26 NOV 1981
cooperi, Ischnochiton Dall, 1879: Proc. U.S. Nat. Mus. 1:296.
NEOTYPE LOT: LACM 1858 (1 dry), designated by Ferreira (1978:23), The Veliger 21(1), 0 m,
intertidal, Cayucos, San Luis Obispo Co., California (35 32 N, 120 54 W), Leg.: A.J. Ferreira,
11 APR 1974.
Remarks: Currently in the genus Lepidozona fide Kaas & Van Belle (1998).
corteziana, Lepidochitona (Lepidochitona) Clark, 2000b: Nemouria 43:10-13, figs. 16-19.
HOLOTYPE: LACM 2878 (coated); PARATYPE: LACM 2879 (1 dry), 12 m, Canal de San
Lorenzo (ca. 40 km N of La Paz), Golfo de California, Baja California Sur, México (24°21'N,
111°15'W), Leg.: M. Medina-Lopez, 2 FEB 1995.
Remarks: Disarticulated valves of holotype mounted on SEM stub.
disalvoi, Rapanuia Dell’Angelo & others, 2004: The Veliger 47(2):135-140.
PARATYPES: LACM 2961 (1 dry, 1 wet), 10-20 m, off Hanga Roa, Isla de Pascua (= Easter
Id.), Leg.: B.K. Raines.
guadalupensis, Lepidozona (Lepidozona) Ferreira, 1978a: The Veliger 21(1):37-38, figs. 13-14.
PARATYPE: LACM 1776 (1 wet), Northeast anchorage, Isla Guadalupe, México (29 11 N,
118 17 W), Leg.: W.L. Lee & A.J. Ferreira, 30 DEC 1974; PARATYPES: LACM 1857 (22
wet), 0 m, intertidal, Sealer's Camp, Isla Guadalupe, México (29 01 N, 118 17 W), Leg.: W.L.
Lee & A.J. Ferreira, 1-2 JAN 1975.
laurae, Lepidozona (Lepidozona) Ferreira, 1985: The Veliger 27(4):423-425.
PARATYPE: LACM 2003 (1 wet), 73-91 m, 3.2 km SE of San Antonio, Guaymas, Sonora,
México (27 57 N, 111 06 W), Leg.: L. & C. Shy, OCT 1977
macleani, Ischnochiton (Ischnochiton) Ferreira, 1978b: Bull. So. Calif. Acad. Sci. 77(1):36-39,
figs., 1-2.
HOLOTYPE: LACM 1853 (dry); PARATYPES: LACM 1854 (8 dry), 55 m, Caleta Tagus, Isla
Isabela (= Albemarle Id.), Islas Galápagos (0 16.63 S, 91 22.73 W), Leg.: R/V Velero III, 13
JAN 1934.
Remarks: Holotype disarticulated; holotype radula and girdle scales mounted on glass slide.
macleaniana, Lepidozona (Lepidozona) Ferreira, 1985: The Veliger 27(4):425-427, figs. 6-10.
HOLOTYPE: LACM 2004 (wet), 100-104 m, off Isla San Pedro Nolasco, Sonora, México
(27 58.53 N, 111 23.62 W), Leg.: R/V Velero III, 6 FEB 1940.
Remarks: Girdle scales mounted on glass slide. Junior synonym of Lepidozona rothi Ferreira,
1983 fide Kaas & Van Bell (1998).
poppei, Ischnochiton (Haploplax) Kass & Van Belle, 1994: Monog. Living chitons 5:72.
PARATYPE: LACM 2719 (1 dry), near Nishino’omote, Kagoshima Pref., Kyushu, Japan, Leg.:
G.T. Poppe, MAR 1988.
psaltes, Dendrochiton Berry, 1963a: Leaflets in Malac., 1(22):135.
PARATYPE: LACM 1448 (1 dry), near Mission Bay entrance, San Diego, San Diego Co.,
California, Leg.: E.N. Wilson, 8 JAN 1944.
Remarks: Junior synonym of Lepidochitona (Dendrochiton) thamnopora (Berry, 1911) fide
Kaas & Van Belle (1998).
rothi, Lepidozona (Lepidozona) Ferreira, 1983a: The Veliger 25(4):316-317, figs. 19-22.
HOLOTYPE: LACM 1818 (dry), 82-91 m, Bahía Sulphur, Isla Clarión, Islas Revillagigedo,
México (18 19.7 N, 114 44.58 W), Leg.: R/V Velero III, 16 MAR 1939.
Remarks: Specimen disarticulated, radula and girdle elements mounted on glass slide.
skoglundi, Ischnochiton Ferreira, 1986b: The Veliger 28(4):449-451.
PARATYPE: LACM 2119 (1 dry), 8-15 m, off Playa Novillero, Nayarit, México (22 23 N,
105 45 W), Leg.: S. & P. Bennett, DEC 1975.
Remarks: Currently in the genus Lepidozona fide Kaas & Van Belle (1998).
stohleri, Lepidozona (Lepidozona) Ferreira, 1985: The Veliger 27(4):427-429.
PARATYPES: LACM 2005 (2 dry), 12-36 m, Isla Smith, Bahía de los Angeles, Golfo de
California, Baja California, México (29 02 N, 113 30 W), Leg.: L. & C. Shy, MAY 1975-MAY
1976.
subtilis, Lepidozona Berry, 1956: Leaflets in Malac., 1(13):74.
PARATYPES: LACM 1449 (2 dry), 0 m, intertidal, San Felipe, Golfo de California, Baja
California, México, Leg.: E.V. Coan & S.S. Berry, 26 JAN 1963.
tomhalei, Ischnochiton (Ischnochiton) Clark, 2000b: Nemouria 43:7-10, figs. 12-14.
HOLOTYPE: LACM 2877 (coated): 12 m, Punta Bajo (ca. 10 mi. N of Loreto), Golfo de
California, Baja California Sur, México (ca. 26°5'N, 112°42'W), Leg.: M. Medina-Lopez, 11
NOV 1994.
Remarks: Disarticulated valves and radula mounted on SEM stub.
tropica, Lepidochitona (Lepidochitona) Pilsbry, 1940: The Nautilus 53(3): pl. 12, fig. 4.
PARATYPE: LACM 1628 (1 dry), Bonefish Key, Monroe Co., Florida, Leg.: B.R. Bales.
Remarks: Junior synonym of Lepidochitona liozonis (Dall & Simpson, 1901) fide Kaas & Van
Belle (1998).
victoria, Ischnochiton (Ischnochiton) Ferreira, 1987: Bull. So. Calif. Acad. Sci. 86(1):42-46.
PARATYPES: LACM 2124 (2 dry),18-34 m, Roca Sucia, near Bahía Wafer, Isla del Coco, East
Pacific, Leg.: D.R. Shasky.
Remarks: Emended to Ischnochiton victoriae Ferreira, 1987 by Kaas & Van Belle (1998).
willetti, Ischnochiton Berry, 1917: Proc. Calif. Acad. Sci., 4th ser., 7(10):236-238.
PARATYPES: LACM 2159 (9 dry), 27-37 m, Forrester Id., Prince of Wales Co., Alaska, Leg.:
G. Willett, MAY 1914-AUG 1916.
Remarks: Currently in the genus Lepidozona fide Kaas & Van Belle (1998). Ex UCLA 22314.
Family Callistoplacidae Pilsbry, 1893
connellyi, Callistochiton Willett, 1937c: The Nautilus 51(1):25-26, pl. 2, fig. 13.
HOLOTYPE: LACM 1048 (dry); PARATYPES: LACM 2145 (6 dry), Punta Arbolitos, near
Ensenada, Pacific Coast Baja California, México, Leg.: Mr. & Mrs. P.M. Connelly, DEC 1956.
Remarks: Junior synonym of Callistochiton palmulatus Carpenter in Pilsbry, 1893 fide Kaas &
Van Belle (1998). Paratypes ex UCLA 22358.
leei, Callistochiton Ferreira, 1979: The Veliger 21(4):451-452.
PARATYPE: LACM 1905 (1 dry), 0 m, intertidal, Sealers' Camp, Isla Guadalupe, México
(29 01 N, 118 13 W), Leg.: A.J. Ferreira & W.L. Lee, 2 JAN 1975.
portobelensis, Callistochiton Ferreira, 1976: The Nautilus 90(1):46-49.
PARATYPE: LACM 1767 (1 dry), 0.3-3 m, intertidal, 5 km W of Portobelo, Colon Prov.,
Panamá (9 30 N, 79 42 W), Leg.: A.J. Ferreira, 15 FEB 1975.
Family Chaetopleuridae Plate, 1899
euryplax, Chaetopleura (Pallochiton) Berry, 1945: Amer. Mid. Nat. 34(2):491-492.
PARATYPE: LACM 1263 (1 dry), Bahía de Adair, Sonora, México, Leg.: E.C. Huffman, 1937.
Remarks: Junior synonym of Chaetopleura (Pallochiton) mixta (Dall, 1919) fide Kaas & Van
Belle (1998).
roddae, Chaetopleura (Chaetopleura) Ferreira, 1983a: The Veliger 25(3):214-216.
PARATYPES: LACM 1977 (3 wet), 0 m, intertidal, Salinas, Guayas Prov., Ecuador (2 12 S,
80 58 W), Leg.: J.H. McLean, 5-6 MAR 1970.
Remarks: Emended to Chaetopleura roddai Ferreira, 1982 by Kaas & Van Belle, 1978.
shyana, Chaetopleura (Chaetopleura) Ferreira, 1983a: The Veliger 25(3):219-220.
PARATYPE: LACM 1978 (1 dry). Isla de Pata, Bahía de los Angeles, Golfo de California, Baja
California, México (29 00 N, 113 30 W), Leg.: L. Shy, MAY 1976.
Family Chitonidae Rafinesque, 1815
arenosa, Acanthopleura Ferreira, 1986a: The Veliger 28(3):255-258.
PARATYPES: LACM 2107 (2 wet), Pebbly Beach (ca. 25 km S of Port Douglas), Queensland,
Australia (16 35 S, 145 22 E), Leg.: A.J. Ferreira & S. Motley, 14 AUG 1981.
Remarks: Currently in the genus Liolophura fide Kaas & Van Belle (1998).
lineatus, Chiton Wood, 1815: Gen. Conch. p.15, pl. 2, figs. 4-5.
NEOTYPE: LACM 2734 (1 dry), designated by Clark (1999, figs. 1-8), Amer. Malac. Bull.
15(1):34. Old Sitka (Starrigaven Bay), 10 km N of Sitka, Baranof Id., Alexander Archipelago,
Sitka Co., Alaska (57 08'N, 135 55'W).
Remarks: Currently in the genus Tonicella fide Kaas & Van Belle (1998).
linsleyi, Rhyssoplax Burghardt, 1973: Proc. Calif. Acad. Sci., 4th ser., 39(21):501-506.
PARATYPES: LACM 1620 (2 dry), Pu‘u o Hulu Beach, S of Ma‘ili, Wai‘anae Dist., O‘ahu,
Hawaiian Ids., Leg.: Mrs. H. Gudnason, 26 FEB 1965
Remarks: Currently in the genus/subgenus Chiton (Rhyssoplax) fide Kaas & Van Belle (1998).
mertensii, Chiton Middendorff, 1847: Bull. Acad. Imp. Sci. St. Petersburg, 6(8):118.
NEOTYPE: LACM 1855 (1 dry), designated by Ferreira (1978:20), The Veliger 21(1),
NEOTYPE LOT: LACM 1856 (10 wet), 0 m, intertidal, 0.5 mi. S of Fort Ross, Sonoma Co.,
California (38 30.7 N, 123 14 W), Leg.: J.H. McLean, 28 DEC 1963.
Remarks: Currently in the genus Lepidozona fide Kaas & Van Belle (1998). Radula mounted on
glass slide.
Suborder Acanthochitonina Bergenhayn, 1930
Superfamily MOPALIOIDEA Dall, 1889
Family Tonicellidae Simroth, 1894
callisto, Micichiton Clark, 2002: Nemouria 46: 2-5, figs. 1-6.
HOLOTYPE: LACM 2884 (dry valves), 10 m, E side entrance to Crescent Bay, NW end of Atka
Id., Andreanof Ids., Aleutian Ids., Alaska (52°2'N, 174°14'W), Leg.: R.N. Clark, 8 JUL 1997.
lokii, Tonicella Clark, 1999: Amer. Malac. Bull. 15(1):39-41, figs. 17-24.
HOLOTYPE: LACM 2623 (wet), PARATYPES: LACM 2624-2625, 0-13 m, Coast Guard
breakwater, Monterey Bay, Monterey Co., California (36 45'N, 122 55'W).
venusta, Tonicella Clark, 1999: Amer. Malac. Bull. 15(1):41-45, figs. 25-32, 34.
HOLOTYPE: LACM 2626 (dry); PARATYPES: LACM 2627-2628 (dry), 1-10 m, Mountain Pt.,
8 km S of Ketchikan, Revillagigedo Id., Alexander Archipelago, Sitka Co., California
(55 17'35″N, 131º32'20″W).
Family Mopaliidae Dall, 1889
ciliata, Mopalia (Sowerby, 1840): Mag. Nat. Hist., n.s. 4:289.
NEOTYPE: LACM 2885 (wet), designated by Clark (2008, figs. 1-10), Amer. Malac. Bull.
25(1/2):75, Whites Pt., Los Angeles Co., California (33º42.8'N, 118º8.3'W), Leg.: S.R. Thorpe,
Jr., 11 DEC 1958.
egretta, Mopalia Berry, 1919a: Proc. Calif. Acad. Sci., 4th ser., 9(1):13-17.
PARATYPES: LACM 2161 (2 dry), 37 m, Forrester Id., Prince of Wales Co., Alaska, Leg.: G.
Willett, 5 AUG 1915.
Remarks: Ex UCLA 22411.
ferreirai, Mopalia Clark, 1991: The Veliger 34(3):309-313, figs. 1-5, 12.
HOLOTYPE: LACM 2329 (dry); PARATYPES: LACM 2330 (2 dry), 0.5-1 m, Rotary Beach, 5
km SE of Ketchikan, Revillagigedo Id., Ketchikan Co., Alaska, (55 16 N, 131 34 N), Leg.: R.N.
Clark, 17 AUG 1986.
Remarks: Holotype radula in 70% ethyl alcohol.
hanselmani, Placiphorella Clark, 1994: The Veliger 34(3):304-308, figs. 23-25, 36-37.
HOLOTYPE: LACM 2707 (dry); PARATYPE: LACM 2708 (1 dry), intertidal, Isla Piojo, Bahía
de los Angeles, Golfo de California, Baja California, México (28 57 N, 113 32 W), Leg.: C. &
L. Shy, May 1976.
kamehamehae, Plaxiphora (Plaxiphora) Ferreira & Bertsch, 1979: Trans. San Diego Soc. Nat.
Hist. 19(7):75-84.
PARATYPES: LACM 1952 (2 wet), 0 m, intertidal, Ft. Kamehameha Beach, Honolulu Dist.,
O‘ahu, Hawaiian Ids. (21 19 N, 57 57 W).
kennerleyi, Mopalia Carpemter, 1864: Rep. Brit. Ass. Adv. Sci. 1863:648.
NEOTYPE: LACM 2886 (wet), designated by Clark (2008, figs. 2-6), Amer. Malac. Bull.
25(1/2):73. Tacoma Narrows, “Puget Sound,” Pierce Co., Washington (47º16'N, 122º31'W),
Leg.: S.R. Thorpe, Jr., 1 JUL 1965.
mirabilis, Placiphorella Clark, 1994: The Veliger 37(3):303-304, figs. 20-22, 34-35.
HOLOTYPE: LACM 2703 (wet), 69 m, 1.2 mi. 130 deg. T from Long Pt., Santa Catalina Id.,
California Channel Ids., California (33 23.5 N, 115 13.3 W), Leg.: R/V Velero IV, 13 FEB 1965;
PARATYPE: LACM 2704 (1 dry), 82-91 m, E of Long Pt., Santa Catalina Id., California
Channel Ids., California (33 23.7 N, 118 20.07 W), Leg.: R/V Velero III, 10 AUG 1941;
PARATYPE: LACM 2705 (1 wet), 66 m, 1.25 mi. E of White Cove, Santa Catalina Id.,
California Channel Ids., California (33 23.54 N, 118 20.9 W), Leg.: R/V Velero IV, 31 OCT
1948; PARATYPE: LACM 2706 (2 wet), 82 m, 0.4-0.7 mi. 110 to 132 deg.T from Ship Rock,
Santa Catalina Id., California Channel Ids. (33 23.5 N, 118 13.3 W), Leg.: R/V Velero IV, 13
FEB 1965.
Remarks: Incorrect longitude cited by Clark for holotype, correct longitude = 118 20.80 W.
pedroana, Mopalia Willett, 1932a: The Nautilus 45(3):101-102, pl. 7, fig. 1.
HOLOTYPE: LACM 1025 (dry), 27 m, off San Pedro, Los Angeles Co., California, Leg.: G.
Willett, AUG 1930.
Remarks: Junior synonym of Mopalia acuta (Carpenter, 1855) fide Kaas & Van Belle (1998).
rufa, Placiphorella (Placiphorella) Berry, 1917: Proc. Calif. Acad. Sci., 4th ser., 7(10):241-244.
PARATYPES: LACM 2160 (11 dry), 27-46 m, Forrester Id., Prince of Wales Co., Alaska,
Leg.: G. Willett, JUL 1914-1916.
Remarks: Ex UCLA 22446.
spectabilis, Mopalia Cowan & Cowan, 1977: Syesis 10:46-52.
PARATYPES: LACM 2051 (2 wet), Mayne Id., Navy Channel, Georgia Strait, Capital Dist.,
British Columbia, Canada, Leg.: G.I.McT. Cowan & I.McT.Cowan, JUN 1970.
Superfamily CRYPTOPLACOIDEA H.Adams & A. Adams, 1858
Family Acanthochitonidae Pilsbry, 1893
burhardtae, Acanthochitona Clark, 2000b: Nemouria 43:13-15, figs. 21, 23-24.
HOLOTYPE: LACM 2880 (coated); PARATYPE: LACM 2881 (1 dry), 12 m, Canal de San
Lorenzo (ca. 40 km N of La Paz), Golfo de California, Baja California México (24°21'N,
111°15'W), Leg.: M. Medina-Lopez, 2 FEB 1995.
Remarks: Disarticulated valves of holotype on SEM stub.
shaskyi, Acanthochitona Ferreira, 1987: Bull. So. Calif. Acad. Sci., 86(1):47-52.
PARATYPES: LACM 2125 (2 wet), 46-69 m, Bahía Chatham, Isla del Coco, East Pacific
(5 33 N, 87 02 W), Leg.: D.R. Shasky.
Remarks: Junior synonym of Acanthochitona angelica Dall, 1919 fide Kaas & Van Belle
(1998).
CLASS SCAPHOPODA Bronn, 1862
Order Dentaliida Da Costa, 1834
Family Dentaliidae Gray, 1834
actiniophorum, Fissidentalium Shimek, 1997: The Veliger 40(2):181-189, fig. 3.
HOLOTYPE: LACM 2792 (shell dry, animal wet), 4100 m, 226 km W of Pt. Arguello, Santa
Barbara Co., California (34 44 N, 123 12 W), Leg.: K.L. Smith, R/V New Horizon, 4 NOV
1993; PARATYPE: LACM 2793 (shell dry, animal wet), 4100 m, 226 km W of Pt. Arguello,
Santa Barbara Co., California (34 42 N, 123 08 W), Leg.: K.L. Smith, R/V New Horizon, 7
NOV 1993; PARATYPES: LACM 2809 (6 dry shells, animals wet), 4100 m, 226 km W of Pt.
Arguello, Santa Barbara Co., California (34 45 N, 123 02 W), Leg.: K.L. Smith, R/V New
Horizon, 24 FEB 1993; PARATYPES: LACM 2810 (4 dry shells, animals wet), 4100 m, 226 km
W of Pt. Arguello, Santa Barbara Co., California (34 44 N, 123 12 W), Leg.: K.L. Smith, R/V
New Horizon, 4 NOV 1993; PARATYPES: LACM 2811 (2 dry shells, animals wet), 4100 m,
226 km W of Pt. Arguello, Santa Barbara Co., California (34 41 N, 123 05 W), Leg.: K.L.
Smith, R/V New Horizon, 5 FEB 1994.
brevicornu, Dentalium (Compressidens) Sharp & Pilsbry, 1897: Tryon's Manual of Conchology,
1st ser., 17:125.
PARATYPE: LACM 1757 (1 dry), 1159 m, Islas Galápagos, Leg.: U.S. Fish Commission.
Remarks: Currently in the genus Compressidens fide Steiner & Kabat (2004).
erosum, Fissidentalium Shimek & Moreno, 1996: The Veliger 39(1):77-81, figs. 2A-B.
HOLOTYPE: LACM 2755 (shell dry, animal wet); PARATYPE: LACM 2756 (shell dry, animal
wet), 3090-3300 m, Pioneer Canyon, WSW of Pigeon Pt., San Mateo Co., California, Leg.: G.
Moreno, 29 FEB 1992.
hancocki, Dentalium Emerson, 1956: Amer. Mus. Novitates, 1787:3-6, fig. 1.
HOLOTYPE: LACM 1758 (dry); PARATYPE: LACM 1759 (1 dry), 9-33 m, 0.5 mi. SE of
Punta Hughes, Bahía Santa Maria, Pacific Coast Baja California Sur, México (24 44.5 N,
112 14.90 W), Leg.: R/V Velero IV, 4 APR 1949; PARATYPES: LACM 2887 (2 dry), 20.1 m,
off Punta Penasco, Sonora, México (31°18.70'N, 113°38.17'W), R/V Velero III, 3 FEB 1940;
PARATYPE: LACM 2888 (1 dry), 36.5 m, 2.25 mi. 162 deg. T from Cabo Pulmo, Golfo de
California, Baja California Sur, México (23°23.90'N, 109°24.30'W), Leg.: R/V Velero III, 13
MAR 1949; PARATYPE: LACM 2889 (1 dry), 36.5 m, Bahía Agua Verde, Golfo de California,
Baja California Sur, México (25°31.0'N, 111°4.0'W), Leg.: R/V Velero III, 18 MAR 1936.
Remarks: Currently in the genus Tesseracme fide Steiner & Kabat (2004). Holotype and
paratype LACM 1759 ex AHF 1787-49.
megathyris, Dentalium Dall, 1890: Proc. U.S. Nat. Mus. 12(773):293.
PARATYPE: LACM 1816 (1 dry), 1485 m, Islas Galápagos, Leg.: U.S. Fish Commission.
Remarks: Currently in the genus Fissidentalium fide Steiner & Kabat (2004).
numerosum, Dentalium oerstedii Dall in Pilsbry & Sharp, 1897: Tryon's Manual of
Conchology, 1st ser., 17:25.
PARATYPE: LACM 1755 (1 dry), 80 m, near Isla Cedros, Pacific coast Baja California, México
(28 12 N, 115 09 W), Leg.: U.S. Fish Commission.
Remarks: Junior synonym of Dentalium oerstedii Mörch, 1861 fide Steiner & Kabat (2004).
panamense, Dentalium megathyris Dall, 1908: Bull. Mus. Comp. Zool. 43(6):357-358.
PARATYPE: LACM 1836 (1 dry), 3241 m, Isla Malpelo, Colombia, Leg.: U.S. Fish
Commission.
Remarks: Junior synonym of Fissidentalium megathyris (Dall, 1889) fide Steiner & Kabat
(2004).
Family Rhabdidae Chistikov, 1975
dalli, Dentalium (Rhabdus) Pilsbry & Sharp, 1897: Tryon's Manual of Conchology, 1st ser.,
17:114-115.
PARATYPE: LACM 1756 (1 dry), 642 m, N of Unalaska Id., Aleutian Ids. Co., Alaska, Leg.:
U.S. Fish Commission.
Remarks: Currently in the genus Rhabdus fide Steiner & Kabat (2004).
Order Gadilida Starobogatov, 1977
Family Pulsellidae Scarabino in Boss, 1982
salishorum, Pulsellum Marshall, 1980: The Veliger 23(2):149-152.
PARATYPES: LACM 1929 (4 dry), 27 m, East Sound, Orcas Id., San Juan Ids., San Juan Co.,
Washington, Leg.: E. Marshall; R/V Caracoles, 16 AUG 1977.
Family Gadilidae Stoliczka, 1868
macleani, Cadulus (Platyschides) Emerson, 1978: The Nautilus 92(3):122, figs. 1-3.
HOLOTYPE: LACM 1886 (dry); PARATYPES: LACM 1887 (3 dry),18-55 m, Bahía de los
Muertos, Golfo de California, Baja California Sur, México (23 55 N, 109 46 W), Leg.: J.H.
McLean, P. Oringer, & L. Marincovich, 8 APR 1966.
Remarks: Currently in the genus Cadulus fide Steiner & Kabat (2004).
nedallisoni, Cadulus (Polyschides) Emerson, 1978: The Nautilus 92(3):120-122.
PARATYPES: LACM 2117 (2 dry), 22-24 m, Corinto, Chinandega Dept., Nicaragua
(12 28.05 N, 87 12.65 W), Leg.: Zaca Expedition, 5 JAN 1938.
Remarks: Currently in the genus Polyschides fide Steiner & Kabat (2004).
splendens, Dischides Raines, 2002a: La Conchiglia 34(304):37-38, fig. 47.
HOLOTYPE: LACM 2949 (dry); PARATYPE: LACM 2950 (1 dry), 50-80 m, off Tahai, Isla de
Pascua (= Easter Id.), Leg.: B.K. Raines.
CLASS GASTROPODA Cuvier, 1791
Clade Patellogastropoda Lindberg, 1986
Superfamily LOTTIOIDEA Gray, 1840
Family Acmaeidae Forbes, 1850
arcuata, Pectinodonta Dall, 1882: Proc. U.S. Nat. Mus. 4(246):409-410.
PARATYPE: LACM 1426 (1 dry), 413 m, off Santa Lucia, St. Thomas, U.S. Virgin Ids.
aupouria, Pectinodonta Marshall, 1985a: New Zealand Jour. Zool. 12(2):277-279.
PARATYPES: LACM 2763 (3 wet), 1075-1100 m, off White Bay, Bay of Plenty, North Id., New
Zealand (37 23.7 S, 177 39.5 E), Leg.: O.M. Moore & P.J. McMillan, R/V Kalinovo, 23 NOV
1981.
kapalae, Pectinodonta Marshall, 1985a: New Zealand Jour. Zool. 12(2):280.
PARATYPE: LACM 2765 (1 wet), 941-978 m, off Ulladulla, New South Wales, Australia
(35 30 S, 150 52 E), Leg.: R/V Kapalae, 8 SEP 1983.
morioria, Pectinodonta Marshall, 1985a: New Zealand Jour. Zool. 12(2):279-280.
PARATYPES: LACM 2764 (3 wet), 800-810 m, Chatham Rise, NE of Mernoo Bank, SE of
Cape Palliser, North Wellington, North Id., New Zealand (42 44 S, 176 07 E), Leg.: P. Grimes,
R/V Kaltan, 28 SEP 1982.
Family Lepetidae Gray, 1850
curumim, Propilidum Leal & Simone, 1998: Bull. Mar. Sci. 63(1):158-164.
PARATYPES: LACM 2806 (6 wet; 5 dry), 166 m, off Chuí, Rio Grande do Sul State, Brazil,
Leg.: R/V Wladimir Besnard, JAN 1972.
lindbergi, Iothia McLean, 1985a: The Veliger 27(3):336-337, fig. 1.
HOLOTYPE: LACM 2063 (dry), PARATYPES: LACM 2064 (2 dry), 183 m, 6 mi. W of Pt.
Piños, Monterey Co., California (36 39 N, 122 01 W), Leg.: J.H. McLean, Hopkins Marine Lab.,
R/V Tage, 2 MAR 1961.
lissocona, "Cocculina" Dall, 1927: Proc. U.S. Nat. Mus. 70:110.
PARALECTOTYPE: LACM 2741 (1 dry), designated by McLean & Harasewych (1995:31),
Nat. Hist. Mus. L.A. Co. Cont. Sci. 453.
538 m, off Fernandina, Nassau Co., Florida.
Family Lottiidae Gray, 1840
acutapex, “Acmaea” Berry, 1960: Leaflets in Malac. 1(19):117.
PARATYPES: LACM 1876 (4 dry), Punta Cholla, W of Puerto Peñasco, Golfo de California,
Sonora, México, Leg.: S.S. Berry, 23-24 MAR 1948.
Remarks: Currently in the genus Lottia.
albomaculata, Lottia gigantea Dall, 1910: The Nautilus 24(8):96.
HOLOTYPE: LACM 1953 (dry), Laguna Beach, Orange Co., California.
Remarks: Junior synonym of Lottia gigantea Sowerby, 1834.
apicina, Acmaea Dall, 1879: Proc. U.S. Nat. Mus. 1:341-342.
PARATYPES: LACM 1425 (2 dry), E side of Simeohoff Id., Shumagin Ids., Aleutian Ids. Co.,
Alaska
Remarks: Currently in the genus Erginus.
borealis, Collisella Lindberg, 1982: Wassmann Jour. Biol. 40(1-2):52-56.
PARATYPES: LACM 2655 (4 wet), S side Gibson Id., Chichagof Harbor, Attu Id., Aleutian Ids.
Co., Alaska (52 57 N, 173 16 W), Leg.: D.R. Lindberg, 6 AUG 1979.
Remarks: Currently in the genus/subgenus Lottia (Collisella).
chamororum, Patelloida Lindberg & Vermeij, 1985: The Veliger 27(4):412-416.
PARATYPES: LACM 2611 (2 dry), Asanite Bay, Guam (13 20 N, 144 46 E), Leg.: G.J.
Vermeij, 10 JUL 1970.
immaculata, Notoacmaea Lindberg & McLean, 1981: Proc. Calif. Acad. Sci., 4th ser., 42(12):
331-332, fig. 14.
HOLOTYPE: LACM 1923 (dry), 0-3 m, intertidal, Caleta del Norte, W side Isla Baltra, off N
end, Isla Santa Cruz (= Indefatigable Id.), Islas Galápagos (0 26 S, 90 17 W), Leg.: R/V Anton
Bruun, 21 SEP 1966
Remarks: Currently in the genus Tectura. Radula mounted on glass slide.
mimica, Lottia (Lottia) Lindberg & McLean, 1981: Proc. Calif. Acad. Sci., 4th ser., 42(12):
332-334, figs. 17-19.
HOLOTYPE: LACM 1925 (shell dry, animal wet), 0 m, intertidal, Isla Coamano, Bahía
Academy, Isla Santa Cruz (= Indefatigable Id.), Islas Galápagos (0 38 S, 90 23 W), Leg.: J.
DeRoy, OCT 1967; PARATYPES: LACM 1926 (16 dry; 18 wet), Bahía Academy, Isla Santa
Cruz (= Indefatigable Id.), Islas Galápagos, Leg.: J. DeRoy, 1967-1969.
Remarks: Holotype radula mounted on glass slide.
morchii, Acmaea scabra var. Dall, 1878: Proc. U.S. Nat. Mus. 1(18):47.
PARATYPES: LACM 1956 (6 dry), Tomales, Marin Co., California, Leg.: H. Hemphill.
Remarks: Junior synonym of Lottia (Colisella) limatula (Carpenter, 1864).
pumila, Notoacmaea Lindberg & McLean, 1981: Proc. Calif. Acad. Sci., 4th ser., 42(12):
328-329, figs. 8-9.
HOLOTYPE: LACM 1919 (dry); PARATYPES: LACM 1920 (58 dry, 2 wet), 0 m, intertidal,
NW side Punta Ancón, Peninsula de Santa Elena, Esmeraldas Prov., Ecuador (2 19.5 S,
80 54 W), Leg.: J.H. McLean, 6 MAR 1970.
Remarks: Currently in the genus Tectura.
puniceus, Erginus (Erginus) Lindberg, 1988: PaleoBios 12(46):3-4, pl. 1, figs. B-D.
HOLOTYPE: LACM 2178 (dry); PARATYPES: LACM 2179 (34 dry), 0 m, intertidal, 2.5 km N
of Tomari, Aomori Pref., Honshu, Japan, Leg.: R.F. Bolland, 16 APR 1978
rothi, Notoacmaea Lindberg & McLean, 1981: Proc. Calif. Acad. Sci., 4th ser., 42(12):329-331,
fig. 11.
HOLOTYPE: LACM 1921 (shell dry, animal wet); PARATYPE: LACM 1922 (1 dry),
0 m, intertidal, Bahía Wreck, Isla San Cristobal (= Chatham Id.), Islas Galápagos (0 54 S,
89 36 W),
Leg.: J. DeRoy, 12 MAY 1968
Remarks: Currently in the genus Tectura. Holotype radula mounted on glass slide.
smithi, Lottia (Lottia) Lindberg & McLean, 1981: Proc. Calif. Acad. Sci., 4th ser., 42(12):
334-337, figs. 20-22.
HOLOTYPE: LACM 1927 (1 dry), 0 m, intertidal, Punta Nuñez, Bahía Academy, Isla Santa
Cruz (= Indefatigable Id.), Islas Galápagos (0 38 S, 90 23 W), Leg.: J. DeRoy, 13 OCT 1969;
PARATYPES: LACM 1928 (9 dry, 13 wet animals), Isla Santa Cruz (= Indefatigable Id.), Islas
Galápagos.
Remarks: Radulae mounted on glass slides.
stanfordiana, Acmaea Berry, 1957: Leaflets in Malac., 1(14):76-77.
PARATYPES: LACM 1012 (3 dry), Punta Peñasco, Golfo de California, Sonora, México,
Leg.: F.B. Howard, FEB 1957.
Remarks: Currently in the genus Lottia.
ubiquita, Notoacmaea Lindberg & McLean, 1981: Proc. Calif. Acad. Sci., 4th ser., 42(12):
326-328, figs. 5-7.
HOLOTYPE: LACM 1917 (dry); PARATYPES: LACM 1918 (2 dry), 0.0-5 m, Playa las Hadas,
S side Peninsula de Santiago, Bahía Manzanillo, Colima, México (19 06.6 N, 104 21.7 W),
Leg.: J.H. McLean & C. Tenney, 21-24 MAR 1963.
Remarks: Currently in the genus Tectura. Radulae mounted on glass slides.
Superfamily NEOLEPETOPSOIDEA McLean, 1990
Family Neolepetopsidae McLean, 1990
clementensis, Paralepetopsis McLean, 2008: Jour Shellfish Res. 27(1):18-19, figs. 2B, 3B, 3E.
HOLOTYPE: LACM 3092 (shell dry, animal wet); PARATYPE: LACM 3093 (dry), 1800 m,
San Clemente Seep, SW of San Diego, San Diego Co., California (32 12 N, 117 44 W), Leg.:
R/V Alvin, C.R. Smith, 29 MAR 2000.
Remarks: Radula of holotype mounted on glass slide.
densata, Neolepetopsis McLean, 1990b: Jour. Zool. Soc., London 222:496-499, pl. 3, figs. a-d.
HOLOTYPE: LACM 2405 (shell dry, animal wet); PARATYPES: LACM 2406 (1 dry, 1 wet, 1
gold coated), 2725 m, East Pacific Rise, East Pacific (11 46 N, 103 47 W), Leg.: R/V Alvin, 25
MAR 1988.
floridensis, Paralepetopsis McLean, 1990b: Jour. Zool. Soc., London 222:510-513, pl. 10, figs.
a-c.
HOLOTYPE: LACM 2420 (dry); PARATYPES: LACM 2421 (2 dry, 5 wet); PARATYPES:
LACM 2422 (6 dry, 13 wet); PARATYPES: LACM 2423 (1 dry, 6 wet, 1 gold coated), 3270 m,
Florida Escarpment, off SW Florida (26 03 N, 84 54 W), Leg.: R/V Alvin, 9 MAR 1984.
Remarks: Radula of paratype LACM 2422 mounted on glass slide.
gordensis, Neolepetopsis McLean, 1990b: Jour. Zool. Soc., London 222:492-496, pl. 1, figs. a-c.
HOLOTYPE: LACM 2401 (shell dry, animal wet); PARATYPE: LACM 2400 (shell dry, animal
wet); PARATYPES: LACM 2402 (2 dry, 5 wet, 1 gold coated); PARATYPES: LACM 2403 (2
dry, 6 wet, 2 gold coated); PARATYPES: LACM 2404 (3 dry, 4 wet), 3200-3250 m, Escanaba
Trough, Gorda Ridge off California, East Pacific (41 00 N, 127 29 W), Leg.: R/V Alvin, 10 JUN
1988.
Remarks: Radulae of paratypes LACM 2402 & LACM 2404 mounted on glass slides.
nicolasensis, Neolepetopsis McLean, 2008: Jour. Shellfish Res. 27(1):16-17, figs. 1C, , 3A, 3C.
HOLOTYPE: LACM 3089 (shell dry, animal wet); PARATYPE: LACM 3090 (dry), 960 m, NW
of San Nicolas Id., California Channel Ids., California (33 20.35 N, 119 58.85 W), Leg.: C.R.
Smith, 30 APR 1995.
Remarks: Radula of paratype mounted on glass slide.
occulta, Neolepetopsis McLean, 1990b: Jour. Zool. Soc., London 222:501-503, pl. 6, figs. a-c.
HOLOTYPE: LACM 2409 (shell dry, animal wet); PARATYPES: LACM 2410 (4 dry, 13 wet, 1
gold coated), 1990 m, Green Seamount, East Pacific (20 49 N, 109 17 W), Leg.: R/V Alvin, 5
FEB 1982.
tunnicliffae, Paralepetopsis McLean, 2008: Jour. Shellfish Res. 27(1):17-18, figs. 2A, 3D.
HOLOTYPE: LACM 3091 (dry), 2145 m, Clam Bed, Chowder Hill, Middle Segment, Juan de
Fuca Ridge, East Pacific (48 27.50 N, 128 42.50 W), Leg.: V. Tunnicliffe, 26 JUN, 1992.
Remarks: Radula of holotype mounted on glass slide.
verruca, Neolepetopsis McLean, 1990b: Jour. Zool. Soc., London 222:499-501, pl. 5, figs. a-d.
HOLOTYPE: LACM 2407 (shell dry, animal wet); PARATYPE: LACM 2408 (1 dry), 2595 m,
East Pacific Rise, East Pacific (20 51 N, 109 04.9 W), Leg.: R/V Alvin, 22 APR 1979.
vitrea, Eulepetopsis McLean, 1990b: Jour. Zool. Soc., London 222:503-509, pl. 7, figs. a-d;
pl. 8, figs. d-g.
HOLOTYPE: LACM 2417 (dry); PARATYPES: LACM 2418 (4 dry, 23 wet), 2618 m, East
Pacific Rise, East Pacific (20 50 N, 109 06 W), Leg.: R/V Alvin, 9 MAY 1982; PARATYPES:
LACM 2411 (3 dry), 2615 m, East Pacific Rise, East Pacific (20 50 N, 109 06 W), Leg.: R/V
Alvin, 17 APR 1982; PARATYPES: LACM 2412 (3 dry), 2617 m, East Pacific Rise, East Pacific
(20 50 N, 109 06 W), Leg.: R/V Alvin, 19 APR 1982; PARATYPES: LACM 2413 (5 dry, 3
wet), 2633 m, East Pacific Rise, East Pacific (20 50 N, 109 06 W), Leg.: R/V Alvin, 20 APR
1982; PARATYPES: LACM 2414 (16 wet), 2612 m, East Pacific Rise, East Pacific (20 50 N,
109 06 W), Leg.: R/V Alvin, 25 APR 1982; PARATYPES: LACM 2415 (27 wet), 2618 m, East
Pacific Rise, East Pacific (20 50 N, 109 06 W), Leg.: R/V Alvin, 4 MAY 1982; PARATYPES:
LACM 2416 (8 dry, 6 wet), 2616 m, East Pacific Rise, East Pacific (20 50 N, 109 06 W), Leg.:
R/V Alvin, 7 MAY 1982; PARATYPES: LACM 2419 (4 dry, 6 wet), 2616 m, East Pacific Rise,
East Pacific (20 50 N, 109 06 W), Leg.: R/V Alvin, 10 MAY 1982, PARATYPE: LACM 2475
(1 dry, 1 wet), 2617 m, East Pacific Rise, East Pacific (20 50 N, 109 `06 W), Leg.: R/V Alvin, 26
APR 1982.
Clade Vetigastropoda Salvini-Plawen, 1987
Superfamily PLEUROTOMARIOIDEA Swainson, 1840
Family Pleurotomariidae Swainson, 1840
maureri, Perotrochus Harasewych & Askew, 1993: The Nautilus 106(4):130-136.
PARATYPES: LACM 2629 (3 wet),198-213 m, 80 naut. mi. E of Charleston, Charleston Co.,
South Carolina (32 43.90 N, 78 06.00 W), Leg.: Askew, Slater, R/V Nekton Delta, 2 MAY
1987.
Superfamily HALIOTOIDEA Rafinesque, 1815
Family Haliotidae Rafinesque, 1815
guadalupensis, Haliotis (Sulculus) fulgens Talmadge, 1964: Trans. San Diego Soc. Nat. Hist.
13(18):375.
PARATYPE: LACM 1210 (1 dry), S tip Morro Sur, Isla Guadalupe, Pacific Coast Baja
California, México, Leg.: R. Clutter, P. Taylor, & C. Boyd, 30 JAN 1960.
Remarks: Junior synonym of Haliotis fulgens Philippi, 1845 fide Geiger & Poppe (2000).
oweni, Haliotis (Sulculus) corrugata Talmadge, 1966: L.A. Co. Mus. Nat. Hist. Cont. Sci.
109:1-4, figs. 1-2.
HOLOTYPE: LACM 1147 (dry); PARATYPES: LACM 1148 (3 dry), 6 m, W anchorage, Isla
Guadalupe, Pacific Coast Baja California, México, Leg.: R.S. Owen, OCT 1965.
Remarks: Junior synonym of Haliotis corrugata Wood, 1828 fide Geiger & Poppe (2000).
patamkanthini, Haliotis Dekker, Regter, & Gras, 2001: Of Sea & Shore 23(4):199-200.
PARATYPE: LACM 2753, (1 dry), 10-20 m, Rahcha Ids., Andaman Sea, Thailand (7°36'N,
98°20'E), Leg.: S. Patamakanthin.
roberti, Haliotis (Padollus) McLean, 1970a: Malac. Review 2:115-116, figs. 1-2.
HOLOTYPE: LACM 1368 (shell dry, animal wet); PARATYPES: LACM 1369 (5 dry shells,
animals wet), 73-128 m, Bahía Chatham, Isla del Coco, East Pacific (5 33.8 N, 86 58.92 W),
Leg.: R/V Velero III, 14 JAN 1938.
splendidula, Haliotis (Sulculus) cracherodii Williamson, 1892: Proc. U.S. Nat. Mus. 15(898):
198.
HOLOTYPE: LACM 1035 (dry); PARATYPE: LACM 1687 (1 dry), Pt. Vicente, Palos Verdes
Peninsula, Los Angeles Co., California, Leg.: Mrs. M.B. Williamson.
Remarks: Junior synonym of Haliotis cratcherodii Leach, 1814 fide Geiger & Poppe (2000).
Superfamily SCISSURELLOIDEA Gray, 1847
Family Scissurellidae Gray, 1847
alto, Scissurella Geiger, 2003: Moll. Res 23(1):27-29, figs1-2.
HOLOTYPE: LACM 2914 (coated); PARATYPE: LACM 2915, 10-20 m, off Punta Rosalia, E
of Anakena, Isla de Pascua (= Easter Id.), Leg.: B.K. Raines, DEC 2000.
carolarum, Sinezona Geiger & McLean, 2010: Zootaxa 2356:8-11.
PARATYPES: LACM 3116 (5 dry), 12-15 m, Île de Clipperton, Eastern Pacific (10 18.17 N,
109 11.52 W), Leg.: K.L. Kaiser; M/V Royal Star, 18 APR 1994.
demispira, Coronadoa Geiger & McLean, 2010: Zootaxa 2356:12-15, fig. 9.
HOLOTYPE: LACM 3113 (dry, coated); PARATYPES: LACM 31143115, 26-40 m,
Farnsworth Bank, S of Santa Catalina Id., California Channel Islands, California (33 21 N,
118 31 W), Leg.: J.H. McLean
epicharis, Scissurella McLean, 1970a: Malac. Review 2:117, fig. 3.
HOLOTYPE: LACM 1370 (dry, lost); PARATYPE: LACM 1371 (1 dry), 22 m, Bahía Cartago,
Isla Isabela (= Albemarle Id.), Islas Galápagos (0 35 N, 90 57 W), Leg.: R/V Velero III, 21 JAN
1938.
Remarks: Currently in the genus Anatoma.
euripes, Temnocinclis McLean, 1989b: Nat. His. Mus. L.A. Co. Cont. Sci. 407:7-9, figs. 1D-F;
4A.
PARATYPE: LACM 2351 (1 wet), 2213 m, Axial Seamount, Endeavor Segment, Juan de Fuca
Ridge off Washington, East Pacific (47 57.0 N, 129 06.0 W), Leg.: R/V Alvin, 3 SEP 1984;
PARATYPE: LACM 2352 (shell dry, animal wet), 1575 m, hydrothermal vents of Axial
Seamount, Juan de Fuca Ridge off Washington, East Pacific (45 59.5 N, 130 03.5 W), Leg.: R/V
Pisces IV, 27 JUL 1986; PARATYPE: LACM 2353 (shell dry, animal wet), Hydrothermal vents
of southern Juan de Fuca Ridge off Oregon, East Pacific (44 39.2 N, 130 22.0 W), Leg.: R/V
Alvin, 27 JUL 1986.
Remarks: Shell of paratype LACM 2351 fragmented, body sectioned (female).
exorum, Depressizona Geiger, 2003: Moll. Res. 23(1):51-52, fig. 14.
HOLOTYPE: LACM 2932 (coated); PARATYPE: LACM 2933, 10-20 m, off Punta Rosalia, E
of Anakena, Isla de Pascua (= Easter Id.), Leg.: B.K. Raines, DEC 2000.
hawaiiensis, Sinezona Geiger & McLean, 2010: Zootaxa 2356:8, fig. 4.
HOLOTYPE: LACM 2466 (dry, coated): 27.4 m, off Waikiki, Honolulu Dist, ‘Oahu, Hawaiian
Islands (21 16.20 N, 157 50.40 W), Leg.: T. Bratcher, OCT 1975.
keenae, Scissurella McLean, 1970a: Malac. Review 2:117, fig. 4.
HOLOTYPE: LACM 1372 (dry); PARATYPES: LACM 1373 (6 dry), 84 m, Isla Angel de la
Guarda, Golfo de California, México (29 32 N, 113 38 W), Leg.: R/V Velero III, 20 MAR 1938.
Remarks: Currently in the genus Anatoma.
parilis, Temnozaga McLean, 1989b: Nat. Hist. Mus. L.A. Co. Cont. Sci. 407:9-11, figs. 2G-H,
5C-D.
PARATYPE: LACM 2354 (shell missing; animal wet), 2615 m, East Pacific Rise, East Pacific
(20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 14 MAY 1982.
senni, Satondella Geiger, 2003: Moll. Res. 23(1):30-31, fig. 3.
HOLOTYPE: LACM 2911 (coated); PARATYPE: LACM 2923 (1 coated, missing), 10-20 m,
off Punta Rosalia, E of Anakena, Isla de Pascua (= Easter Id.), Leg.: B.K. Raines, DEC 2000.
theca, Sutilizona McLean, 1989b: Nat. His. Mus. L.A. Co. Cont. Sci. 407:15, figs. 3A-B, 5E-F.
PARATYPE: LACM 2355 (2 dry), 2715 m, on sulfide mounds at base of inactive chimney, East
Pacific Rise, East Pacific (11 46 N, 103 47 W), Leg.: R/V Alvin, 25 MAR 1988.
zimmeri, Sinezona Geiger, 2003: Moll. Res. 23(1):33-37, figs. 5-9.
HOLOTYPE: LACM 2912 (coated); PARATYPES: LACM 2913 (3 coated), 10-20 m, off Punta
Rosalia, E of Anakena, Isla de Pascua (= Easter Id.), Leg.: B.K. Raines, DEC 2000.
Family Anatomidae McLean, 1989
baxteri, Anatoma McLean, 1984a: The Veliger 26(3):233, fig. 1.
HOLOTYPE: LACM 1991 (dry); PARATYPES: LACM 1992 (25 dry), 9 m, off N side Hesketh
Id., Kachemak Bay, Kenai Peninsula, Kenai-Cook Inlet Co., Alaska (59 30.5 N, 151 31 W),
Leg.: J.H. McLean & Rae Baxter, 2 AUG 1973.
rainesi, Anatoma Geiger, 2003: Moll. Res. 23(1):44-46, figs. 11-12.
HOLOTYPE: LACM 2909 (coated); PARATYPE: LACM 2910 (1 coated), 10-20 m, off Punta
Rosalia, E of Anakena, Isla de Pascua (= Easter Id.), Leg.: B.K. Raines, DEC 2000.
Superfamily FISSURELLOIDEA Fleming, 1822
Family Fissurellidae Fleming, 1822
angusta, Emarginula McLean, 1970c: The Veliger 12(3):363-364, pl. 54, figs. 14-15.
HOLOTYPE: LACM 1308 (dry), 60-170 m, off Isla San Félix, Islas Desventurados, East Pacific
(26 20 S, 80 02 W), Leg.: R/V Anton Bruun, 12 JUL 1966
arnoldi, Diodora McLean, 1966: L.A. Co. Mus. Nat. Hist. Cont. Sci. 100:5-7, figs. 3-4.
HOLOTYPE: LACM 1143 (dry); PARATYPES: LACM 1144 (9 dry), 21 m, 3 mi. S of Carmel,
San Jose Beach Creek, Monterey Co., California, Leg.: J.H. McLean, 6 FEB 1964
barbouri, Fissurella (Cremides) Farfante, 1943: Johnsonia 1(10):9.
PARATYPE: LACM 1218 (1 dry), 1 mi. W of Nassau, New Providence, Bahamas, Leg.: P.D.
Ford, 1943.
chilensis, Tugali McLean, 1970c: The Veliger 12(3):362-363, pl. 54, figs. 8-9.
HOLOTYPE: LACM 1306 (dry), 0 m, intertidal, Bahía Carvajal, S end of Isla Más a Tierra, Islas
Juan Fernández, East Pacific (34 50 S, 70 00 W), Leg.: R. Seapy, 17 JUL 1966.
decorata, Puncturella Cowan & McLean, 1968: The Veliger 11(2):105-108.
PARATYPES: LACM 1175 (2 dry), 94 m, off W coast of Queen Charlotte Ids., British
Columbia, Canada (53 21.3 N, 133 04.1 W), Leg.: F.R. Bernard, 11 AUG 1967.
Remarks: Currently in the genus Cranopsis.
decemcostata, Fissurella (Cremides) McLean, 1970c: The Veliger 12(3):365-366, pl. 54, figs.
1-2.
HOLOTYPE: LACM 1313 (dry); PARATYPE: LACM 1314 (1 dry), Playa Caleta, Acapulco,
Guerrero, México, Leg.: E.C. Huffman, 18 DEC 1937.
depressa, Clathrosepta McLean & Geiger, 1998: Nat. Hist. Mus. L.A. Co. Cont. Sci. 475:13-14,
figs. 7A-D.
HOLOTYPE: LACM 2784 (shell dry, animla wet), 1160 m, eastern slope of Volcano 5, East
Pacific Rise, East Pacific (12 58.0'N, 103 26.0'W), Leg.: L. Levin, R/V Alvin, 15 JUN 1984.
deroyae, Fissurella (Cremides) McLean, 1970a: Malac. Review 2:118, figs. 7-8.
HOLOTYPE: LACM 1375 (dry); PARATYPES: LACM 1376 (5 dry), Bahía Academy, Isla
Santa Cruz (= Indefatigable Id.), Isla Galápagos (0 45 S, 90 20 W), Leg.: J. DeRoy, OCT 1967.
dictya, Emarginula McLean, 1970c: The Veliger 12(3):364, pl. 54, figs. 16-17.
HOLOTYPE: LACM 1309 (dry); PARATYPE: LACM 1310 (1 dry), 160-170 m, off Isla San
Félix, Islas Desventurados, East Pacific (26 20 S, 80 02 W), Leg.: R/V Anton Bruun, 12 JUL
1966.
eleanorae, Lucapinella McLean, 1967: The Veliger 9(3):350-351, pl. 49, figs. 2-4.
HOLOTYPE: LACM 1153 (dry),18 m, off La Cruz, Bahía Banderas, Jalisco, México, Leg.: J.H.
McLean & A.M. Keen, 24 MAR 1965.
fumarium, Puncturella Hedley, 1911: Report on the Mollusca obtained by the F.I.S.
"Endeavor," ... 1909-1910 under H.C. Dannevig, Commonwealth Director of Fisheries
1:100-101.
PARATYPE: LACM 1964 (1 dry), 183 m, 64 km S of Cape Wiles, South Australia, Australia.
galapagensis, Nesta McLean, 1970c: The Veliger 12(3):363, pl. 54, fig. 18-19.
HOLOTYPE: LACM 1307 (dry), 146-183 m, Caleta Tagus, Isla Isabela (= Albemarle Id.), Islas
Galápagos (0 16 S, 91 22 W), Leg.: R/V Velero III, 15 JAN 1934.
hessleri, Manganesepta McLean & Geiger, 1998: Nat. Hist. Mus. L.A. Co. Cont. Sci. 475:7,
figs. 2A-G.
HOLOTYPE: LACM 2785 (dry); PARATYPES: LACM 2786 (3 dry), 4500 m, North Equitorial
Pacific, northwest of Ile de Clipperton, East Pacific (14 40'N, 125 25'W), R/V Melville, JUN
1983.
howensis, Scutus Iredale, 1940: Australian Zool. 9(4):429-430.
PARATYPES: LACM 1963 (2 dry), Lord Howe Id., Australia, Leg.: R. Bell.
levinae, Cornisepta McLean & Geiger, 1998: Nat. Hist. Mus. L.A. Co. Cont. Sci. 475:22-23,
figs. 13A-F.
HOLOTYPE: LACM 2788 (dry), 1775 m, summit of Volcano 6, East Pacific Rise, East Pacific
(12 44.0'N, 102 33.0'W), Leg.: Lisa Levin, R/V Alvin, 3 JUN 1994.
longifissa, Puncturella Dall, 1914: The Nautilus 28(5):63.
PARATYPE: LACM 1689 (1 dry), 18 m, off Ostrov Beringa (= Bering Id.), Komandorskiye
Ostrova, western Bering Sea, Russia.
microsticta, Diodora Dall, 1927: Proc. U.S. Nat. Mus. 70(2268):9.
SYNTYPES: LACM 1878 (2 dry), Florida.
morrisoni, Fissurella (Clypidella) McLean, 1970a: Malac. Review 2:117-118, figs. 5-6.
HOLOTYPE: LACM 1374 (1 dry), 0 m, intertidal, Bahía Honda, Isla Coiba, Veraguas Prov.,
Panamá (7 44.42 N, 81 32.7 W), Leg.: R/V Velero III, 1 MAR 1938.
platyactis, Amblychilepas McLean & Kilburn, 1986: Nat. Hist. Mus. L.A. Co. Cont. Sci.
379:9-11.
PARATYPE: LACM 2108 (1 wet), Port Alfred, Cape Prov., South Africa, Leg.: E. Warren, JUL
1912.
punctifissa, Diodora McLean, 1970c: The Veliger 12(3):365, pl. 54, figs. 5-7.
HOLOTYPE: LACM 1311 (1 dry); PARATYPES: LACM 1312 (5 dry), 183-274 m, off Isla
Wenman (= Wolf Id.), Islas Galápagos (1 23.1 N, 91 48.7 W), Leg.: R/V Velero III, 11 JAN
1934.
pusilla, Diodora Berry, 1959: Leaflets in Malac. 1(18):109.
PARATYPE: LACM 1721 (1 dry), 11-18 m, off Acapulco, Guerrero, México, Leg.: M.W.
Williams, 3-7 APR 1937.
roseola, Leurolepas McLean, 1970c: The Veliger 12(3):366-367, pl. 54, figs. 12-13.
HOLOTYPE: LACM 1315 (shell dry, animal wet), 24 m, Isla Magdalena, Islas Tres Marias,
Nayarit, México (21 25.6 N, 106 21.1 W), Leg.: R/V Velero III, 9 MAY 1939.
rothi, Puncturella McLean, 1984a: The Veliger 26(3):234.
PARATYPE: LACM 1993 (1 dry), 283-521 m, Delgada Canyon, off Buck Creek, Humboldt Co.,
California (40 05'N, 124 08'W), Leg.: R/V N.B. Scofield, 6 OCT 1950.
verenae, Cornisepta McLean & Geiger, 1998: Nat. Hist. Mus. L.A. Co. Cont. Sci. 475:23-24,
figs. 14A-F.
HOLOTYPE: LACM 2787 (wet), 1530 m, Axial Seamount, Juan de Fuca Ridge off Oregon, East
Pacific (45 56.2'N, 130 04'W), Leg.: V. Tunnicliffe, R/V Pisces, 31 JUL 1996.
Superfamily LEPETODRILOIDEA McLean, 1988
Family Lepetodrilidae McLean, 1988
corrugatus, Lepetodrilus McLean, 1992b: The Veliger 36(1):29-32, figs. 11-16.
HOLOTYPE: LACM 2256 (wet), 2420 m, Heinecken Hollow, Middle Valley, Juan de Fuca
Ridge off Washington (48 25.8 N, 128 40.9 W), Leg.: R/V Alvin, 5 AUG 1990.
cristatus, Lepetodrilus McLean, 1988c: Phil. Trans. Roy. Soc., London, ser. B, 319(1192):15-16,
figs. 57-60.
HOLOTYPE: LACM 2130 (dry), 2615 m, East Pacific Rise, East Pacific (20 50.0 N, 9 06.0 W),
Leg.: R/V Alvin, 17 APR 1982; PARATYPE: LACM 2495 (1 wet), 2483 m, Galapagos Rift, East
Pacific (0 47.1 N, 86 08.5 W), Leg.: R/V Alvin, 13 FEB 1979; PARATYPES: LACM 2496 (1
dry; 2 wet), 2482 m, Galapagos Rift, East Pacific (0 48.0 N, 86 09.0 W), Leg.: R/V Alvin, 6
DEC 1979; PARATYPE: LACM 2497 (1 wet), 2490 m, Galapagos Rift, East Pacific (0 48.0 N,
86 09.0 W), Leg.: R/V Alvin, 8 DEC 1979; PARATYPES: LACM 2498 (8 dry, 5 wet), 2615 m,
East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 17 APR 1982;
PARATYPE: LACM 2499 (1 wet), 2617 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 19 APR 1982; PARATYPE: LACM 2500 (1 wet), 2633 m, East
Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 20 APR 1982;
PARATYPE: LACM 2501 (1 wet), 2618 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 4 MAY 1982; PARATYPES: LACM 2502 (2 wet), 2614 m, East
Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 6 MAY 1982;
PARATYPES: LACM 2503 (5 wet), 2616 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 7 MAY 1982; PARATYPES: LACM 2504 (10 wet), 2618 m,
East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 9 MAY 1982;
PARATYPES: LACM 2505 (14 wet), 2616 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 10 MAY 1982.
elevatus, Lepetodrilus McLean, 1988c: Phil. Trans. Roy. Soc., London, ser. B, 319(1192):11-13,
figs. 36-42.
HOLOTYPE: LACM 2127 (shell dry, animal wet), 2633 m, East Pacific Rise, East Pacific,
(20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 20 APR 1982; PARATYPE: LACM 2529 (1 wet),
2622 m, East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 2 NOV 1979;
PARATYPES: LACM 2530 (5 wet), 2493 m, Galapagos Rift, East Pacific (0 47.6 N,
86 06.4 W), Leg.: R/V Alvin, 21 JAN 1979; PARATYPES: LACM 2531 (5397 wet), 2615 m,
East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 17 APR 1982;
PARATYPES: LACM 2532 (7 wet), 2617 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 19 APR 1982; PARATYPES: LACM 2533 (333 dry, 13611 wet),
2633 m, East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 20 APR 1982;
PARATYPES: LACM 2534 (29 wet), 2616 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 21 APR 1982; PARATYPES: LACM 2535 (20 wet), 2618 m,
East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 24 APR 1982;
PARATYPES: LACM 2536 (2570 wet), 2612 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 25 APR 1982; PARATYPES: LACM 2537 (2 dry), 2617 m, East
Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 26 APR 1982;
PARATYPES: LACM 2538 (4027 wet), 2618 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 4 MAY 1982; PARATYPES: LACM 2539 (446 wet), 2614 m,
East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 6 MAY 1982;
PARATYPES: LACM 2540 (2393 wet); 2616 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 7 MAY 1982; PARATYPES: LACM 2541 (19740 wet), 2618 m,
East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 9 MAY 1982;
PARATYPES: LACM 2542 (751 wet), 2616 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 10 MAY 1982; PARATYPES: LACM 2543 (4 wet), 2615 m,
East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 14 MAY 1982.
emarginatus, Gorgoleptis McLean, 1988c: Phil. Trans. Roy. Soc., London, ser. B, 319(1192):
22-23, figs. 84-88, 91.
HOLOTYPE: LACM 2133 (dry), 2618 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 9 MAY 1982; PARATYPES: LACM 2477 (3 dry), 2615 m, East
Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 17 APR 1982;
PARATYPE: LACM 2478 (1 dry), 2612 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 25 APR 1982; PARATYPES: LACM 2479 (14 dry), 2618 m,
East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 4 MAY 1982;
PARATYPES: LACM 2480 (2 wet), 2614 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 6 MAY 1982; PARATYPES: LACM 2481 (3 wet [1 w/o shell], 1
dry), 2618 m, East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 9 MAY
1982; PARATYPE: LACM 2482 (1 dry), 2616 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 10 MAY 1982.
Remarks: Radula of paratype LACM 2479 mounted on glass slide.
fucensis, Lepetodrilus McLean, 1988c: Phil. Trans. Roy. Soc., London, ser. B, 19(1192):18-19,
figs. 75-77, 79-81.
HOLOTYPE: LACM 2132 (shell dry, animal wet), 2208 m, unnamed vent field, Endeavor
Segment, Juan de Fuca Ridge, off Washington, East Pacific (47 57.0 N, 129 04.0 W), Leg.: R/V
Alvin, 25 JUL 1984; PARATYPES: LACM 2506 (595 wet), 2380 m, southern Juan de Fuca
Ridge off Oregon, East Pacific (45 13.0 N, 130 09.0 W), Leg.: R/V Alvin, 15 JUL 1984;
PARATYPES: LACM 2507 (378 wet, 3 dry), 2208 m, Endeavor Segment, Juan de Fuca Ridge
off Washington, East Pacific (47 57.0 N, 129 04.0 W), Leg.: R/V Alvin, 25 JUL 1984;
PARATYPES: LACM 2508 (22 wet),1599 m, Axial Seamount, mid Juan de Fuca Ridge off
Oregon, East Pacific (45 59.5 N, 130 03.5 W), Leg.: R/V Pisces IV, 10 AUG 1983;
PARATYPES: LACM 2509 (1 dry, 9 wet),1587 m, Axial Seamount, mid Juan de Fuca Ridge off
Oregon, East Pacific (45 59.5 N, 130 03.5 W), Leg.: R/V Pisces IV, 12 AUG 1983;
PARATYPES: LACM 2510 (2 wet), 1592 m, Axial Seamount, mid Juan de Fuca Ridge off
Oregon, East Pacific (45 59.5 N, 130 03.5 W), Leg.: R/V Pisces IV, 17 AUG 1983;
PARATYPES: LACM 2511 (23 wet), 2199 m, Endeavor Segment, Juan de Fuca Ridge off
Washington, East Pacific (47 57.0 N, 129 06.0 W), Leg.: R/V Alvin, 06 SEP 1984;
PARATYPES: LACM 2512 (1 dry, 11 wet), 1818 m, Magic Mountain, Explorer Ridge, off
Vancouver Id., Juan de Fuca Ridge, East Pacific (49 45.6 N, 130 16.1 W), Leg.: R/V Pisces IV,
1 JUL 1984; PARATYPES: LACM 2513 (12 wet), 1808 m, Magic Mountain, Explorer Ridge,
off Vancouver Id., Juan de Fuca Ridge, East Pacific (49 45.6 N, 130 16.1 W), Leg.: R/V Pisces
IV, 2 JUL 1984; PARATYPES: LACM 2514 (13 wet), 1812 m, Upper Magic Mountain,
Explorer Ridge, off Vancouver Id., Juan de Fuca Ridge, East Pacific (49 45.5 N, 130 16.1 W)
Leg.: R/V Pisces IV, 4 JUL 1984
galriftensis, Lepetodrilus elevatus McLean, 1988c: Phil. Trans. Roy. Soc., London, ser. B,
319(1192):13-14, figs. 45-48.
HOLOTYPE: LACM 2128 (shell dry, animal wet), 2451 m, Rose Garden rift field, Galapagos
Rift, East Pacific (0 48.0 N, 86 13.0 W), Leg.: R/V Alvin, 7 DEC 1979; PARATYPES: LACM
2515 (2 wet), 2495 m, Galapagos Rift, East Pacific (0 48.2 N, 86 04.1 W), Leg.: R/V Alvin, 20,
JAN 1979; PARATYPE: LACM 2516 (1 wet), 2493 m, Galapagos Rift, East Pacific (0 47.0 N,
86 08.0 W), Leg.: R/V Alvin, 24 JAN 1979; PARATYPES: LACM 2517 (185 wet, 1 dry), 2482
m, Galapagos Rift, East Pacific (0 48.1 N, 86 07.0 W), Leg.: R/V Alvin, 25 JAN 1979;
PARATYPES: LACM 2518 (6 wet), 2488 m, Galapagos Rift, East Pacific (0 48.5 N,
86 09.1 W), Leg.: R/V Alvin, 12 FEB 1979; PARATYPES: LACM 2519 (7 wet), 2483 m,
Galapagos Rift, East Pacific (0 47.1 N, 86 08.5 W), Leg.: R/V Alvin, 13 FEB 1979;
PARATYPES: LACM 2520 (4 wet), 2447 m, Galapagos Rift, East Pacific (0 48.9 N,
86 13.3 W), Leg.: R/V Alvin, 15 FEB 1979; PARATYPES: LACM 2521 (3 wet), 2457 m,
Galapagos Rift, East Pacific (0 48.2 N, 86 14.9 W), Leg.: R/V Alvin, 19 FEB 1979;
PARATYPES: LACM 2522 (2 wet), 2460 m, Galapagos Rift, East Pacific (0 48.2 N,
86 13.6 W), Leg.: R/V Alvin, 21 FEB 1979; PARATYPES: LACM 2523 (2 wet), 2457 m,
Galapagos Rift, East Pacific (0 48.0 N, 86 13.0 W), Leg.: R/V Alvin, 30 NOV 1979;
PARATYPES: LACM 2524 (7 wet), 2451 m, Galapagos Rift, East Pacific (0 48.0 N,
86 13.0 W), Leg.: R/V Alvin, 1 DEC 1979; PARATYPES: LACM 2525 (13 wet), 2482 m,
Galapagos Rift, East Pacific (0 48.0 N, 86 09.0 W), Leg.: R/V Alvin, 6 DEC 1979;
PARATYPES: LACM 2526 (15 wet), 2451 m, Galapagos Rift, East Pacific (0 48.0 N,
86 13.0 W), Leg.: R/V Alvin, 7 DEC 1979; PARATYPES: LACM 2527 (8 wet), 2490 m,
Galapagos Rift, East Pacific (0 48.0 N, 86 09.0 W), Leg.: R/V Alvin, 8 DEC 1979;
PARATYPES: LACM 2528 (112 wet), 2518 m, Galapagos Rift, East Pacific (0`47.0'N,
86`08.0'W), Leg.: R/V Alvin, 10 DEC 1979.
gordensis, Lepetodrilus Johnson, Young, Jones, Warén, & Vrijenhoek, 2006: Biol. Bull. 210:
151-157, figs. 3-5.
PARATYPES: LACM 1974 (5 wet), 2716 m, Gorda Ridge, East Pacific (42º45'N, 126º42'W),
Leg.: R/V Tiburon.
guaymasensis, Lepetodrilus McLean, 1988c: Phil. Trans. Roy. Soc., London, ser. B, 319(1192):
16-18, figs. 66-68,70-72.
HOLOTYPE: LACM 2131 (shell dry, animal wet), PARATYPES: LACM 2493 (3 dry, 2 wet)
2019 m, southern trough of Guaymas Basin, Golfo de California, Baja California Sur, Mexico
(27 01.0 N, 111 25.0 W), Leg.: R/V Alvin, 12 JAN 1982; PARATYPE: LACM 2494 (1 wet),
2014 m, southern trough of Guaymas Basin, Golfo de California, Baja California Sur, Mexico
(27 02.0 N, 111 24.0 W), Leg.: R/V Alvin, 20 JAN 1982.
ovalis, Lepetodrilus McLean, 1988c: Phil. Trans. Roy. Soc., London, ser. B, 319(1192):14-15,
figs. 51-55.
HOLOTYPE: LACM 2129 (shell dry, animal wet), 2633 m, East Pacific Rise, East Pacific
(20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 20 APR 1982; PARATYPES: LACM 2483 (2 wet),
2615 m, East Pacific Rise, East Pacific (20 50.0 N, 109 `06.0 'W), Leg.: R/V Alvin, 17 APR
1982; PARATYPES: LACM 2484 (96 wet, 46 dry), 2633 m, East Pacific Rise, East Pacific
(20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 20 APR 1982; PARATYPES: LACM 2485 (7 wet),
2626 m, East Pacific Rise, East Pacific (20 50.0 N, 109 06.6 W), Leg.: R/V Alvin, 21 APR 1982;
PARATYPES: LACM 2486 (5 wet), 2618 m, East Pacific Rise, East Pacific (20 59.0 N,
109 06.0 W), Leg.: R/V Alvin, 24 APR 1982; PARATYPES: LACM 2487 (5 dry; 52 wet),
2612 m, East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 25 APR 1982;
PARATYPES: LACM 2488 (73 wet), 2618 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 4 MAY 1982; PARATYPES: LACM 2489 (3 wet), 2614 m, East
Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 6 MAY 1982;
PARATYPES: LACM 2490 (31 wet), 2616 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 7 MAY 1982; PARATYPES: LACM 2491 (29 wet), 2618 m,
East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 9 MAY 1982;
PARATYPES: LACM 2492 (16 wet), 2616 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 10 MAY 1982.
patulus, Gorgoleptis McLean, 1988c: Phil. Trans. Roy. Soc., London, ser. B, 319(1192):24-25,
figs. 98-102.
HOLOTYPE: LACM 2134 (dry), 2482 m, Mussel Bed vent field, Galapagos Rift, East Pacific
(0 48.0 N, 86 09.0 W), Leg.: R/V Alvin, 6 DEC 1979; PARATYPE: LACM 2476 (1 wet),
2490 m, Mussel Bed vent field, Galapagos Rift, East Pacific (0 48.0 N, 86 09.0 W), Leg.: R/V
Alvin, 8 DEC 1979.
pustulosus, Lepetodrilus McLean, 1988c: Phil. Trans. Roy. Soc., London, ser. B, 319(1192):
8-11, figs. 25-30.
HOLOTYPE: LACM 2126 (dry), 2493 m, Mussel Bed vent field, Galapagos Rift, East Pacific
(0 47.6 N, 86 06.4 W), Leg.: R/V Alvin, 21 JAN 1979; PARATYPE: LACM 2544 (1 wet), 2496
m, Galapagos Rift, East Pacific (0 47.3 N, 86 07.8 W), Leg.: R/V Alvin, 16 MAR 1977;
PARATYPES: LACM 2545 (209 wet), 2493 m, Galapagos Rift, East Pacific (0 47.6 N,
86 06.4 W), Leg.: R/V Alvin, 21 JAN 1979; PARATYPES: LACM 2546 (71 wet), 2482 m,
Galapagos Rift, East Pacific (0 48.1 N, 86 07.0 W), Leg.: R/V Alvin, 25 JAN 1979;
PARATYPES: LACM 2547 (28 wet), 2488 m, Galapagos Rift, East Pacific (0 48.5 N,
86 09.1 W), Leg.: R/V Alvin, 12 FEB 1979; PARATYPES: LACM 2548 (64 wet), 2483 m,
Galapagos Rift, East Pacific (0 47.1 N, 86 08.5 W), Leg.: R/V Alvin, 13 FEB 1979;
PARATYPES: LACM 2549 (67 wet), 2447 m, Galapagos Rift, East Pacific (0 48.9 N,
86 13.3 W), Leg.: R/V Alvin, 15 FEB 1979; PARATYPES: LACM 2550 (4 wet), 2488 m,
Galapagos Rift, East Pacific (0 48.3 N, 86 13.4 W), Leg.: R/V Alvin, 16 FEB 1979;
PARATYPE: LACM 2551 (1 wet), 2454 m, Galapagos Rift, East Pacific (0 48.3 N, 86 13.8 W),
Leg.: R/V Alvin, 17 FEB 1979; PARATYPES: LACM 2552 (2 wet), 2457 m, Galapagos Rift,
East Pacific (0 48.2 N, 86 14.9 W), Leg.: R/V Alvin, 19 FEB 1979; PARATYPES: LACM 2553
(7 wet), 2460 m, Galapagos Rift, East Pacific (0 48.2 N, 86 13.6 W), Leg.: R/V Alvin, 21 FEB
1979; PARATYPES: LACM 2554 (9 wet), 2457 m, Galapagos Rift, East Pacific (0 48.0 N,
86 13.0 W), Leg.: R/V Alvin, 30 NOV 1979; PARATYPES: LACM 2555 (97 wet), 2451 m,
Galapagos Rift, East Pacific (0 48.0 N, 86 13.0 W), Leg.: R/V Alvin, 1 DEC 1979;
PARATYPES: LACM 2556 (19 wet), 2482 m, Galapagos Rift, East Pacific (0 48.0 N,
86 09.0 W), Leg.: R/V Alvin, 6 DEC 1979; PARATYPES: LACM 2557 (46 wet), 2451 m,
Galapagos Rift, East Pacific (0 48.0 N, 86 13.0 W), Leg.: R/V Alvin, 7 DEC 1979;
PARATYPES: LACM 2558 (68 wet), 2490 m, Galapagos Rift, East Pacific (0 48.0 N,
86 09.0 W), Leg.: R/V Alvin, 8 DEC 1979; PARATYPES: LACM 2559 (12 wet), 2518 m,
Galapagos Rift, East Pacific (0 47.0 N, 86 08.0 W), Leg.: R/V Alvin, 10 DEC 1979;
PARATYPES: LACM 2560 (199 wet), 2615 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 17 APR 1982; PARATYPES: LACM 2561 (1183 wet), 2633 m,
East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 20 APR 1982;
PARATYPES: LACM 2562 (3 wet); 2616 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 21 APR 1982; PARATYPES: LACM 2563 (177 wet), 2612 m,
East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 25 APR 1982;
PARATYPES: LACM 2564 (108 wet), 2618 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 4 MAY 1982; PARATYPES: LACM 2565 (27 wet), 2614 m,
East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 6 MAY 1982;
PARATYPES: LACM 2566 (10 wet), 2616 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 7 MAY 1982; PARATYPES: LACM 2567 (145 wet), 2618 m,
East Pacific Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 9 MAY 1982;
PARATYPES: LACM 2568 (50 wet), 2616 m, East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 10 MAY 1982.
tevnianus, Lepetodrilus McLean, 1993 : The Veliger 36(1):28-29, figs. 1-10.
HOLOTYPE: LACM 2254 (wet); PARATYPES: LACM 2255 (2 wet), 2536 m, East Pacific
Rise, East Pacific (10 56.3 N, 103 41.4 W), Leg.: R/V Alvin, 8 MAR 1988.
Family Clypeosectidae McLean, 1989
curvus, Clypeosectus McLean, 1989b: Nat. Hist. Mus. L.A. Co. Cont. Sci. 407:21-22, figs.
10G-H, 11F.
PARATYPE: LACM 2363 (1 dry), 2213 m, Axial Seamount, Endeavor Segment, Juan de Fuca
Ridge off Washington, East Pacific (47 57.0 N, 129 06.0 W), Leg.: R/V Alvin, 3 SEP 1984;
PARATYPES: LACM 2364 (4 dry), 1575 m, hydrothermal-vent community of Axial Seamount,
Juan de Fuca Ridge off Oregon, East Pacific (45 59.5 N, 130 03.5 W), Leg.: R/V Pisces IV, 31
JUL 1986; PARATYPE: LACM 2365 (1 wet), 1575 m, hydrothermal-vent community of Axial
Seamount, Juan de Fuca Ridge off Oregon, East Pacific (45 59.5 N, 130 03.5 W), Leg.: R/V
Pisces, 3 AUG 1986.
delectus, Clypeosectus McLean, 1989b: Nat. Hist. Mus. L.A. Co. Cont. Sci., 407:18-21, figs.
7F-H, 8E-H, 11B.
PARATYPES: LACM 2356 (2 wet), 2493 m, hydrothermal vents of Galapagos Rift, East Pacific
(0 47.6 N, 86 06.4 W), Leg.: R/V Alvin, 21 JAN 1979; PARATYPES: LACM 2357 (2 wet),
2482 m, hydrothermal vents of Galapagos Rift, East Pacific (0 47.9 N, 86 09.3 W), Leg.: R/V
Alvin, 20 FEB 1979; PARATYPES: LACM 2358 (1 wet), 2451 m, hydrothermal vents of
Galapagos Rift, East Pacific (0 48.0 N, 86 13.0 W), Leg.: R/V Alvin, 1 DEC 1979;
PARATYPE: LACM 2359 (1 wet), 2482 m, hydrothermal vents of Galapagos Rift, East Pacific
(0 48.0 N, 86 09.0 W), Leg.: R/V Alvin, 6 DEC 1979; PARATYPE: LACM 2360 (1 wet), 2490
m, hydrothermal vents of Galapagos Rift, East Pacific (0 48.0 N, 86 09.0 W), Leg.: R/V Alvin, 8
DEC 1979; PARATYPES: LACM 2361 (3 wet), 2618 m, hydrothermal vents of East Pacific
Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 4 MAY 1982; PARATYPES:
LACM 2362 (6 wet), 2616 m, hydrothermal vents of East Pacific Rise, East Pacific (20 50.0 N,
109 06.0 W), Leg.: R/V Alvin, 10 MAY 1982.
marianae, Pseudorimula McLean, 1989b: Nat. Hist. Mus. L.A. Co. Cont. Sci. 407:24-25, figs.
12F-H.
PARATYPES: LACM 2366 (2 dry), 3660 m, Snail Pit vents Mariana Back-Arc Basin, West
Pacific (18 10.95 N, 144 43.2 E), Leg.: R/V Alvin, 27 APR 1987; PARATYPE: LACM 2367 (1
wet), 3640 m, Snail Pit vents, Mariana Back-Arc Basin, West Pacific (18 12.6 N, 144 42.4 E),
Leg.: R/V Alvin, 4 MAY 1987.
midatlantica, Pseudorimula McLean, 1992a: The Nautilus 106(3):115-118.
PARATYPES: LACM 2424 (2 wet), 3478 m, Snake Pit hydrothermal field, Mid-Atlantic Ridge
(23 22 N, 47 57 W), Leg.: R/V Nautile, 28 JUN 1988.
Superfamily NEOMPHALOIDEA McLean, 1981
Family Neomphalidae McLean, 1981
exquisitus, Lacunoides Warén & Bouchet, 1989: Zool. Scripta 18(1):72-74.
PARATYPES: LACM 2282 (10 dry), 2482 m, Galapagos Rift Zone, East Pacific (0 48.1 N,
86 07 W), Leg.: R/V Alvin, 25 JAN 1979.
fretterae, Neomphalus McLean, 1981: Malacologia 21(1-2):294-309, figs. 1, 4, 6-9.
PARATYPES: LACM 1966 (7 dry, 10 wet), 2478-2490 m, Oyster Bed, Galapagos Rift, East
Pacific (0 47.2 N, 86 07.7 W), Leg.: R/V Alvin, 27 FEB 1977; PARATYPES: LACM 2574 (8
dry, 7 wet), 2493 m, Oyster Bed, Galapagos Rift, East Pacific (0 47.2 N, 86 07.7 W), Leg.: R/V
Alvin, 09 MAR 1977; PARATYPES: LACM 2575 (10 dry, 7 wet), 2496 m, Garden of Eden,
Galapagos Rift, East Pacific (0 47.3 N, 86 07.8 W), Leg.: R/V Alvin, 16 MAR 1977
Remarks: Radula of paratype LACM 2575 mounted on glass slide.
laevis, Pachydermia Warén & Bouchet, 1989: Zool. Scripta 18(1):76-80.
PARATYPES: LACM 2283 (3 dry), 2635 m, East Pacific Rise, East Pacific (12 48.6 N,
103 56.7 W), Leg.: BIOCYARISE 1984, 23 MAR 1984.
naticoides, Cyathermia Warén & Bouchet, 1989: Zool. Scripta 18(1):70-72.
PARATYPES: LACM 2281 (54 wet), 2635 m, East Pacific Rise, East Pacific (12 48.6 N,
103 56.7 W), Leg.: BIOCYARISE, 1984.
regularis, Symmetromphalus McLean, 1990a: The Nautilus 104(3):79-81, figs. 1-6.
HOLOTYPE: LACM 2432 (shell dry, animal wet); PARATYPES: LACM 2433 (3 dry, 11 wet, 1
gold coated), 3640 m, Alice Springs vents, Mariana Back-Arc Basin, Northern Mariana Ids.
(18 12.6 N, 144 42.4 E), Leg.: R/V Alvin, 4 MAY 1987.
Family Peltospiridae McLean, 1989
concentrica, Rhynchopelta McLean, 1989a: Zool. Scripta 18(1):58, figs. 32-34.
HOLOTYPE: LACM 2318 (dry); PARATYPES: LACM 2319 (1 dry, 148 wet), 2633 m, East
Pacific Rise, East Pacific (20 50.0 N, 105 06.0 W), Leg.: R/V Alvin, 20 APR 1982;
PARATYPES: LACM 2320 (3 dry, 25 wet), 2635 m, East Pacific Rise, East Pacific (12 48.6 N,
103 56.7 W), Leg.: R/V Cyana, 24 MAR 1984.
delicata, Peltospira McLean, 1989a: Zool. Scripta 18(1):53.
PARATYPES: LACM 2312 (2 dry, 5 wet), 2635 m, East Pacific Rise, East Pacific (12 48.6 N,
103 56.7 W), Leg.: R/V Cyana, 15 MAR 1984
fistulosa, Echinopelta McLean, 1989a: Zool. Scripta 18(1):60, figs. 41-43.
HOLOTYPE: LACM 2321 (shell dry, animal wet); PARATYPES: LACM 2322 (2 dry, 4 wet, 1
gold coated), 2615 m, East Pacific Rise, East Pacific (20 50.0 N, 105 06.0 W), Leg.: R/V Alvin,
6 NOV 1979; PARATAYPES: LACM 2323 (2 dry, 6 wet), 2616 m, East Pacific Rise, East
Pacific (20 50.0 N, 105 06.0 W), Leg.: R/V Alvin, 7 MAY 1982; PARATYPES: LACM 2324 (1
wet, 1 gold coated), 2616 m, East Pacific Rise, East Pacific (20 50.0 N, 105 06.0 W), Leg.: R/V
Alvin, 10 MAY 1982.
globulus, Depressigyra Warén & Bouchet, 1989: Zool. Scripta 18(1):80-81.
PARATYPES: LACM 2284 (10 dry), 2208 m, Endeavour Segment, Juan de Fuca Ridge off
Washington, East Pacific (45 57 N, 129 04 W), Leg.: R/V Alvin, 25 JUL 1984.
heminoda, Nodopelta McLean, 1989a: Zool. Scripta, 18(1):53-56 figs. 17-19.
HOLOTYPE: LACM 2313 (shell dry, animal wet); PARATYPES: LACM 2314 (6 dry, 5 wet),
2622 m, East Pacific Rise, East Pacific (20 50.0 N, 105 06.0 W), Leg.: R/V Alvin, 2 NOV 1979;
PARATYPES: LACM 2315 (1 dry, 19 wet), 2616 m, East Pacific Rise, East Pacific (20 50.0 N,
105 06.0 W), Leg.: R/V Alvin, 7 MAY 1982; PARATYPES: LACM 2316 (2 wet), 2635 m, East
Pacific Rise, East Pacific (12 48.6 N, 103 56.7 W), Leg.: R/V Cyana, 15 MAR 1984.
Remarks: Radula of paratype LACM 2315 mounted on glass slide.
humata, Lirapex Warén & Bouchet, 1989: Zool. Scripta 18(1):84-85.
PARATYPES: LACM 2287 (2 dry), 2616 m, East Pacific Rise, East Pacific (20 50 N,
109 06 W), Leg.: R/V Alvin, 11 MAY 1982.
operculata, Peltospira McLean, 1989a: Zool. Scripta 18(1):51-53, figs. 1-3.
HOLOTYPE: LACM 2307 (dry); PARATYPES: LACM 2308 (2 dry), 2617 m, East Pacific
Rise, East Pacific (20 50.0 N, 109 06.0 W), Leg.: R/V Alvin, 19 APR 1982; PARATYPES:
LACM 2309 (10 wet), 2616 m, East Pacific Rise, East Pacific (20 50.0 N, 105 06.0 W), Leg.:
R/V Alvin, 7 MAY 1982; PARATYPES: LACM 2310 (37 wet), 2616 m, East Pacific Rise, East
Pacific (20 50.0 N, 105 06.0 W), Leg.: R/V Alvin, 10 MAY 1982; PARATYPES: LACM 2311
(4 wet), 2630-2635 m, East Pacific Rise, East Pacific (12 48.6 N, 103 56.7 W), Leg.: R/V
Cyana, 12 MAR 1982.
planispira, Depressigyra Warén & Bouchet, 1989: Zool. Scripta 18(1):81-82.
PARATYPES: LACM 2285 (4 wet), 2633 m, East Pacific Rise, East Pacific (20 50 N,
109 06 W), Leg.: R/V Alvin, 20 APR 1982.
reticulata, Solutigyra Warén & Bouchet, 1989: Zool. Scripta 18(1):82-84.
PARATYPES: LACM 2286 (3 dry), 2617 m, East Pacific Rise, East Pacific (20 50 N,
109 06 W), Leg.: R/V Alvin, 26 APR 1982.
subnoda, Nodopelta McLean, 1989a: Zool. Scripta 18(1):56.
PARATYPES: LACM 2317 (5 dry, 15 wet), 2635 m, East Pacific Rise, East Pacific (12 48.6 N,
103 56.7 W), Leg.: R/V Cyana, 15 MAR 1984.
Superfamily LEPETELLOIDEA Dall, 1882
Family Pseudococculinidae Hickman, 1983
askewi, Kaiparapelta McLean & Harasewych, 1995: Nat. Hist. Mus. L.A. Co. Cont. Sci.
453:28-29.
PARATYPE: LACM 2740 (1 dry), 194 m, 165 km E of Charleston, Charleston County, South
Carolina (32 43.68 N, 78 05.72 W), Leg.: T.M. Askew & M.G. Harasewych, R/V Clelia, 6 JUL
1993.
careyi, Yaquinabyssia McLean, 1988a: Zool. Scripta 17(2):159-160, figs. 12-14.
HOLOTYPE: LACM 2069 (shell dry, animal wet); PARATYPES: LACM 2070 (2 dry, 1 wet),
2774 m, 171 km W of Cape Foulweather, Cascadia Abyssal Plain off Oregon, East Pacific
(44 53.7 N, 126 33.4 W), Leg.: Oregon State Univ., R/V Yaquina, 19 JAN 1970.
georgiana, Tentaoculus Dall, 1927: Proc. U.S. Nat. Mus. 70:121.
PARALECTOTYPES: LACM 2739 (2 dry), designated by McLean & Harasewych (1995:28),
Nat. Hist. Mus. L.A. Co. Cont. Sci. 453. 805 m, off Georgia, Leg.: U.S. Fish Commission.
gordensis, Amphiplica (Gordabyssia) McLean, 1991: The Veliger 34(1):44-46, figs. 25-26.
HOLOTYPE: LACM 2440 (shell dry, animal wet); PARATYPES: LACM 2441 (1 dry, 36 wet, 1
gold coated), 3305 m, Escanaba Trough, Gorda Ridge off California, East Pacific (41 00.4 N,
127 29.3 W), Leg.: R/V Alvin, 5 JUN 1988.
gracilis, Notocrater Marshall, 1985b: New Zealand Jour. of Zool. 12(4):528-530.
PARATYPES: LACM 2773 (3 dry), 55-73 m, off Hicks Bay, East Coast, North Id., New
Zealand, Leg.: A.S. Voss & K. Hipkins.
gregaria, Pseudococculina Marshall, 1985b: New Zealand Jour. Zool. 12(4):522-524.
PARATYPES: LACM 2768 (3 wet), 800-810 m, Chatham Rise, SE of Cape Pelliser,
Wellington, North Id., New Zealand (42 43.9 S, 176 08 E), Leg.: R/V Kaltan, 28 SEP 1982.
haptricola, Tentaoculus Marshall, 1985b: New Zealand Jour. Zool. 12(4):534-535.
PARATYPES: LACM 2770 (3 wet), 1070-1198 m, off Castlepoint, Wellington, North Id., New
Zealand (41 11.0 S, 176 40.0 E), Leg.: R/V James Cook, 4 APR 1984.
houbricki, Notocrater McLean & Harasewych, 1995: Nat. Hist. Mus. L.A. Co. Cont. Sci.
453:24-26.
PARATYPE: LACM 2737 (1 dry), 412 m, S of Settlement Pt., Grand Bahama Id., Bahamas
(26 37.52 N, 78 58.93 W), Leg.: C. Messing, R/V Johnson-Sea-Link II, 18 MAY 1992.
plutonica, Amphiplica Leal & Harasewych, 1999: Invert. Biol. 118(2):131-133.
PARATYPES: LACM 2808 (5 wet), 6646-6600 m, Cayman Trench, SE of Grand Cayman,
Cayman Ids. (19°38'N, 76°18.1'W), Leg.: R/V Gillis, 21 JUL 1975.
ponderi, Notocrater Marshall, 1985b: New Zealand Jour. Zool. 12(4):530 .
PARATYPES: LACM 2769 (3 wet), 384 m, E of Brush Id., New South Wales, Australia
(35 33 S, 150 44 E), Leg.: R/V Kapala, 30 NOV 1976.
similaris, Copulabyssia Hasegawa, 1997: Nat. Sci. Mus. Tokyo, Memoirs 12:86-89.
PARATYPES: LACM 2839 (3 dry shells, animals wet), 430-710 m, off Toi, Suruga-wan,
Shizuoka Pref., Honshu, Japan (34 56.7 N, 138 42.3 E), Leg.: R/V Soro-Maru, 31 NOV 1993.
tibbettsi, Punctabyssia McLean, 1991: The Veliger 34(1):39, figs. 1-5.
HOLOTYPE: LACM 2434 (shell dry, animal wet); PARATYPES: LACM 2435 (1 dry, 4 wet, 2
gold coated), 2700 m, along axis of East Pacific Rise, East Pacific (11 51 N, 103 50 W), Leg.:
R/V Alvin, 22 MAR 1988.
venezuelensis, Amphiplica (Amphiplica) McLean, 1988a: Zool. Scripta 17(2):155-157, figs. 2-5.
PARATAYPES: LACM 2265 (12 dry; 3 wet), 5057 m, Venezuela Basin, due N of Caracas,
Federal District, Venezuela (14 20 N, 67 00 W), Leg.: R/V Bartlett, 30 NOV 1981;
PARATYPES: LACM 2266 (1 dry, 5 wet), 3476-3518 m, Venezuela Basin, due N of Caracas,
Federal District, Venezuela (13 30 N, 64 45 W), Leg.: R/V Bartlett, 27 NOV 1981.
youngi, Notocrater McLean & Harasewych, 1995: Nat. Hist. Mus. L.A. Co. Cont. Sci. 453:
26-27.
PARATYPES: LACM 2738 (3 dry), 518 m, Off Southwest Reef, New Providence Id., Bahamas
(24 54.04 N, 77 33.23 W), Leg.: M. Rice, R/V Johnson-Sea-Link I, 10 MAY 1993.
Family Lepetellidae Dall, 1882
hickmanae, Choristella McLean, 1982: The Veliger 35(4):285-286, figs. 39-40.
HOLOTYPE: LACM 2249 (dry); PARATYPE: LACM 2250 (1 dry), 2176 m, 172 km W of
Strait of Juan de Fuca, Cascadia Abyssal Plain off Washington, East Pacific (43 38.1 N,
126 58 W), Leg.: Oregon State University, 11 SEP 1971.
Remarks: Originally two paratypes, one sectioned, shell destroyed, remaining paratype badly
damaged. Sectioned animal in 70% ETOH.
marshalli, Choristella McLean, 1992b: The Veliger 35(4):278-281.
PARATYPE: LACM 2247 (1 dry), 1097-1116 m, SE of Banks Peninsula, Canterbury, South Id.,
New Zealand (44 55.4 S, 174 04.9 E), Leg.: R/V James Cook, 15 JUN 1984.
nofronii, Choristella McLean, 1992b: The Veliger 35(4):281-282.
PARATYPES: LACM 2248 (2 dry), 50-100 m, Alboran Sea, westernmost Mediterranean, SE of
Cabo de Gata, Almeria Prov., Spain (ca. 36 00 N, 1 30 W).
ponderi, Choristella McLean, 1992b: The Veliger 35(4):282-285.
PARATYPE: LACM 2630 (1 dry), 494-585 m, off Shoalhaven Heads, New South Wales,
Australia (35 56 S, 151 09.5 E), Leg.: R/V Kapala, 10 SEP 1986
Family Addisoniidae Dall, 1882
brophyi, Addisonia McLean, 1985b: The Veliger 28(1):105-106, fig. 6.
HOLOTYPE: LACM 2082 (dry); PARATYPES: LACM 2083 (7 wet), 155-165 m, off Gaviota,
Santa Barbara Co., California (34 12 N, 120 12 W), Leg.: P. Brophy, 28 AUG 1968;
PARATYPES: LACM 2084 (3 dry), 155-165 m, off Gaviota, Santa Barbara Co., California
(34 12 N, 120 12 W), Leg.: P. Brophy, 17 JUL 1968.
Family Bathyphytophilidae Moskalev, 1978
diegensis, Bathyphytophilus Haszprunar & McLean, 1996: Zool. Scripta 25(1):38, figs. 1-7.
HOLOTYPE: LACM 2398 (dry); PARATYPES: LACM 2399 (3 wet), 1224 m, San Diego
Trough at base of Coronado Canyon, off Tijuana, Baja California, México (32 18.2 N,
117 29.8 W), Leg.: P.A. Jumars, 6 DEC 1971.
Family Caymanabyssidae Moskalev, 1978
fosteri, Caymanabyssia (Dictyabysia) McLean, 1991: The Veliger 34(1):41, figs. 9-11.
HOLOTYPE: LACM 2436 (shell dry, animal wet); PARATYPES: LACM 2437 (1 dry, 8 wet, 1
gold coated), 2700 m, along axis of East Pacific Rise, East Pacific (11 51 N, 103 50 W), Leg.:
R/V Alvin, 22 MAR 1988.
hestica, Colotrachelus Marshall, 1985b. New Zealand Jour. Zool. 12(4 ):540 .
PARATYPE: LACM 2772 (1 dry), 1174-1208 m, off Castlepoint, Wellington, North Island, New
Zealand (41 9.9 S, 176 26.5 E), Leg.: R/V James Cook, 4 APR 1984.
sinespina, Caymanabyssia Marshall, 1985b: New Zealand Jour. Zool. 12(4):539-540.
PARATYPE: LACM 2771 (1 dry), 833-891 m, off Jackson Bay, Westland, South Id., New
Zealand (43 26.8 S, 168 54.2 E), Leg.: R/V James Cook, 15 DEC 1983.
vandoverae, Caymanabyssia (Caymanabyssia) McLean, 1991: The Veliger 34(1):41-44, figs.
17-18.
HOLOTYPE: LACM 2438 (shell dry, animal wet); PARATYPES: LACM 2439 (1 dry, 2 wet, 1
gold coated), Escanaba Trough, Gorda Ridge off California, East Pacific (41 00.4 N,
127 29.3 W), Leg.: R/V Alvin, 4 JUN 1988.
Family Pyropeltidae McLean & Haszprunar, 1983
corymba, Pyropelta McLean & Haszprunar, 1987: The Veliger 30(2):200-202, figs. 10-11.
HOLOTYPE: LACM 2277 (shell dry, animal wet), 2022 m, southern trough of Guaymas Basin,
Golfo de California, Baja California Sur, México (27 01.0 N, 111 25.0 W), Leg.: R/V Alvin, 19
JAN 1982.
musaica, Pyropelta McLean & Haszprunar, 1987: The Veliger 30(2):198-200, figs. 5-8.
HOLOTYPE: LACM 2275 (shell dry, animal wet); PARATYPES: LACM 2276 (3 dry, 37 wet),
1575 m, Not-so-miserable Vent, Axial Seamount, Juan de Fuca Ridge off Oregon, East Pacific
(45 59.5 N, 130 03.5 W), Leg.: R/V Pisces IV, 3 AUG 1986; PARATYPES: LACM 2569 (2
dry, 4 wet), 1575 m, Hammond's Hell Vent, Axial Seamount, Juan de Fuca Ridge off Oregon,
East Pacific (45 59.5 N, 130 03.5 W), Leg.: R/V Pisces IV, 19 JUL 1986; PARATYPES: LACM
2570 (9 wet), 1575 m, Southern Axial Vent, Axial Seamount, Juan de Fuca Ridge off Oregon,
East Pacific (45 59.5 N, 130 06.5 W), Leg.: R/V Pisces IV, 29 JUL 1986; PARATYPES: LACM
2571 (2 dry; 7 wet), 1575 m, Anemone Ridge, Axial Seamount, Juan de Fuca Ridge off Oregon,
East Pacific (45 59.5 'N, 130 03.5 W), Leg.: R/V Pisces IV, 30 JUL 1986; PARATYPES:
LACM 2572 (8 wet), 1575 m, Eastern Axial, Axial Seamount, Juan de Fuca Ridge off Oregon,
East Pacific (45 59.5 N, 130 03.5 W), Leg.: R/V Pisces IV, 31 JUL 1986; PARATYPE: LACM
2573 (1 dry), 1575 m, Post Taylor's Vent, Axial Seamount, Juan de Fuca Ridge off Oregon, East
Pacific (45 59.5 N, 130 03.5 W), Leg.: R/V Pisces, 1 AUG 1986.
wakefieldi, Pyropelta McLean, 1992c: Jour. Moll. Stud. 58:410-412, figs. 25-27.
HOLOTYPE: LACM 2462 (dry); PARATYPES: LACM 2463 (7 dry), 940 m, off Pt. Sur,
Monterey Co., California (36 17.6 N, 122 12.2 W), Leg.: R/V David Starr Jordan, JAN-FEB
1987.
Family Osteopeltidae Marshall, 1987
mirabilis, Osteopelta Marshall, 1987: Jour. Moll. Stud. 53(2):122-126.
PARATYPES: LACM 2205 (2 wet), 880-882 m, NE of Chatham Id., New Zealand (42 56.2 S,
175 32.6 W), Leg.: D.A. Robertson; R/V Otago Buccaneer, 17 JUL 1985.
Superfamily TROCHOIDEA Rafinesque, 1815
Family Trochidae Rafinesque, 1815
akuana, Ethminolia Raines, 2007: Visaya 2(1):71-72.
HOLOTYPE: LACM 3011 (dry); 50 m, off Tahai, Isla de Pascua (= Easter Id.) (27º8'24"S,
109º27'30"W), Leg.: B.K. Raines, NOV 2002; PARATYPES: LACM 3012 (6 dry); 50 m, off
Orongo, Isla de Pascua (= Easter Id.) (27º11'58"S, 109º29'28"W), Leg.: B.K. Raines, NOV 2002.
aniesae, Priotrochus Moolenbeek & Dekker, 1992: Bull. Zool. Mus. Amsterdam 13(15): 71-72.
PARATYPE: LACM 2728 (1 wet), Umm Rasas (Rusays), E coast of Masirah Id., Oman, Leg.:
R.G. Moolenbeek & H. Dekker, 21 NOV 1991
bairdii, Turcicula Dall, 1889: Bull. Mus. Comp. Zool. 18:377-378.
PARATYPES: LACM 1207 (5 dry), 757 m, off San Clemente Id., California Channel Ids.,
California, Leg.: R/V Albatross.
Remarks: Currently in the genus Bathybembix.
bergeroni, Tegula (Agathistoma) McLean, 1970a: Malac. Review 2:119, figs. 12-13.
HOLOTYPE: LACM 1379 (dry); PARATYPE: LACM 1380 (1 dry), Bahía Cabita, near Cabo
Corrientes, Choco Dept., Colombia (5 29.3 N, 77 29.58 W), Leg.: R/V Velero III, 13 FEB 1934.
bermudensis, Euchelus Moolenbeek & Faber, 1989: Found. Sci. Res. in Surinam & Neth.
Antilles 123:224.
PARATYPE: LACM 2469 (1 dry), Palm Cave, Bermuda, Leg.: T.M. Iliffe, 20 JAN 1982.
bernardi, Calliostoma McLean, 1984a: The Veliger 26(3):236-237, fig. 5.
HOLOTYPE: LACM 1996 (shell dry, animal wet), 241-271 m, off SE end of Santa Catalina Id.,
California Channel Ids., California (33 15.6 N, 118 13.42 W), Leg.: R/V Velero III, 5 JUL 1940.
Remarks: Currently in the genus Otukaia.
bicostata, Lirularia McLean, 1964: The Veliger 7(2):129-130.
PARATYPES: LACM 1134 (4 dry), 14 m, off the N side of Coronado del Medio, Islas los
Coronados, Pacific Coast, Baja California, México, Leg.: J.H. McLean, 10 AUG 1963.
cookiana, Thoristella chathamensis Powell, 1934: Trans. Proc. Roy. Soc. New Zealand
64(2):154-155.
PARATYPE: LACM 1360 (1 dry), Island Bay, Wellington, North Id., New Zealand.
corteziana, Tegula (Agathistoma) McLean, 1970a: Malac. Review 2:119-120, figs. 14-15.
HOLOTYPE: LACM 1381 (dry); PARATYPES: LACM 1382 (67 dry), 0 m, intertidal, S side of
Cabo Tepoca, Sonora, México (30 16 N, 112 30 W), Leg.: J.H. McLean, 26 MAR 1967.
delli, Calliostoma McLean & Andrade, 1982: Nat. Hist. Mus. L.A. Co. Cont. Sci. 342:7-8,
figs. 14-15.
PARATYPES: LACM 1981 (26 dry, 3 wet), 400 m, Los Vilos, Coquimbo Prov., Chile (31 56 S,
71 54 W), Leg.: H. Andrade, R/V Goden Wind, 29 MAY 1977.
Remarks: Currently in the subgenus Otukaia.
dentatum, Calliostoma Quinn, 1992: The Nautilus 106(3):87.
PARATYPE: LACM 2293 (1 dry), 51 m, off Freeport, Brazoria Co., Texas (approx. 28 05 N,
94 35 W), Leg.: A. Knight, NW Gulf Survey, 17 DEC 1986.
discors, Lirularia McLean, 1984a: The Veliger 26(3):237, fig. 7.
HOLOTYPE: LACM 1997 (dry); PARATYPES: LACM 1998 (49 dry), 3-9 m, S side of Punta
Banda, near the Blowhole, Pacific Coast Baja California, México (31 43.6 N, 116 43 W), Leg.:
J.H. McLean, 12 AUG 1963.
eloiseae, Umbonium Dance, Moolenbeek, & Dekker, 1992: Jour. Conch. 34(4):231-235.
PARATYPES: LACM 2725 (3 dry), Sur Masirah, W coast of Masirah Id., Oman, Leg.: R.G.
Moolenbeek & H. Dekker, 22 NOV 1991.
felipensis, Tegula (Agathistoma) McLean, 1970a: Malac. Review 2:121, figs. 16-17.
HOLOTYPE: LACM 1383 (dry); PARATYPES: LACM 1384 (15 dry), Punta San Felipe, Golfo
de California, Baja California, México (31 02 N, 114 49 W), Leg.: J.H. McLean, APR
1961-DEC 1962.
galapagensis, Mirachelus McLean, 1970a: Malac. Review 2:118, figs. 9, 11.
HOLOTYPE: LACM 1377 (dry); PARATYPE: LACM 1378 (2 dry), 75-100 m, off Canal
Bolivar, near Caleta Tagus, Isla Isabela (= Albemarle Id.), Islas Galápagos (0 16 S, 91 22 W),
Leg.: A. & J. DeRoy, 23 JAN 1968.
genecoani, Halistylus McLean, 1984a: The Veliger 26(3):237-238, fig. 10.
HOLOTYPE: LACM 1999 (dry); PARATYPES: LACM 2000 (30 dry), 21-24 m, cove at S side
of Punta San Pablo, Pacific Coast Baja California Sur, Mexico (27 12.92 N, 114 27.5 W), Leg.:
C.C. Swift & R. Huddleston, R/V Searcher, 24 OCT 1971.
gordanum, Calliostoma McLean, 1970d: The Veliger 12(4):422-423.
PARATYPE: LACM 1268 (1 dry), 128 m, Gorda Bank, Golfo de California, Baja California Sur,
México (23 02 N, 109 31 W), Leg.: Templeton Crocker Exped., 1936.
guttarosea, Euchelus Dall, 1889: Bull. Mus. Comp. Zool. 18:382.
PARATYPE: LACM 1427 (1 dry), 212 m, Bahía Samana, Santo Domingo Prov., Dominican
Republic, Leg.: W.H. Dall.
harrisi, Gibbula Baker & Spicer, 1930: Trans. San Diego Soc. Nat. Hist. 6(6):177-178.
PARATYPES: LACM 1014 (2 dry), 7 m, Basilan Channel, off Zamboanga, Zamboanga del Sur
Prov., Mindanao Id., Philippines, Leg.: F. Baker, 24 FEB 1914.
Remarks: Currently in the genus Solariella.
hickmanae, Margarites (Margarites) McLean, 1984a: The Veliger 26(3):234-236.
PARATYPES: LACM 1994 (2 dry), 99 m, Petrel Bank, near Semisopochnoi Id., Rat Ids.,
Aleutian Ids. Co., Alaska (52 11 N, 179 57 W), Leg.: R/V Albatross, 5 JUN 1906.
hummelincki, Euchelus Moolenbeek & Faber, 1989: Found. Sci. Res. in Surinam & Neth.
Antilles 123:220-224.
PARATYPE: LACM 2468 (1 dry), 100 m, off Holetown, St. James Parish, Barbados, Leg.: P.W.
Hummelinck, 19 FEB 1964.
jacquelinae, Calliostoma McLean, 1970d: The Veliger 12(4):425.
PARATYPE: LACM 1274 (1 dry), 150 m, Bahía Academy del Sur, Isla Santa Cruz (=
Indefatigable Id.), Islas Galápagos (0 45 S, 90 20 W), Leg.: A. & J. DeRoy, 10 JUN 1969.
keenae, Calliostoma McLean, 1970d: The Veliger 12(4):424-425, pl. 62, fig. 13-14.
HOLOTYPE: LACM 1272 (dry), 106-110 m, Fifty-eight Fathom Bank, 12 mi. off Laguna
Beach, Orange Co., California (33 23.78 N, 117 59.78 W), Leg.: R/V Velero III, 12 FEB 1949.
korkosi, Clanculus Singer, Mienis, & Geiger, 2000: La Conchiglia 32(294-295):33-38.
PARATYPE: LACM 2872 (1 dry); PARATYPE: LACM 2873 (shell dry, animal wet),18 m,
Blue Hole, 11 km N of Dahab, Gulf of Aqaba, Sinai Peninsula, Egypt; PARATYPE: LACM
2874 (2 dry, 1 coated), 20 m, Dahab area, Gulf of Aqaba, Sinai Peninsula, Egypt.
Remarks: Radula of paratype 2873 on SEM stub.
lewisae, Solariella (Solariella) Willett, 1946: Bull. So. Calif. Acad. Sci. 45(1):32, pl. 7.
HOLOTYPE: LACM 1084 (dry), 27 m, Spiridon Bay, Kodiak Id., Kodiak Co., Alaska, Leg.: Mr.
& Mrs. F. Lewis.
Remarks: Junior synonym of Solariella (Solariella) obscurra (Couthouy, 1838).
marshalli, Calliostoma Lowe, 1935: Trans. San Diego Soc. Nat. Hist. 8(6):19.
PARATYPES: LACM 2184 (15 dry), San Felipe, Golfo de California, Baja California, México,
Leg.: Quayle & Glassell.
Remarks: Ex UCLA 5188.
mcleani, Calliostoma Shasky & Campbell, 1964: The Veliger 7(2):117-118.
PARATYPES: LACM 1258 (2 dry), 5.4 m, S of Punta Colorado, Guaymas, Sonora, México,
Leg.: B. Dillworth, JUL 1964.
mendella, Tegula (Agathistoma) McLean, 1964: The Veliger 7(2):131.
PARATYPE: LACM 1135 (1 dry), Mission Bay, San Diego, San Diego Co., California, Leg.:
J.H. McLean, 1962.
Remarks: Junior synonym of Tegula eiseni Jordan, 1936.
nortoni, Agathodonta McLean, 1984b: Nautilus 98(3):122-123, figs. 1-3.
HOLOTYPE: LACM 2031 (dry), 96-320 m, Baltazar Id., Marinduque Prov., Philippines
(13 14 N, 121 49 E).
parvum, Calliostoma canaliculatum Williamson, 1905b: Bull. So. Calif. Acad. Sci. 4(8):128.
SYNTYPES: LACM 2206 (10 dry), San Diego Bay, San Diego Co., California, Leg.: F.W.
Kelsey.
Remarks: Junior synonym of Calliostoma canaliculatum (Lightfoot, 1786).
picta, Tegula (Agathistoma) McLean, 1970a: Malac. Review 2:121-122, figs. 18-19.
HOLOTYPE: LACM 1385 (dry); PARATYPES: LACM 1386 (13 dry), W of Manta, Manabi
Prov., Ecuador (0 56.72 S, 80 44.72 W), Leg.: R/V Velero III, 20 JAN 1935
sanjaimense, Calliostoma McLean, 1970d: The Veliger 12(4):423, pl. 62, fig. 11.
HOLOTYPE: LACM 1269 (dry); PARATYPES: LACM 1270 (2 dry), 137 m, San Jaime Bank,
off Cabo San Lucas, Golfo de California, Baja California Sur, México (22 50.5 N, 110 15 W),
Leg.: R/V Velero III, 3 MAR 1937.
scalenum, Calliostoma Quinn, 1992: The Nautilus 106(3):92-93.
PARATYPE: LACM 2292 (1 dry), 21-27 m, Stetson Bank, SE of Galveston, Galveston Co.,
Texas (28 09.90 N, 94 18.0 W), Leg.: NW Gulf Survey, AUG 1971.
scurra, Calliostoma Quinn, 1992: The Nautilus 106(3):110-111.
PARATYPE: LACM 2253 (1 dry), 33-35 m, off Orinoco River mouth, Delta Amacuro Terr.,
Venezuela (9 04.1 N, 60 10.7 W), Leg.: R/V John Elliott Pillsbury, 30 JUN 1969.
symplector, Bathymargarites Warén & Bouchet, 1989: Zoologica Scripta 18(1):91-94.
PARATYPES: LACM 2288 (7 dry), 2612 m, East Pacific Rise, East Pacific (20 50 N,
106 06 W), Leg.: R/V Alvin, 25 APR 1982.
titanium, Calliostoma McLean, 1984a: The Veliger 26(3):236, fig. 4.
HOLOTYPE: LACM 1995 (shell dry, animal wet), 256-274 m, SE of Santa Catalina Id.,
California Channel Ids., California (33 15.2 N, 118 14 W), Leg.: R/V Velero III, 10 DEC 1939.
Remarks: Currently in the subgenus Otukaia.
toreuta, Cataegis McLean & Quinn, 1987: The Nautilus 101(3):113-115.
PARATYPES: LACM 2264 (2 dry), 604-733 m, off Bolivar Prov., Colombia (10 17 N,
79 59.9 W), Leg.: R/V John Elliott Pillsbury, 14 JUL 1966.
veleroae, Calliostoma McLean, 1970d: The Veliger 12(4):423-424, pl. 62, fig. 12.
HOLOTYPE: LACM 1271 (1 dry), 99 m, 3 mi. S of Isla Ladrones, Chiriqui Prov., Panamá
(7 49 N, 82 23.5 W), Leg.: R/V Velero III, 27 MAR 1939.
verdispira, Tegula (Agathistoma) McLean, 1970a: Malac. Review 2:122, figs. 20-21.
HOLOTYPE: LACM 1387 (dry); PARATYPE: LACM 1388 (1 dry), 0 m, intertidal, S
anchorage, Isla Maria Magdalena, Islas Tres Marias, Nayarit, México (21 35 N, 106 35 W),
Leg.: C. Miller, J.H. McLean, A.M. Keen, R/V Gringa, 15 MAR 1965.
verrucosa, Tegula (Agathistoma) McLean, 1970a: Malac. Review 2:122-123, figs. 22-23.
HOLOTYPE: LACM 1389 (dry); PARATYPES: LACM 1390 (5 dry), 0 m, intertidal, Palo Seco,
Pacific Coast Panama Canal Zone, Panamá (8 55 N, 79 34 W), Leg.: E. Bergeron, 1963.
Superfamily TURBINOIDEA Rafinesque, 1815
Family Turbinidae Rafinesque, 1815
cordellensis, Homalopoma McLean, 1995: Tax. Atlas Benthic Fauna Santa Maria Basin & W.
Santa Barbara Ch. 9(2)(1)28-29, fig. 1.4E.
HOLOTYPE: LACM 2779 (dry); PARATYPES: LACM 2780 (356 dry), 37-55 m, unnamed
bank 3 mi. WSW of Pt. Sur, Monterey Co., California (approx. 36 17 N, 121 58 W), Leg.: R.W.
Schmieder, R/V Cordell Explorer, SEP 1988-OCT 1989.
draperi, Homalopoma (Homalopoma) McLean, 1984a: The Veliger 26(3):238-239, fig. 8.
HOLOTYPE: LACM 2001 (dry); PARATYPES: LACM 2002 (164 dry), 9-30 m, Isthmus Cove,
Santa Catalina Id., California Channel Ids., California (33 26.5 N, 118 29 W), Leg.: J.H.
McLean, JUN-JUL 1971.
engbergi, Leptothyra Willett, 1929b: The Nautilus 43(1):27-28, pl. 3, fig. 4.
HOLOTYPE: LACM 1017 (dry); PARATYPES: LACM 2141 (22 dry), Olga, Orcas Id., San
Juan Ids., San Juan Co., Washington, Leg.: C.C. Engberg.
Remarks: Currently in the genus Homalapoma. Paratypes ex UCLA 20960.
jonathani, Turbo Dekker, Moolenbeek, & Decker, 1992: Jour. Conch. 34(4):225-229.
PARATYPE: LACM 2724 (1 dry), 7 km east of Marbat, Dhofar Prov., Oman, Leg.: R.G.
Moolenbeek & H. Dekker, 14 NOV 1991.
kachemakensis, Spiromoelleria Baxter & McLean, 1984: The Veliger 27(2):223-225, fig. 6.
HOLOTYPE: LACM 1989 (dry); PARATYPES: LACM 1990 (43 dry), 9 m, off the N side of
Hesketh Id., Kachemak Bay, Kenai Peninsula, Kenai-Cook Inlet Co., Alaska (59 30.5 N,
151 31 W), Leg.: J.H. McLean & R. Baxter, 2 AUG 1973.
mimicum, Homalopoma (Homalopoma) LaFollette, 1976: The Veliger 19(1):71-76, figs. 7-10,
26.
HOLOTYPE: LACM 1765 (dry); PARATYPES: LACM 1766 (5 dry), 24-34 m,
Six-and-one-half Fathom Bank, 4 mi. off Isla San Martin, Pacific Coast Baja, California, México
(30 25.38 N, 116 08.20 W), Leg.: J.H. McLean, R/V Searcher, 17 OCT 1971.
morini, Leptothyra Viader, 1951: Mauritius Inst. Bull. 3(2):150.
SYNTYPE: LACM 2804 (1 dry), St. Brandon Id., Cargados Carajos Shoals, Mascarene Ids.,
Indian Ocean (16 38 S, 59 38 E), Leg.: G. Morin & R. Viader.
Family Phasianellidae Swainson, 1840
diantha, Tricolia (Eulithidium) McLean, 1970a: Malac. Review 2:125-126, fig. 34.
HOLOTYPE: LACM 1397 (dry); PARATYPES: LACM 1398 (34 dry), 110 m, E of the S end of
Isla Isabela (= Albemarle Id.), Islas Galápagos (0 55 S, 90 30 W), Leg.: R/V Velero III, 26 JAN
1934.
macleani, Tricolia (Eulithidium) Marincovich, 1973: Nat. Hist. Mus. L.A. Co. Sci. Bull. 16:25,
fig. 47.
HOLOTYPE: LACM 1585 (dry); PARATYPES: LACM 1586 (275 dry), 0 m, intertidal, Patillos,
Tarapaca Prov., Chile (20 48 S, 70 12 W), Leg.: L. Marincovich, JUL 1964.
roni, Gabrielona Moolenbeek & Dekker, 1992: Venus 52(2):143-145.
PARATYPES: LACM 2727 (2 wet), Ra's Janjali, Dhofar Prov., Oman, Leg.: R.G. Moolenbeek
& H. Dekker, 14 NOV 1991.
Family Liotiidae Gray, 1850
adusta, Arene (Arene) McLean, 1970a: Malac. Review 2:123, figs. 24-25.
HOLOTYPE: LACM 1391 (dry); PARATYPES: LACM 1392 (18 dry), 0-1.8 m, intertidal,
Bahía Partida, between Islas Partida & Espíritu Santo, Golfo de California, Baja California Sur,
México (24 25 N, 110 25 W), Leg.: J.H. McLean, 10 APR 1966.
balboai, Liotia Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):236-237.
PARATYPES: LACM 1731 (2 dry), 5-16 m, off Isla Taboga, Panamá Prov., Panamá, Leg.: L.G.
Hertlein.
Remarks: Currently in the genus Marevalvata.
digitata, Macrarene McLean, Absalao, & Cruz, 1988: The Nautilus 102(3):100-101.
PARATYPE: LACM 2377 (1 dry), 146 m, off Cabo San Roque, Rio Grande do Norte State,
Brazil (4 30 S, 50 03 W), Leg.: R/V Almarante Saldanha, 8 MAY 1968.
echinata, Arene (Arene) McLean, 1970b: The Veliger 12(3):311.
PARATYPES: LACM 1277 (2 dry), 0 m, intertidal, Punta Espinosa, Isla Fernandina (=
Narborough Id.), Islas Galápagos, Leg.: J. DeRoy, 30 JAN 1968.
ferruginosa, Arene (Arene) McLean, 1970a: Malac. Review 2:123-124, figs. 26-27.
HOLOTYPE: LACM 1393 (dry), 46 m, S of White Friars, Guerrero, México (17 30.8 N,
101 29.92 W), Leg.: R/V Velero III, 2 MAR 1934.
glassi, Bathyliotina McLean, 1988b: The Veliger 30(4):409-410, fig. 1.
HOLOTYPE: LACM 2298 (dry); PARATYPES: LACM 2299 (2 dry), 90-180 m, Bohol Strait,
Philippines.
guttata, Arene (Arene) McLean, 1970b: The Veliger 12(3):310-311, pl. 46, figs. 1-2.
HOLOTYPE: LACM 1275 (dry); PARATYPES: LACM 1276 (87 dry), 0 m, intertidal rocks,
Bahía Academy, Isla Santa Cruz (= Indefatigable Id.), Islas Galápagos, Leg.: J. DeRoy, 30 MAY
1969.
heimi, Liotia Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):238.
PARATYPE: LACM 1736 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
lepidotera, Macrarene McLean, 1970a: Malac. Review 2:125, figs. 32-33.
HOLOTYPE: LACM 1396 (dry); 128-137 m, Bahía Braithwaite, Isla Socorro, Islas
Revillagigedo, Southern México (18 40.70 N, 110 55.97 W), Leg.: R/V Velero III, 18 MAR
1939.
miniata, Liotia Dall, 1889: Bull. Mus. Comp. Zool. 18:390.
PARATYPES: LACM 1428 (3 dry), Nassau, New Providence, Bahamas, Leg.: W.H. Dall.
spectabilospina, Macrarene Shasky, 1970: The Veliger 13(2):188-189, figs. 1-2.
HOLOTYPE: LACM 1409 (dry), 82 m, Golfo de Tehuantepec, Chiapas, México, Leg.: D.R.
Shasky, 10 JUL 1963.
springsteeni, Pseudoliotina McLean, 1988b: The Veliger 30(4):410, fig. 2.
HOLOTYPE: LACM 2300 (dry); PARATYPE: LACM 2301 (1 dry), 200-350 m, Bohol Strait,
Philippines.
stellata, Arene (Arene) McLean, 1970a: Malac. Review 2:124-125, figs. 30-31.
HOLOTYPE: LACM 1394 (dry); PARATYPES: LACM 1395 (4 dry), 6-9 m, Bahia Partida,
adjoining the W sides of Islas Partida & Espíritu Santo, Golfo de California, Baja California Sur,
México (24 25 N, 110 25 W), Leg.: J.H. McLean, 10 APR 1966.
winslowae, Arene (Arene) Pilsbry & Lowe, 1932a: Proc. Acad. Nat. Sci. Philadelphia 84:86.
PARATYPES: LACM 1262 (3 dry); PARATYPES: LACM 2328 (4 dry), 0 m, intertidal, San
Juan del Sur, Rivas Dept., Nicaragua, Leg.: H.N. Lowe, 1931.
Family Skeneidae Clark, 1851
fallax, Dillwynella Hasegawa, 1997: Nat. Sci. Mus. Tokyo Mem. 12:97-99.
PARATYPES: LACM 2842 (3 dry), 180-450 m, off Heda, Suruga-wan, Shizuoka Pref., Honshu,
Japan (34 59.4 N, 138 43.4 E), Leg.: R/V Soro-Maru, 25 OCT 1993
planorbis, Dillwynella Hasegawa, 1997: Nat. Sci. Mus. Tokyo Mem. 12:94-96.
PARATYPES: LACM 2841 (3 dry), 140-400 m, 3.5 km ENE of Osezaki, Suruga-wan, Shizuoka
Pref., Honshu, Japan (35 00.5 N, 138 45.1 E), Leg.: R/V Soro-Maru.
rehderi, Parviturbo Pilsbry & McGinty, 1945: The Nautilus 59(2):54-55.
PARATYPE: LACM 2233 (1 dry), N inlet to Lake Worth, Palm Beach Co., Florida, Leg. T.L.
McGinty, 7 JUL 1945.
seishinmaruae, Dillwynella Hasegawa, 1997: Nat. Sci. Mus. Tokyo Mem. 12:99-103.
PARATYPES: LACM 2843 (2 dry), 460 m, off Heda, Suruga-wan, Shizuoka Pref., Honshu,
Japan, Leg.: H. Numanami, R/V Kotoku-Maru, 20 FEB 1991.
vitrea, Dillwynella Hasegawa, 1997: Nat. Sci. Mus. Tokyo Mem. 12:90-94.
PARATYPE: LACM 2840 (1 dry), 400-740 m, off Heda, Sugura-wan, Shizuoka Pref., Honshu,
Japan (35 00.4 N, 138 43.2 E), Leg.: R/V Soro-Maru, 17 NOV 1993.
Superfamily SEGUENZIOIDEA Verrill, 1884
Family Seguenziidae Verrill, 1884
inermis, Carencia Quinn, 1983: Proc. Biol. Soc. Washington 96(3):362, figs. 13-14.
HOLOTYPE: LACM 1806 (dry), 2820 m, 181.7 mi. 263.5 degrees T from Cape Perpetua, near
Yachats, Lincoln Co., Oregon (44 33 N, 128 20.2 W), Leg.: Oregon State Univ.; R/V Cayuse, 3
NOV 1973.
megaloconcha, Seguenzia Rokop, 1972: The Veliger 15(1):15.
PARATYPE: LACM 1550 (1 dry), 3592-3798 m, W of Pioneer Seamount and SW of the
Farallon Ids., California (37 22 N, 123 54 W), Leg.: R/V Thomas Washington, 14-15 JUN 1967.
quinni, Seguenzia McLean, 1985a: The Veliger 27(3):337-338, fig. 2.
HOLOTYPE: LACM 2065 (dry), 3900 m, 451.7 mi. 264.3 degrees T from Cape Foulweather,
Tufts Abyssal Plain off Oregon, East Pacific (45 02 N, 134 42.2 W), Leg.: Oregon State Univ.,
R/V Yaquina, 9 OCT 1972.
Subclass Orthogastropoda Ponder & Lindberg, 1998
Clade Cocculiniformia Haszprunar, 1987
Superfamily COCCULINOIDEA Dall, 1882
Family Cocculinidae Dall, 1882
baxteri, Cocculina McLean, 1987:Zool. Scripta 16(4):330, fig. 14.
HOLOTYPE: LACM 2067 (shell dry, animal wet); PARATYPES: LACM 2068 (78 wet),
424-430, Prince William Sound, Valdez-Cordova Co., Alaska (60 42.6 N, 148 04 W), Leg.: R.
Baxter, R/V Oregon, 26 APR 1978.
cowani, Cocculina McLean, 1987: Zool. Scripta 16(4):328-330.
PARATYPES: LACM 2066 (4 wet animals, dry shells), 1370 m, off W side of Moresby Id.,
Queen Charlotte Ids., British Columbia, Canada (52 30.5 N, 131 55 ’W), Leg.: Fisheries Res.
Board of Canada, R/V G.B. Reed, OCT 1966.
craigsmithi, Cocculina McLean, 1992c: Jour. Moll. Stud. 58:402-405, figs.1-8.
HOLOTYPE: LACM 2459 (shell dry, animal wet); PARATYPES: LACM 2460 (2 dry, 6 wet);
PARATYPES: LACM 2461 (shells dry, animals wet), 1240 m, Santa Catalina Basin, between
San Clemente Id. and Santa Catalina Id., California Channel Ids., California (33 12 N,
118 30 W), Leg.: C.R. Smith, R/V Alvin, 10 NOV 1987; PARATYPES: LACM 2391 (5 wet),
1240 m, Santa Catalina Basin, between San Clemente Id. and Santa Catalina Id., California
Channel Ids., California (33 12 N, 118 30 W), Leg.: C.R. Smith, R/V Alvin, 21 FEB 1991;
PARATYPES: LACM 2392 (5 wet, 3dry shells), 1240 M, Santa Catalina Basin, between San
Clemente Id. and Santa Catalina Id., California Channel Ids., California (33 12 N, 118 30 W),
Leg.: C.R. Smith, R/V Alvin, 22 FEB 1991.
crinita, Coccopigya Marshall, 1985b: New Zealand Jour. Zool. 12(4):515-516.
PARATYPES: LACM 2767 (3 wet), Chatham Rise, SE of Cape Pelliser, Wellington, North Id.,
New Zealand (42 43 S, 176 08 E), Leg.: R/V Kaltan, 28 SEP 1982.
emsoni, Cocculina McLean & Harasewych, 1995: Nat. Hist. Mus. L.A. Co. Cont. Sci. 453:
13-14.
PARATYPE: LACM 2736 (1 dry), 518 m, Off Southwest Reef, New Providence Id., Bahamas
(24 54.04 N, 77 33.14 W), Leg.: R. Emson, C.M. Young, & P.A. Tyler, R/V Johnson-Sea-Link
II, 9 MAY 1992.
hispida, Coccopigya Marshall, 1985b: New Zealand Jour. Zool. 12(4):513-514.
PARATYPE: LACM 2766 (wet), 1198-1211 m, off Castlepoint, Wellington, North Id., New
Zealand (41 09.7 S 176 31.3 E), Leg.: R/V James Cook, 4 APR 1984 .
messingi, Cocculina McLean & Hararsewych, 1995: Nat. Hist. Mus. L.A. Co. Cont. Sci. 453:
11-13.
PARATYPE: LACM 2735 (1 dry), 412 m, S of Settlement Pt., Grand Bahama Id., Bahamas.
(26 37.52 N, 78 58.93 W), Leg.: C. Messing, R/V Johnson-Sea-Link II, 18 MAY 1992.
moskalevi, Macleaniella Leal & Harasewych, 1999: Invert. Biol. 118(2):123-127.
PARATYPES: LACM 2807 (3 wet), 7961-7970 m, western Puerto Trench, N of Puerto Rico
(19°42.5'N, 67°5'W), Leg.: R/V Pillsbury, 21 JAN 1970.
okutanii, Coccopigya Hasegawa, 1997: Nat. Sci. Mus. Tokyo, Memoirs 12:79-81.
PARATYPES: LACM 2838 (3 dry shells, animals wet), 275-500 m, 5.1 km off Kumomi,
Suruga-wan, Shizuoka Pref., Honshu, Japan (34 46.3 N, 138 42.8 E), Leg.: R/V Soro-Maru, 23
OCT 1994.
Clade Cycloneritimorpha
Superfamily HELICINOIDEA Férussac, 1822
Family Helicinidae Férussac, 1822
chionea, Helicina (Helicina) Pilsbry, 1949: The Nautilus 62(3):101.
PARATYPES: LACM 2052 (5, dry), 2900 m, Mejorada, near Huancayo, Rio Mantaro, Pasco
Dept., Peru, Leg.: W. Weyrauch.
Clade Caenogastropoda Cox, 1959
Zygopleuroid Group
Family Provannidae Warén & Ponder, 1991
admetoides, Provanna Warén & Ponder, 1991: Zoologica Scripta 20(1):54-55.
PARATYPES: LACM 2442 (2 dry), 624-631 m, W of St. Petersburg, Pinellas Co., Florida
(28 08 N, 85 52 W), Leg.: Mineral Management Services, 16 MAY 1985.
laevis, Provanna Warén & Ponder, 1991: Zoologica Scripta 20(1):53.
PARATYPES: LACM 2443 (3 dry), 004 m, Guaymas Basin, Golfo de California, México
(27 00.7 N, 111 24.4 W), Leg.: R/V Alvin, 6 AUG 1985.
macleani, Provanna Warén & Bouchet, 1989: Zoologica Scripta 18(1):94-95, fig. 112.
HOLOTYPE: LACM 2289 (dry); PARATYPES: LACM 2290 (3 dry), 2713 m, off Cape
Foulweather, Cascadia Abyssal Plain off Oregon, East Pacific (44 52.4 N, 126 40.8 W), Leg.:
Oregon State Univ., R/V Yaquina, 16 JAN 1970.
Remarks: Radula of paratype mounted on glass slide.
nautilei, Ifremeria Bouchet & Warén, 1991: Comp. Rend. Seances de L'Acad. Sci., Paris, 3rd
ser., Sci. Vie 312:495-501.
PARATYPES: LACM 2245 (2 wet), 1750 m, Vailili Site, Valufa Ridge, Lau Basin, SE of Fiji
(23 13 S, 176 38 W), Leg.: Biolau Exped., R/V Nautile, 12 MAY, 1989.
sculpta, Provanna Warén & Ponder, 1991: Zoologica Scripta 20(1):53-54.
PARATYPES: LACM 2444 (2 dry), 576 m, "Bush Hill" hydrocarbon seep, Louisiana Slope,
Gulf of Mexico (27 46.94 N, 90 30.34 W), Leg.: R/V Johnson-Sea-Link I, 28 SEP 1986.
segonzaci, Provanna Warén & Ponder, 1991: Zoologica Scripta 20(1):51-52.
PARATYPES: LACM 2445 (2 dry), 900 m, Valufa Ridge, Fiji Back-Arc, Lau Basin, SE of Fiji
(22 32 S, 176 43 W), Leg.: Biolau Exped.
variabilis, Provanna Warén & Bouchet, 1986: Zoologica Scripta 15(2):163-164.
PARATYPE: LACM 2291 (1 dry), 2212 m, Endeavor Segment, Juan de Fuca Ridge off,
Washington, East Pacific (47 57 N, 129 04 W), Leg.: R/V Alvin, 25 JUL 1984.
Informal Group Architaenioglossa Haller, 1892
Superfamily CYCLOPHOROIDEA Gray, 1847
Family Cyclophoridae Gray, 1847
barbouri, Farcimen majusculum Alcalde, 1945a: Revista Sociedad Malac. 3(1):16.
PARATYPES: LACM 1637 (2 dry), Ceno de Bastillos, La Sierra, Cienfuegos, Cienfuegos Prov.,
Cuba.
Remarks: Currently in the genus Megalomastoma.
cisnerosi, Farcimen Alcalde, 1945b: Revista Sociedad Malac. 3(2):45-46.
PARATYPES: LACM 1642 (5 dry), Quemado de los Cedros, San Jose, La Sierra, Cienfuegos,
Cienfuegos Prov., Cuba.
Remarks: Currently in the genus Megalomastoma.
corralilloense, Farcimen auriculatum Alcalde, 1945a: Revista Sociedad Malac. 3(1):12-13.
PARATYPES: LACM 1636 (2 dry), Corralillo, Guaos, Cienfuegos, Cienfuegos Prov., Cuba.
Remarks: Currently in the genus Megalomastoma.
deflexa, Cyclophorus (Eucyclotus) auriculatus var. Möllendorff, 1893: Seneck. natur. Gess.
Berichte, p. 108.
PARATYPE: LACM 2620 (1 dry), Jaro, Leyte Prov., Philippines.
edouardi, Farcimen cisnerosi Alcalde, 1945b: Revista Sociedad Malac. 3(2):46.
PARATYPE: LACM 1643 (1 dry), Baulla, San Juan, Trinidad, Sancti Spiritus Prov., Cuba.
Remarks: Currently in the genus Megalomastoma.
imperator, Farcimen Alcalde, 1945b: Revista Sociedad Malac. 3(2):44.
PARATYPES: LACM 1641 (2 dry),Valle de Delicias, Siguanea, Villa Clara Prov., Cuba.
Remarks: Currently in the genus Megalomastoma.
jaumei, Farcimen Alcalde, 1945b: Revista Sociedad Malac. 3(2):43.
PARATYPES: LACM 1639 (5 dry), Varallones de Briche, Yaguanabo, Trinidad, Sancti Spiritus
Prov., Cuba.
Remarks: Currently in the genus Megalomastoma.
kibleri, Cyclophorus (Cyclohelix) Fulton, 1907: Ann. Mag. Nat. Hist., 7th ser., 19:156.
SYNTYPES: LACM 2618 (2 dry), Nias Id., Kepulauan Mentawai, Indonesia.
martensianus, Cyclophorus (Cyclophorus) Möllendorff, 1875: Jahr. Deut. Malak. Gess. 2:120.
PARATYPE: LACM 2621 (1 dry), Hubei Prov., China.
obtecta, Cyclophorus (Cricophorus) philippinarum var. Kobelt & Möllendorff, 1898: Nachr.
Deut. Malak. Gess. 29:111.
PARATYPES: LACM 2622 (4 dry), Ilocos Id., NW Luzon Id., Philippines, Leg.: O.F.
Möllendorff.
Remarks: Junior synonym of Cyclophoras philippinarum (Sowerby, 1842).
parvulum, Farcimen jaumei Alcalde, 1945b: Revista Sociedad Malac. 3(2):44.
PARATYPES: LACM 1640 (5 dry), Agunda de la Botella, Cumanayagua, Villa Clara Prov.,
Cuba.
Remarks: Currently in the genus Megalomastoma.
rugulosum, Farcimen guitarti Alcalde, 1945a: Revista Sociedad Malac. 3(1):42-43.
PARATYPES: LACM 2041 (4 dry), Las Veguetas del Cuajani, Buenos Aires, Cienfuegos Prov.,
Cuba.
Remarks: Currently in the genus Megalomastoma.
Family Neocyclotidae Kobelt & Möllendorff, 1897
chocolatum, Incidostoma Morrison, 1955: Jour. Washington Acad. Sci. 45(5):158-159.
PARATYPE: LACM 1654 (1 dry), Papallacta, Napo-Pastaza Prov., Ecuador.
diminutum, Incidostoma Morrison, 1955: Jour. Washington Acad. Sci. 45(5):159.
PARATYPES: LACM 1655 (2 dry), Papallacta, Napo-Pastaza Prov., Ecuador.
jacksoni, Incidostoma Morrison, 1955: Jour. Washington Acad. Sci. 45(5):158.
PARATYPES: LACM 1656 (3 dry), Near Mera, Napo-Pastaza Prov., Ecuador.
Clade Sobreoconcha Ponder & Lindberg, 1997
Superfamily CERITHIOIDEA Fleming, 1822
Family Cerithiidae Fleming, 1822
scobiniforme, Cerithium Houbrick, 1992: Smithsonian Contr. to Zool. 510:177-179.
PARATYPES: LACM 2654 (6 dry), 2-9 m, 1.5 mi. S of Port Dimalonsa, Estagno Id., Cagayan
Prov., Luzon Id., Philippines, Leg.: J. Norton, 26 JUN 1964.
Family Diastomatidae Cossmann, 1894
veraguaensis, Alabina Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):218-219.
PARATYPE: LACM 1711 (2 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
Family Pachychilidae Fischer & Cross, 1892
clenchi, Goniobasis Goodrich, 1924: The Nautilus 38(2):46-48.
PARATYPES: LACM 1649 (3 dry), Choctawatchee River, Dale Co., Alabama, Leg.: W.J.
Clench, 23 JUL 1923.
Remarks: Currently in the genus Elimia.
praerosa, Leptoxis (Leptoxis) Say, 1821: Jour. Acad. Nat. Sci. Philadelphia 2(1):177-178.
PARATYPE: LACM 2021 (1 dry), Ohio River, United States.
Family Turritellidae Lovén, 1847
ahiparana, Turritella Powell, 1927b: Trans. Proc. New Zealand Inst. 58(3):297.
PARATYPES: LACM 1319 (18 dry); PARATYPES: LACM 2057 (7 dry), 42 m, off Ahipara,
Northland, North Id., New Zealand.
Remarks: Currently in the genus Zeacolpus.
anactor, Turritella Berry, 1957: Leaflets in Malacology 1(14):78.
PARATYPE: LACM 1447 (1 dry), Playa de Almejas, N side of San Felipe, Golfo de California,
Baja California, México, Leg.: F.B. Howard, APR 1957.
frisbeyae, Vermicularia McLean, 1970b: The Veliger 12(3):311-312, pl. 46, figs. 5-6.
HOLOTYPE: LACM 1278 (dry); PARATYPE: LACM 1279 (1 dry), 55-73 m, off the lighthouse
at Manzanillo, Colima, México (19 03 N, 104 20 W), Leg.: J. Frisbey, FEB 1969.
kowiensis, Turritella Sowerby, 1900: Proc. Malac. Soc. London 4(1):6.
SYNTYPES [as "COTYPES"]: LACM 1322 (2 dry), Port Alfred (= The Kowie), Cape Prov.,
South Africa.
lyonsi, Turritella Garcia, 2006: Novapex 7(4):78-79.
PARATYPES: LACM 3072 (2 dry), 28-48 m, Bahía de Campeche, Golfo de México, NE of
Campeche, Campeche, México (20 46.97 N, 91 55.86 W), Leg.: E.F. Garcia, 11 JUN 2005.
parkeri, Turritella McLean, 1970a: Malac. Review 2:127, fig. 37.
HOLOTYPE: LACM 1401 (dry); PARATYPES: LACM 1402 (24 dry), 82-119 m, Bahía de La
Paz, W of Isla Espíritu Santo, Golfo de California, Baja California Sur, México (24 24.3 N,
110 23.7 W), Leg.: R/V Thomas Washington.
willetti, Turritella McLean, 1970b: The Veliger 12(3):312-313, pl. 46, figs. 7-8.
HOLOTYPE: LACM 1280 (dry); PARATYPES: LACM 1281 (2 dry), 13-22 m, Bahía Santiago,
Manzanillo, Colima, México (19 06 N, 104 23 W), Leg.: L. & C. Shy, NOV-DEC 1968.
Family Siliquariidae Anton, 1838
levispinosum, Stephopoma Bieler, 1997: Mar. Flora Fauna Houtman Abrolhos Ids., Aust., p.
271-272.
PARATYPE: LACM 2802 (1 dry), 3 m, Isla Taboguilla, Bahía de Panamá, Panamá Prov.,
Panamá (8°48'N, 79°31'W), Leg.: R. Bieler, 5 AUG 1983.
Clade Littorinimorpha
Superfamily LITTORINOIDEA Children, 1834
Family Littorinidae Children, 1834
albicarinata, Littorina (Littorina) McLean, 1970a: Malac. Review 2:127, fig. 36.
HOLOTYPE: LACM 1399 (dry); PARATYPES: LACM 1400 (263 dry), El Requeson, Bahía
Concepción, Golfo de California, Baja California Sur, México (26 38 N, 111 50 W), Leg.: R.
Maynard, AUG 1963.
atkana, Littorina (Littorina) sitkana Dall, 1886: Proc. USNM 9(562):211-212.
PARATYPES: LACM 1203 (3 dry), Kiska Id., Rat Ids., Aleutian Ids. Co., Alaska, Leg.: W.H.
Dall.
Family Pickworthiidae Iredale, 1917
bollandi, Reynellona LeRenard & Bouchet, 2003: Zoosystema 25(4):582-584, fig. 8.
HOLOTYPE: LACM 2455 (dry); PARATYPES: LACM 2456 (13 dry), 46-55 m, 1 km WNW of
Onna Village, Horseshoe Cliffs, Okinawa-jima (26˚30 N, 127˚51 E).
circumserrata, Mareleptopoma Raines, 2002a: La Conchiglia 34(304):18-19, fig. 13.
HOLOTYPE: LACM 2938 (dry); PARATYPE: LACM 2939 (1 dry), 15 m, in cave off HangaTeo, Isla de Pascua (= Easter Id.), Leg.: B.K. Raines.
Family Pomatiidae Newton, 1891
alcaldei, Chondrothyrium (Plicathyrium) Jaume & Fuentes, 1943b: Revista Sociedad Malac.
1(1):24-25.
PARATYPES: LACM 1560 (4 dry), Palmarito de Gonzalez, La Sierra Cienfuegos, Cienfuegos
Prov., Cuba.
affinis, Chondrothyretes Torre & Bartsch, 1938: Proc. USNM 85(3039):303-304.
PARATYPES: LACM 1355 (7 dry), Dos Hermanos marker, Vinales, Pinar del Rio Prov., Cuba.
aguayoi, Chondrothyrium (Plicathyrium) torrei Jaume & Fuentes, 1943c: Revista Sociedad
Malac. 1(1):30-31.
PARATYPES: LACM 1561 (3 dry), Hoyo del Infierno, San Blas, Cienfuegos, Cienfuegos Prov.,
Cuba.
arubana, Tudora fossor H.B. Baker, 1924: Occas. Pap. Mus. Zool., Univ. Mich. 152:68-69.
PARATYPES: LACM 1672 (27 dry), Spur of Seroe Pretoen, between Rolis Spoki and Hundoe,
Aruba, Netherlands Antilles, Leg.: H.B. Baker, SUM 1962.
Remarks: Currently in the genus Licina.
barbouri, Chondrothyretes Torre & Bartsch, 1938: Proc. USNM 85(3039):299-300.
PARATYPE: LACM 1349 (5 dry), Luis Lazo, Pinar del Rio Prov., Cuba.
boqueronensis, Annularia (Fossularia) Torre & Bartsch, 1941: Proc. USNM 89(3096):309-310,
pl. 34, fig. 6.
PARATYPES: LACM 1635 (2 dry), Boqueron del Jatibonico, Guantanamo Prov., Cuba.
britoi, Eutudora limbifera Jaume & Fuentes, 1943a: Revista Sociedad Malac. 1(1):19-20.
PARATYPES: LACM 2042 (4 dry), El Hatillo, San Miguel de los Banos, Matanzas Prov., Cuba,
Leg.: L. Fuentes.
capestanyi, Annularia (Bermudezia) Torre & Bartsch, 1941: Proc. USNM 89(3096):358-359.
PARATYPES: LACM 1353 (3 dry), Palenque de Taguayabon, near Remedios, Villa Clara Prov.,
Cuba.
cerina, Chondrothyretes Torre & Bartsch, 1938: Proc. USNM 85(3039):298-299.
SYNTYPES [as COTYPES]: LACM 1350 (6 dry), Sitio del Infierno, Sierro del Iniferno,
Vinales, Pinar del Rio Prov., Cuba.
emmae, Torrella (Torrella) Jaume & Fuentes, 1943d: Revista Sociedad Malac. 1(1):32-34.
PARATYPES: LACM 2054 (3 dry), Loma de Matanegro, Finca "Odisio," between Coliseo and
Jovellanos, Matanzas Prov., Cuba.
fuentesi, Chondropoma (Chondropoma) Jaume & Alcalde, 1944: Revista Sociedad Malac.
2(2):39-40.
PARATYPES: LACM 1638 (4 dry), Finca de Lewis, Guanaroca, Cienfuegos, Cienfuegos Prov.,
Cuba.
fuentesi, Chondrothyrium crenimargo Alcalde, 1943: Revista Sociedad Malac. 1(1):11-13.
PARATYPE: LACM 1559 (2 dry), Loma #3 del Potrero "Maranon," Cienfuegos, Cienfuegos
Prov., Cuba.
grandiensis, Tudora muskusi H.B. Baker, 1924: Occas. Pap. Mus. Zool., Univ. Michigan
152:53-54.
PARATYPES: LACM 1673 (45 dry), N half of Seroe Grandi, Curaçao, Netherlands Antilles
Leg.: H.B. Baker, SUM 1922,
Remarks: Currently in the genus Licina.
obilterata, Annularia (Bermudezia) Torre & Bartsch, 1941: Proc. USNM 89(3096):358.
PARATYPE: LACM 1354 (1 dry), Cerro de Guajbana, Villa Clara Prov., Cuba.
pilsbryi, Tudora H.B. Baker, 1924: Nautilus 37(3):94.
PARATYPES: LACM 1670 (18 dry), Tafelberg, Santa Barbara, Curaçao, Netherlands Antilles.
Remarks: Currently in the genus Licina.
spreitensis, Tudora megachelios H.B. Baker, 1924: Occas. Pap. Mus. Zool., Univ. Michigan
152:58-59.
PARATYPES: LACM 1674 (20 dry), Seroe Spreit, Curaçao, Netherlands Antilles, Leg.: H.B.
Baker, SUM 1922.
Remarks: Currently in the genus Licina.
wassauensis, Tudora aurantia H.B. Baker, 1924: Occas. Pap. Mus. Zool., Univ. Michigan
152:48-49.
PARATYPES: LACM 1671 (28 dry), Seroe Wassau, W of the entrance to the Goto, Bonaire,
Netherlands Antilles, Leg.: H.B. Baker, SUM 1922.
Remarks: Currently in the genus Licina.
Superfamily CINGULOPSOIDEA Fretter & Patil, 1958
Family Cingulopsidae Fretter & Patil, 1958
atacamae, Eatonina (Saginofusca) Marincovich, 1973: Nat. Hist. Mus. L.A. Co. Sci. Bull.
16:27-28, fig. 52.
HOLOTYPE: LACM 1589 (dry); PARATYPES: LACM 1590 (2374 dry), 0 m, intertidal,
Iquique, Tarapaca Prov., Chile (20 13 S, 70 10 W), Leg.: L. Marincovich, AUG 1964.
hallae, Skenella Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci. 446:21, fig.
15A.
HOLOTYPE: LACM 2672 (dry); PARATYPES: LACM 2673 (8 dry), 17 m, Punta Valparaiso,
Canal Cockburn, Magallanes Prov., Chile (54 22.2 S, 71 21.7 W), Leg.: P. Dayton, R/V Hero,
17 MAY 1973.
Family Eatoniellidae Ponder, 1965
castanea, Eatoniella (Eatoniella) Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib.
Sci. 446:15, fig. 11H.
HOLOTYPE: LACM 2664 (dry); PARATYPES: LACM 2665 (6 dry, 28 wet), Intertidal, NW
arm Bahía York, Isla de los Estados, Tierra del Fuego, Argentina (54 47.15 S, 64 17.9 W),
Leg.: R/V Hero, 5 MAY 1971.
denticula, Eatoniella (Eatoniella) Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib.
Sci. 446:7-9, 11, fig. 5B.
HOLOTYPE: LACM 2658 (dry); PARATYPES: LACM 2659 (9 wet), Intertidal, Puerto Cook,
Isla de los Estados, Tierra del Fuego, Argentina (54 45.25 S, 64 2.3 W), Leg.: R/V Hero, 17
MAY 1971.
ebenina, Eatoniella (Eatoniella) Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib.
Sci. 446:12, fig. 5E.
HOLOTYPE: LACM 2660 (dry); PARATYPES: LACM 2661 (12 dry), 14 m, Bahía Tom,
Magallanes Prov., Chile (50 11.3 S, 74 47.9 W), Leg.: P. Dayton, R/V Hero, 21 MAY 1971.
glomerosa, Eatoniella (Eatoniella) Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib.
Sci. 446:15, 17-18, fig. 11A.
HOLOTYPE: LACM 2666 (dry); PARATYPES: LACM 2667 (28 dry, 202 wet), Intertidal,
Puerto Cook, Isla de los Estados, Tierra del Fuego, Argentina (54 45.25 S, 64 02.3 W), Leg.:
R/V Hero.
latina, Eatoniella (Eatoniella) Marincovich, 1973: Nat. Hist. Mus. L.A. Co. Sci. Bull., 16:26-27
fig. 51.
HOLOTYPE: LACM 1587 (dry); PARATYPES: LACM 1588 (100 dry), Intertidal, Iquique,
Tarapaca Prov., Chile (20 13 S, 70 10 W), Leg.: L. Marincovich, AUG 1964.
magellanica, Pupatonia Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci.
446:19-21, fig. 11I.
HOLOTYPE: LACM 2670 (dry); PARATYPES: LACM 2671 (3 wet), Intertidal, Bahía
Crossley, Isla de los Estados, Tierra del Fuego, Argentina (54 46.2 S, 64 42.7 W), Leg.: R/V
Hero, 27 APR 1971; PARATYPES: LACM 2176 (5 dry), 23 m, Isla Westhoff, Chiloe Prov.,
Chile (43 54 S, 73 43.5 W), Leg.: P. Dayton, R/V Hero, 25 MAY 1973.
mcleani, Eatoniella (Albosabula) Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib.
Sci. 446:19, fig. 11G.
HOLOTYPE: LACM 2668 (dry); PARATYPES: LACM 2669 (295 dry), Intertidal, Los Molles,
Aconcagua Prov., Chile (32 14 S, 71 32 W), Leg.: J.H. McLean, 16-18 OCT 1975.
picea, Eatoniella (Eatoniella) Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci.
446:12-14, fig. 5D.
HOLOTYPE: LACM 2662 (dry); PARATYPES: LACM 2663 (100 wet), Intertidal, Isla
Observatorio, N of Isla Estados, Tierra del Fuego, Argentina (54 39.5 S, 64 08 W), Leg.: R/V
Hero, 19 OCT 1971.
turricula, Eatoniella (Eatoniella) Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib.
Sci. 446:6-7, fig. 5A.
HOLOTYPE: LACM 2656 (dry); PARATYPES: LACM 2657 (69 dry, 136 wet), 135-137 m, 6.4
km N of Cabo San Juan, Isla de los Estados, Tierra del Fuego, Argentina, (54 39.1 S,
63 50.1 W), Leg.: R/V Hero, 26 OCT 1971.
zigzag, Eatoniella (Eatoniella) Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci.
446:55-56.
PARATYPE: LACM 2700 (1 dry), 9-12 m, Isla Juan Fernández, Eastern Pacific (33 42 S,
78 55 W), Leg.: S. Earle, 15 DEC 1965
Superfamily RISSOOIDEA Gray, 1847
Family Rissoidae Gray, 1847
algida, Onoba (?) Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci. 446:47, figs.
20G, 27A.
HOLOTYPE: LACM 2692 (dry); PARATYPES: LACM 2693 (14 dry), 67-71 m, Bahía Capitan
Canepa, Isla de los Estados, Tierra del Fuego, Argentina (54 50.2 S, 64 29.4 W), Leg.: R/V
Hero, 4 NOV 1971.
averni, Pusillina (Haurakia) Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci.
446:23-25, figs. 16A-B.
HOLOTYPE: LACM 2674 (dry); PARATYPES: LACM 2675 (5 dry), 38 m, NW arm Bahía
York, Isla de los Estados, Tierra del Fuego, Argentina (54 47.2 S, 64 18.4 W), Leg.: R/V Hero,
1 NOV 1971.
basteriae, Cingula (Cingula) Moolenbeek & Faber, 1986: Basteria 50(4-6):177-179.
PARATYPES: LACM 2273 (5 dry), Playa de Montana Roja, Lanzarote Id., Canary Ids., Leg.:
R.G. Moolenbeek & W.C.M. van der Hijden, SEP 1984.
bermudensis, Alvania Faber & Moolenbeek, 1987: Beaufortia 37(4):69-70.
PARATYPES: LACM 2350 (2 dry), Gibbons Bay, Bermuda, Leg.: A.T. Guest, APR 1985.
Remarks: Currently in the genus Manzonia.
boucheti, Rissoina (Rissoina) Sleurs in Crosnier & Bouchet, 1991: Resultats des Campagnes
MUSORSTOM, v. 7, Mem. Mus. Natn. Hist. Nat., ser. A, 150:164-174.
PARATYPE: LACM 2702 (1 dry), 440-450 m, SE of New Caledonia (22 47 S, 167 14 E),
Leg.: Biocal Exped., R/V New-Charcot, 30 AUG 1985.
delicatissima, Rissonia, 2002a: La Conchiglia 34(304):16, fig. 9.
HOLOTYPE: LACM 2934 (dry); PARATYPE: LACM 2935 (1 dry), 10-20 m, off Punta Rosalia,
E of Anakena, Isla de Pascua (= Easter Id.), Leg.: B.K. Raines.
erugata, Onoba Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci. 446:45-46,
fig. 19G.
HOLOTYPE: LACM 2690 (dry); PARATYPES: LACM 2691 (32 dry, 62 wet), Intertidal, W
side Bahía Buen Suceso, Tierra del Fuego, Argentina (54 47.8 S, 65 16 W), Leg.: R/V Hero, 23
APR 1971.
eyerdami, Cingula Willett, 1934a: The Nautilus 47(3):103, pl. 8, fig. 9.
HOLOTYPE: LACM 1037 (dry), Elrington Id., Prince William Sound, Valdez-Cordova Co.,
Alaska, Leg.: W.J. Eyerdam; PARATYPES: LACM 1174 (3 dry), Evans Id., Prince William
Sound, Valdez-Cordova Co., Alaska, Leg.: W.J. Eyerdam, JUL 1924; PARATYPES: LACM
2142 (11 dry), Elrington Id., Prince William Sound, Valdez-Cordova Co., Alaska, Leg.: W.J.
Eyerdam, JUL 1924.
Remarks: Currently in the genus Onoba. Junior synonym of Onoba kyskensis (Bartsch, 1911).
Paratype lot LACM 2142 ex UCLA 21372.
forresterensis, Cingula Willett, 1934a: The Nautilus 47(3):103-104, pl. 8, fig. 8.
HOLOTYPE: LACM 1038 (dry); PARATYPES: LACM 2143 (7 dry), Forrester Id., Prince of
Wales-Outer Ketchikan Co., Alaska, Leg.: G. Willett.
Remarks: Junior synonym of Onoba cerinella (Dall, 1886). Paratypes ex UCLA 21368.
guesti, Alvania (Flemellia) Faber & Moolenbeek, 1987: Beaufortia 37(4):70-71.
PARATYPES: LACM 2349 (2 dry), Gibbons Bay, Bermuda, Leg.: A.T. Guest, APR 1985.
hannai, Rissoina Smith & Gordon, 1948: Proc. Calif. Acad. Sci., 4th ser., 26(8):236.
PARATYPES: LACM 1004 (2 dry), 46 m, Carmel Bay, Monterey Co., California, Leg.: W.L.
Scofield.
Remarks: Currently in the genus Schwartziella.
lacuniformis, Onoba (?) Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci.
446:47-48, figs. 20A, C, 27B, E.
HOLOTYPE: LACM 2694 (dry); PARATYPES: LACM 2695 (2 dry), Intertidal, Puerto Cook,
Isla de los Estados, Tierra del Fuego, Argentina (54 45.25 S, 64 2.3 W), Leg.: R/V Hero, 17
MAY 1971.
limensis, Manzonia (Alvinia) Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci.
446:25, figs. 16D-E.
HOLOTYPE: LACM 2676 (dry); PARATYPES: LACM 2677 (2 dry), 6-12 m, Isla Chincha
Norte, Ica Prov., Peru (13 38 S, 76 25 W), Leg.: J.H. McLean & V. Alamo, 1 APR 1972 .
microlirata, Powellisetia Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci.
446:26, fig. 17B.
HOLOTYPE: LACM 2678 (dry); PARATYPES: LACM 2679 (175 dry), 14 m, Bahía Tom,
Magallanes Prov., Chile (50 11.3 S, 74 47.9 W), Leg.: P. Dayton, R/V Hero, 21 MAY 1973.
myriosirissa, Lapsigyrus Shasky, 1970: The Veliger 13(2):189, fig. 3.
HOLOTYPE: LACM 1410 (dry), 0 m, intertidal, Punta Tiburon, Mazatlán, Sinaloa, México
(23 12.5 N, 106 26.5 W), Leg.: D.R. Shasky, 22 DEC 1962.
okinawensis, Rissoina (Moerchiella) Sleurs, 1993: Bull. Inst. Roy. Sci. Nat. Belg., Biol.
63:97-98, figs. 23A-B.
HOLOTYPE: LACM 2729 (dry); PARATYPES: LACM 2730 (5 dry),2 km SSW of Fitzwoody
Beach, Naha Air Force Base, Okinawa-jima, Ryukyu-shoto, Japan (26 11.1 N, 127 38.0 E),
Leg.: R.F. Bolland, 30 SEP 1977; PARATYPES: LACM 2731 (12 dry), "Onna Flats," 1.5 km
WSW of Onna, Okinawa-jima, Ryukyu-shoto, Japan (26 29.3 N, 127 50.2 E), Leg.: R.F.
Bolland, 24 SEP & 14 OCT 1977; PARATYPES: LACM 2732 (2 dry), Fitzwoody Beach, Naha
Air Force Base, Okinawa-jima, Ryukyu-shoto, Japan (26 11.6 N, 27 38.4 E), Leg.: R.F. Bolland,
30 SEP 1977; PARATYPES: LACM 2733 (5 dry), 1 km W of Onna, Okinawa-jima,
Ryukyu-shoto, Japan (26 29.7 N, 127 50.6 E), Leg.: R.F. Bolland, 27 MAY 1978.
protofimbriata, Onoba Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci.
446:26-27, 30-31, figs. 19F, 20B, D.
HOLOTYPE: LACM 2680 (dry), PARATYPES: LACM 2681 (11 dry), Subtidal, Bahía San
Andres, N of Golfo de Penas, Aisen Prov., Chile (46 35.5 S, 75 30.6 W), Leg.: P. Dayton, R/V
Hero, 23 MAY 1973.
striola, Onoba Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci. 446:36, figs.
25A-F.
HOLOTYPE: LACM 2684 (dry); PARATYPES: LACM 2685 (2 dry, 2 wet), Intertidal, Puerto
Cook, Isla de los Estados, Tierra del Fuego, Argentina (54 45.25 S, 64 2.3 W), Leg.: R/V Hero.
subaedonis, Onoba Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci. 446:36-39,
figs. 26B-C.
HOLOTYPE: LACM 2686 (dry); PARATYPES: LACM 2687 (33 dry), Subtidal, Bahía San
Andres, N of Golfo de Penas, Aisen Prov., Chile (46 35.3 S, 75 30.6 W), Leg.: P. Dayton, R/V
Hero, 23 MAY 1973.
subincisa, Onoba Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci. 446:31, 34,
36, figs. 19E.
HOLOTYPE: LACM 2682 (dry); PARATYPES: LACM 2683 (37 dry), Intertidal, Punta Santa
Ana, Fuerte Bulnes, Peninsula Brunswick, Estrecho Magallanes, Magallanes Prov., Chile
(53 38 S, 70 54.5 W), Leg.: J.H. McLean, 16 NOV 1975.
sulcula, Onoba Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci. 446:39, figs.
20E-F, 27C.
HOLOTYPE: LACM 2688 (dry); PARATYPE: LACM 2689 (1 dry), 771-903 m, 8 km S of
Punta Ventana, S side of Isla de los Estados, Tierra del Fuego, Argentina (54 54.5 S, 63 56 W),
Leg.: R/V Hero, 27 OCT 1971; PARATYPES: LACM 2177 (10 dry), 121 m, off Cabo de
Hornos, Magallanes Prov., Chile (56 19.3 S, 67 09.45 W), Leg.: Discovery Exped., 16 APR
1930.
Family Barleeidae Gray, 1857
meridionalis, Barleeia Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib. Sci.
446:48-49, figs. 27F-G.
HOLOTYPE: LACM 2696 (dry); PARATYPES: LACM 2697 (4 dry), 0-4.5 m, Rocky point N
of naval base, midway on NE side Isla San Lorenzo, Lia Prov., Perú (12 5.75 S, 77 12.9 W),
Leg.: J.H. McLean, J.A. Coyer, & J.M. Engle, 29 JAN 1974; PARATYPES: LACM 2754 (9
dry), 0-6 m, Pucusana (Chilca), Lima Prov., Perú (12 30 S, 76 49 W), Leg.: J.H. McLean,
MAR-APR 1972.
ovata, Scrobs Powell, 1927a: Trans. New Zealand Inst. 57:546.
PARATYPES: LACM 1362 (2 dry), 22 m, off Awanui, Ranganu Bay, Northland, North Id., New
Zealand.
Remarks: Currently in the genus Anabathron.
stephensae, Amphithalamus (Amphithalamus) Bartsch, 1927: Proc. USNM 70(2660):28-29.
PARATYPE: LACM 2183 (9 dry), Bahía Magdalena, Pacific Coast Baja California Sur, México
Leg.: C.L. Simons.
Remarks: Ex UCLA 46030.
trosti, Amphithalamus (Amphithalamus) Strong & Hertlein, 1939: Allan Hancock Pac. Exped.
2(12):228-229.
PARATYPES: LACM 1730 (2 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
zeteki, Barleeia Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):228.
PARATYPES: LACM 1716 (2 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
Remarks: Junior synonym of Barleeia paupercula (C.B. Adams, 1856).
Family Iravadiidae Thiele, 1928
lignicola, Nozeba Hasegawa, 1997: Nat. Sci. Mus. Tokyo Mem. 12:108-110.
PARATYPES: LACM 2844 (2 dry), 140-400 m, 3.5 km ENE of Osezaki, Suruga-wan, Shizuoka
Pref., Honshu, Japan (35 00.5 N, 138 45.1 E), Leg.: R/V Soro-Maru, 26 OCT 1994.
Family Hydrobiidae Stimpson, 1865
adamantina, Tryonia (Paupertryonia) Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull.
116:41-42, fig. 20.
HOLOTYPE: LACM 2089 (wet), Diamond Y Spring, Pecos Co., Texas, Leg.: D.W. Taylor, 26
JUN 1968.
alamosae, Tryonia (Paupertryonia) Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull.
116:42-44, fig. 21.
HOLOTYPE: LACM 2188 (wet), Ojo Caliente, Socorro Co., New Mexico, Leg.: D.W. Taylor,
13 APR 1979.
alcaldei, Nanivitrea Jaume & Abbott, 1948: Revista Sociedad Malac. 6(1):5-6.
PARATYPES: LACM 1609 (12 dry), Arroyo del Naranjo, Trinidad, Cienfuegos Prov., Cuba.
arizonae, Apachecoccus Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull. 116:32-34, fig.
15.
HOLOTYPE: LACM 2203 (wet), unnamed spring on N side of Gila River, ca. 2 mi. N of Bylas,
Graham Co., Arizona, Leg.: D.W. Taylor, 20 APR 1971.
Remarks: Currently in the genus Pyrgulopsis.
bernardinus, Yaquicoccus Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull. 116:34-35,
fig. 16.
HOLOTYPE: LACM 2186 (wet), Cochise Co., Arizona, Leg.: D.W. Taylor, 27 JUL 1968
Remarks: Currently in the genus Pyrgulopsis.
brandi, Amnicola Drake, 1953: Jour. Washington Acad. Sci. 43(1):26-27.
PARATYPES: LACM 2231 (9 dry), Las Palomas, Distrito Galeana, Chihuahua, México, Leg.:
R.J. Drake & C.C. Hoff, 15 APR 1949.
brunei, Tryonia (Paupertryonia) Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull.
116:44-45, fig. 22.
HOLOTYPE: LACM 2251 (wet), outflow of Phantom Lake Spring, at Joe Kingston Ranch, Jeff
Davis Co., Texas, Leg.: D.W. Taylor, 5 NOV 1981.
chupaderae, Fontelicella Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull. 116:24-26, fig.
11.
HOLOTYPE: LACM 2218 (wet), S end of Chupadera Mountains., 5 mi. W of Bosc del Apache
National Wildlife Refuge, Socorro Co., New Mexico, Leg.: D.W. Taylor, 23 APR 1979.
Remarks: Currently in the genus Pyrgulopsis.
davisi, Fontelicella Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull. 116:10-12, fig. 4.
HOLOTYPE: LACM 2211 (wet), tributary of Limpia Creek, ca. 5 mi. NE of Fort Davis, Jeff
Davis Co., Texas, Leg.: D.W. Taylor, 16 APR 1981.
Remarks: Currently in the genus Pyrgulopsis.
gilae, Fontelicella Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull. 116:16-18, fig. 7.
HOLOTYPE: LACM 2214 (wet), springs on N side of East Fork of Gila River, Grant Co., New
Mexico.
Remarks: Currently in the genus Pyrgulopsis.
gilae, Tryonia Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull. 116:36-37, fig. 17.
HOLOTYPE: LACM 2187 (wet), on N side of Gila River, ca. 2 mi. N of Bylas, Graham Co.,
Arizona, Leg.: D.W. Taylor, 20 APR 1971.
hertleini, Lyrodes Drake, 1956: Bull. So. Calif. Acad. Sci. 55(1):44-46.
PARATYPES: LACM 1657 (2 dry), Las Palomas, Chihuahua, México, Leg.: R.J. Drake, 14
APR 1949.
kolobensis, Fontelicella Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull. 116:19-20, fig.
8.
HOLOTYPE: LACM 2216 (wet), Toquerville Springs, Washington Co., Utah, Leg.: D.W.
Taylor, 13 NOV 1984.
Remarks: Currently in the genus Pyrgulopsis.
kosteri, Tryonia (Paupertryonia) Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull.
116:45-47, fig. 23.
HOLOTYPE: LACM 2252 (wet), Sago Spring, Chaves Co., New Mexico, Leg.: D.W. Taylor, 29
OCT 1981.
letsoni, Amnicola Walker, 1901: The Nautilus 14(10):113-114.
SYNTYPES: LACM 2023 (3 dry), Goat Id., Niagara River, Niagara Co., New York, Leg.: E. J.
Letson.
metcalfi, Fontelicella Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull. 116:12-14, fig. 5.
HOLOTYPE: LACM 2212 (wet), Naegele Springs, 5.3 mi. NNW of Ruidosa, Presidio Co.,
Texas, Leg.: D.W. Taylor, 15 APR 1981.
Remarks: Currently in the genus Pyrgulopsis.
pecosensis, Fontelicella Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull. 116:27-28, fig.
12.
HOLOTYPE: LACM 2220 (wet), Blue Spring, Eddy Co., New Mexico, Leg.: D.W. Taylor, 20
APR 1981.
Remarks: Currently in the genus Pyrgulopsis.
pinetorum, Fontelicella Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull. 116:20-21, fig.
9.
HOLOTYPE: LACM 2217 (wet), spring tributary to Leeds Creek, Washington Co., Utah, Leg.:
D.W. Taylor, 13 NOV 1984.
Remarks: Currently in the genus Pyrgulopsis.
roswellensis, Fontelicella Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull. 116:14-16, fig.
6.
HOLOTYPE: LACM 2213 (wet), seepage, Chaves Co., New Mexico, Leg.: D.W. Taylor.
Remarks: Currently in the genus Pyrgulopsis.
stocktonensis, Tryonia Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull. 116:37-38, fig.
18.
HOLOTYPE: LACM 2090 (wet), Diamond Y Draw, 9 mi. N of Fort Stockton, 0.5 mi. W of
State Highway. 18, Pecos Co., Texas, Leg.: D.W. Taylor, 27 OCT 1984.
thermalis, Fontelicella Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull. 116:28-30, fig.
13.
HOLOTYPE: LACM 2224 (wet), E side of Gila River, Grant Co., New Mexico, Leg.: D.W.
Taylor, 10 APR 1981.
Remarks: Currently in the genus Pyrgulopsis.
trivialis, Fontelicella Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull. 116:30-32, fig. 14.
HOLOTYPE: LACM 2225 (wet), Apache Co., Arizona, Leg.: D.W. Taylor, 5 NOV 1980.
Remarks: Currently in the genus Pyrgulopsis.
Family Elachisinidae Ponder, 1985
floridanus, Microdonchus Rehder, 1943: Proc. USNM 93(3161):193-194.
PARATYPES: LACM 1317 (11 dry); PARATYPES: LACM 2059 (7 dry), Missouri Key,
Monroe Co., Florida, Leg.: B.R. Bales.
Remarks: Currently in the genus Elachisina.
hertleini, Rissoella Smith & Gordon, 1948: Proc. Calif. Acad. Sci., 4th ser., 26(8):224-225.
PARATYPE: LACM 1003 (1 dry),18 m, off Cabrillo Pt., Monterey Bay, Monterey Co.,
California, Leg.: A.G. Smith, JUL 1913.
Remarks: Currently in the genus Elachisina.
Family Assimineidae H. Adams & A. Adams, 1856
pecos, Assiminea Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull. 116:8-9, fig. 2.
HOLOTYPE: LACM 2088 (dry), seepage on Bitter Lake National Wildlife Refuge, Chaves Co.,
New Mexico, Leg.: D.W. Taylor, 5 MAY 1981.
takanoshimana, Omphalotropis japonica Pilsbry, 1924: Proc. Acad. Nat. Sci. Philadelphia
76:13.
PARATYPES: LACM 2020 (6 dry), Takanoshima, Japan
Family Pomatiopsidae Stimpson, 1865
hinkleyi, Pomatiopsis Pilsbry, 1896b: The Nautilus 10(4):37-38.
PARATYPES: LACM 2055 (16 dry), Black Falls, above Florence, Lauderdale Co., Alabama,
Leg.: A.A. Hinkley, 1894.
Remarks: Junior synonym of Pomatiopsis lapidaria (Say, 1817).
Family Tornidae Sacco, 1896
bakeri, Circulus Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):240-241.
PARATYPE: LACM 1740 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
Remarks: Currently in the genus Solariorbis.
bartschi, Cyclostrema Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):240.
PARATYPE: LACM 1739 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá
Leg.: L.G. Hertlein.
Remarks: Currently in the genus Cyclostremiscus. Junior synonym of S. veleronis (Strong
& Hertlein, 1947).
chamizal, Cyclostremiscus Myers, Hertz, & Gemmell, 1991: Venus 50(1):31-33.
PARATYPE: LACM 2368 (1 dry), 18-27 m, N side Bahía de las Banderas, 2-5 mi. SE Punta de
Mita & off La Cruz, Jalisco, México (20 45 N, 105 30 W), Leg.: J.H. McLean, C. Miller, A.M.
Keen, R/V Gringa, 20-24 MAR 1965.
cornis, Cochliolepis Hertz, Myers, & Gemmell, 1992: The Veliger 35(1):70-71.
PARATYPES: LACM 2576 (6 dry), Bahía San Felipe, Golfo de California, Baja California,
México (31 03 N, 114 48 W), Leg.: J. Gemmell, 1965-1976.
cortezi, Solariorbis Myers, Hertz, & Gemmell, 1991: Venus 50(1):33-34.
PARATYPE: LACM 2369 (1 dry), Low intertidal, Bahía Concepción, Golfo de California, Baja
California Sur, México (26 42 N, 111 55 W), Leg.: R. Maynard, JUN 1963
eshnauri, Vitrinella (Vitrinellops) Bartsch, 1907: Proc. USNM 32(1520):168.
PARATYPES: LACM 2165 (5 dry), Off Terminal Id., Los Angeles Co., California, Leg.: W.H.
Eshnaur.
Remarks: Ex UCLA 20969.
hannai, Delphinoidea Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):242-243.
PARATYPE: LACM 1741 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
Remarks: Currently in the genus Solariorbis.
hemphilli, Teinostoma (?Teinostoma) Strong & Hertlein, 1939: Allan Hancock Pac. Exped.
2(12):244.
PARATYPE: LACM 1742 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
mccullochae, Cyclostrema Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):239.
PARATYPES: LACM 1738 (2 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
Remarks: Currently in the genus Solariorbis. Junior synonym of Solariorbis concinnus (C.B.
Adams, 1852).
ochsneri, Teinostoma (Pseudorotella) Strong & Hertlein, 1939: Allan Hancock Pac. Exped.
2(12):244-245.
PARATYPE: LACM 1743 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
salvatierrensis, Cyclostremiscus Hertz, Myers, & Gemmell, 1992: The Veliger 35(1):71-72.
PARATYPES: LACM 2577 (2 dry), Bahía Cuastecomate, 3 air mi. NW of Barra de Navidad,
Jalisco, México (65 16.8 N, 104 44.88 W, Leg.: J.H. McLean, P.M. Oringer, & G. Radwin, 21
OCT 1968; PARATYPE: LACM 2578 (1 dry), Off anchorage at Bahía de los Muertos, Golfo de
California, Baja California Sur, México (23 58 N, 109 46 W), Leg.: J.H. McLean, P.M. Oringer,
& L. Marincovich, 8 APR 1966; PARATYPE: LACM 2579 (1 dry), N side Frailes, Bahía Pulmo,
Golfo de California, Baja California Sur, México (23 24 N, 109 25.5 W), Leg.: J.H. McLean &
P.M. Oringer, 6-7 APR 1966; PARATYPE: LACM 2580 (1 dry), Under boat anchorage at S end
of Cabo Pulmo, N side Frailes, Golfo de California Sur, México (23 24 N, 109 25.5 W), Leg.:
J.H. McLean, P.M. Oringer, & L. Marincovich, 6-7 APR 1966; PARATYPE: LACM 2581 (1
dry), 44 m, off Isla Cresciente, Bahía Magdalena, Pacific Coast Baja California Sur, México
(24 19.3 N, 111 37.5 W), Leg.: R/V Velero III, 17 FEB 1936 ; PARATYPE: LACM 2582 (1
dry), Bahía Cocos, S of Puerto Culebra, Guanacaste Prov., Costa Rica, Leg.: R/V Velero III, 13
MAR 1933; PARATYPES: LACM 2583 (3 dry), Off Punta Arena, Golfo de California, Baja
California Sur, México (23 32 N, 109 28 N), Leg.: J.H. McLean, P.M. Oringer, & L.
Marincovich, 7 APR 1966; PARATYPES: LACM 2584 (5 dry), Off the house and hotel at Los
Tezos Ranch, 1.5 mi. S of Punta Arena, Golfo de California, Baja California Sur, México
(23 31 N, 109 29 W), Leg.: J.H. McLean, 2 FEB 1971; PARATYPE: LACM 2585 (1 dry),
Bahía Partida, between Isla Partida & Isla Espíritu Santo, Golfo de California, Baja California
Sur, México (24 31 N, 110 23 W), Leg.: J.H. McLean et al., 10 APR 1966.
Family Caecidae Gray, 1850
bahiahondaense, Caecum Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):219, 224.
PARATYPES: LACM 1712 (2 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá
Leg.: L.G. Hertlein.
boucheti, Meioceras Pizzini & Raines, 2011: Boll. Malacolog. 47:42-43.
PARATYPE: LACM 3133 (dry), 18-26 m, W side Arutanga, Aitutaki, Lower Cook Ids., Cook
Ids. (18°52.30’N, 154°50’W).
bounty, Caecum Pizzini & Raines, 2011: Boll. Malacolog. 47:31-33.
PARATYPE: LACM 3132 (dry), 1 m, Ahe Atoll, Tuamotu Arch. French Polynesia (14°28 N,
146°22’E).
brennani, Caecum Raines & Pizzini, 2009: Boll. Malacolog. 45:117, 120, figs. 3A-D, 4B, 5B,
6C-D.
HOLOTYPE: LACM 3103 (dry); PARATYPES: LACM 3104-3105 (3 dry), 18.2-28.9 m, Marpi,
Saipan, No. Mariana Ids. (15°17’30 N, 145°48’15 E), Leg.: B.K. Raines.
campanulatum, Caecum Raines & Pizzini, 2005: Iberus 12(1):62-63, fig. 12.
HOLOTYPE: LACM 3027 (dry); PARATYPES: LACM 3028 (2 dry), 20 m, off Hanga Nui, Isla
de Pascua (= Easter Id.) (27°7.46’S, 109°16.36’W), Leg.: B.K. Raines, DEC 2000.
cooki, Caecum Pizzina & Raines, 2011: Bol. Malacolog. 47:29-31.
PARATYPE: LACM 3130 (1 dry), 5-20 m, of Ajer & Meno islets, NW side of Lombok, Nusa
Tenggara Barat, Indonesia (28°22’S, 116°4’E); PARATYPES: LACM 3131 (4 dry), near Club
Mediterranean, E side of Moorea, Society Arch., French Polynesia (17°30’S, 149°46’W).
danielei, Caecum Pizzini & Raines, 2011: Boll. Malacolog. 4725-26.
PARATYPE: LACM 3129 (1 dry), 34 m, 1 mi. W of Onna Village, Okinawa-jima, Okinawashoto, Ryukyu-shoto, Okinawa Pref., Japan (26°29.7’N, 127°50.60’E).
heterochromum, Caecum Raines & Pizzini, 2005: Iberus 23(1):57-59, figs. 7-8.
HOLOTYPE: LACM 3021 (dry); PARATYPES: LACM 3022 (6 dry), 20 m, off Hanga Nui, Isla
de Pascua (= Easter Id.) (27°7.46’S, 109°16.35’W), Leg.: B.K. Raines, DEC 2000.
limnetes, Caecum (Micranellum) Long, 1972: The Veliger 14(3):291.
PARATYPE: LACM 1446 (1 dry), Estero Cholla, Sonora, México (31 20.52 N, 113 36.5 W)
Leg.: Garrell Long, 23 OCT 1970.
lohri, Micranellum Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):225-226.
PARATYPE: LACM 1714 (1 dry), 5-16 m, off Isla Taboga, Panamá Prov., Panamá, Leg.: L.G.
Hertlein.
Remarks: Curently in the genus Caecum.
massambabensis, Caecum (Elephantulum) Absalão, 1994: Jour. Conch. 35(2):138-139.
PARATYPE: LACM 2431 (1 dry), 31 m, off Massambaba Beach, Rio de Janeiro State, Brazil
(22 57 30 S, 42 19 30 W), Leg.: Brazilian Naval R/V Suboficial Oliveira, 25 MAR 1986.
moorei, Fartulum Marincovich, 1973: Nat. Hist. Mus. L.A. Co. Sci. Bull. 16:30-31, fig. 64.
HOLOTYPE: LACM 1591 (dry); PARATYPES: LACM 1592 (1050 dry), 0 m, intertidal,
Iquique, Tarapaca Prov., Chile (20 13 S, 70 10 W), Leg.: L. Marincovich, AUG 1964.
pascuanum, Caecum Raines & Pizzini, 2005: Iberus 23(1):60-61, fig. 9.
HOLOTYPE: LACM 3023 (dry); PARATYPE: LACM 3024 (1 dry), 15 m, in cave off HangaTeo, Isla de Pascua (= Easter Id.) (27°3.37'S, 109°21.58'W), Leg.: B.K. Raines, DEC 2000.
rapanuiense, Caecum Raines & Pizzini, 2005: Iberus 23(1):61-62, fig. 10.
HOLOTYPE: LACM 3025 (dry); PARATYPES: LACM 3026 (2 dry), 30 m, off Tahai, Isla de
Pascua (= Easter Id.) (27°7.20'S, 109°26.30'W), Leg.: B.K. Raines, DEC 2000.
rehderi, Caecum Raines & Pizzini, 2005: Iberus 23(1):52-55, fig. 2.
HOLOTYPE: LACM 3019 (dry); PARATYPE: LACM 3020 (1 dry), 10-20 m, Punta Rosalia, E
of Anakena, Isla de Pascua (= Easter Id.) (27°4.18'S, 109°19.45'W), Leg.: B.K. Raines, APR
1988.
richthofeni, Caecum Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):224-225.
PARATYPE: LACM 1713 (1 dry), 5-16 m, off Isla Taboga, Panamá Prov., Panamá, Leg.: L.G.
Hertlein.
skoglundae, Caecum Pizzini, Raines, & Nofroni, 2007: Iberus 25(2):2-6, figs. 1A-F.
HOLOTYPE LACM 3063 (dry); PARATYPES LACM 3064(3 dry), Punta Chame, Bahía de
Panamá, Panamá Prov., Panamá 8º41'N, 79º39'W), Leg.: D.R. Shasky, JUN 1977.
tahitianum, Caecum Pizini & Raines, 2011: Boll. Malacolog. 47:27-28.
PARATYPE: LACM 3134 (1 dry), Tiarei, Tahiti, Society Archipelago, French Polynesia, Leg.:
J. Letournex.
Superfamily STROMBOIDEA Rafinesque, 1815
Family Strombidae Rafinesque, 1815
nanus, Strombus (Tricornis) raninus Bales, 1942: The Nautilus 56(1):19.
PARATYPE: LACM 1358 (1 dry), Lake Worth, Palm Beach Co., Florida
Remarks: Junior synonym of Strombus raninus Gmelin, 1791.
Superfamily VANIKOROIDEA Gray, 1840
Family Vanikoridae Gray, 1840
schmiederi, Megalomphalus McLean, 1995: Tax. Atlas Benthic Fauna Santa Maria Basin & W.
Santa Barbara Ch. 9(2)(1):51, fig. 1.9B.
HOLOTYPE: LACM 2777 (dry); PARATYPES: LACM 2778 (56 dry), 37-55 m, unnamed bank
WSW of Pt. Sur, Monterey Co., California (approx. 36 17 N, 21 58 W), Leg.: R.W. Schmieder,
R/V Cordell Explorer, SEP 1988-OCT 1989.
Family Haloceratidae Warén & Bouchet, 1991
rugosa, Haloceras Warén & Bouchet, 1991: Mem. Mus. Natn. Hist. Nat., Paris (A)150:145-147,
figs. 72-73, 91-92.
HOLOTYPE: LACM 2279 (dry), 457 m, 8 mi. NW of Isla Santa Maria (= Charles Id.), Islas
Galápagos (1 09 S, 90 36 W), Leg.: R/V Velero III, 23 JAN 1938.
Superfamily CALYPTREAOIDEA Lamarck, 1809
Family Calyptraeidae Lamarck, 1809
coei, Crepidula Berry, 1950: Leaflets in Malacology 1(8):35-39.
PARATYPE: LACM 1008 (1 dry), Near Sunset Beach, Orange Co., California, Leg.: E.P. Chace.
naticarum, Crepidula rugosa Williamson, 1905a: The Nautilus 19(5):50.
SYNTYPES: LACM 1026 (2 dry), San Pedro, Los Angeles Co., California.
Remarks: Junior synonym of Crepidula onyx Sowerby, 1824.
norrisiarum, Crepidula rugosa Williamson, 1905a: The Nautilus 19(5):50-51.
SYNTYPES: LACM 1027 (4 dry), San Pedro, Los Angeles Co., California.
Remarks: Junior synonym of Crepidula onyx Sowerby, 1824.
Superfamily CAPULOIDEA Fleming, 1822
Family Capulidae Fleming, 1822
octocarinatus, Trichosirius Powell, 1931: Rec. Canterbury Mus. 3(6):376.
PARATYPE: LACM 1359 (1 dry), 91 m, Bounty Ids., New Zealand.
Remarks: Currently in the genus Trichotropis.
Superfamily XENOPHOROIDEA Troschel, 1852
Family Xenophoridae Troschel, 1852
tulearensis, Xenophora Stewart & Kosuge, 1993: Bull. Inst. Malac., Tokyo 3(1):4-5.
PARATYPE: LACM 2716, Tulear, Toliara Prov., Madagascar.
Superfamily CYPRAEOIDEA Rafinesque, 1815
Family Cypraeidae Rafinesque, 1815
dayritiana, Cypraea Cate, 1963: The Veliger 5(4):141-142.
PARATYPES: LACM 2025 (2 dry), Demang Id., Calamian Group, Palawan Prov., Philippines
Leg.: J. Norton, 26 MAY 1962.
Remarks: Currently in the genus Blasicrura fide Lorenz & Hubert (2000).
eugeniae, Notadusta Cate, 1975: The Veliger 17(3):260, fig. 5.
HOLOTYPE: LACM 1704 (dry), Hopetoun, Western Australia (38 07 S, 144 27 E).
Remarks: Junior synonym of Erronea xanthodon (Sowerby, 1832) fide Lorenz & Hubert
(2000). Incorrect coordinates cited by Cate, correct coordinates = ca. 33 57 S, 120 7 E.
garciai, Cribrarula Lorenz & Raines, 2001: La Conchiglia 33(299):27-29, pl. 1.
HOLOTYPE: LACM 2898 (dry), 30 m, cave roof, near Hanga Roa, Hanga Roa Bay, Isla de
Pascua (= Easter Id.) (ca. 27º8.15’S, 109º26.45’W), M. Garcia, SUM 200.
greegori, Cypraea cruenta Ford, 1893: The Nautilus 6(10):112-113.
PARATYPE: LACM 2397 (1 dry),"Ceylon" [= Sri Lanka].
Remarks: Junior synonym of Ovatipsa coloba (Melvill, 1888) fide Lorenz & Hubert (2000).
maricola , Pustularia Cate, 1976a: The Veliger 18(4):383, fig. 3.
HOLOTYPE: LACM 1706 (dry), Santa Cruz, Marinduque Prov., Philippines (13 28 N,
122 01 E), Leg.: R.M. Lumawig, 28 FEB 1964.
Remarks: Junior synonym of Nesiocypraea lisetae (Kilburn, 1975) fide Lorenz & Hubert
(2000).
perlae, Zoila Lopez & Chiang, 1975: The Veliger 18(1):84, figs. 1a-c, 2b.
HOLOTYPE: LACM 1558 (dry),182-274 m, 18 deg. to 25 deg. S paralleling the off-shore coast
of North West Cape, Western Australia (21 00 S, 114 10 E).
Remarks: Erroneously published as LACM 1706.
piscatorum, Erronea (Adusta) Schilder, 1965: Arch. Moll. 94(3/4):148-149, fig. 2.
PARATYPE: LACM 2081 (1 dry), Kii-suido, Wakayama Pref., Japan, NOV 1963.
Remarks: Junior synonym of Erronea subviridis (Reeve, 1835) fide Lorenz & Hubert (2000).
takahashii, Pustularia cicercula Moretzsohn, 2007: Viasaya 2(1):44-46.
PARATYPES: LACM 3051 (4 dry), Haleiwa, Waialua Dist., O‘ahu, Hawai‘i, Hawaiian Ids.
Family Ovulidae Fleming, 1822
aboriginea, Margovula Cate, 1973: The Veliger, v. 15 supplement, p. 20.
PARATYPE: LACM 1893 (1 dry), Geralia [= Giralia], Exmouth Gulf, Western Australia
(22 00 S, 114 15 E). , Leg.: L. Figges.
Remarks: Currently in the genus Diminovula fide Lorenz & Fehse (2009). Incorrect latitude
cited by Cate, correct latitude = 22 19 S.
advena, Spiculata Cate, 1978a: The Nautilus 92(4):164, fig. 6.
HOLOTYPE: LACM 1794 (dry), Off Sand Key, 7 mi. SW of Key West, Monroe Co., Florida
(24 33 N, 81 47 W).
albomarginata, Pseudosimnia (Pseudosimnia) Cate, 1978b: Venus 37(4):195-196, pl. 2, fig. 5.
HOLOTYPE: LACM 1841 (dry), Malaita Id., Solomon Ids. (9 00 S, 161 00 E).
Remarks: Junior synonym of Pseudosimnia pyrulina (A. Adams, 1854) fide Lorenz & Fehse
(2009).
alleneae, Cyphoma Cate, 1973: The Veliger, v. 15 supplement, p. 67-68, fig. 151, pl. 4, fig.
151c.
HOLOTYPE: LACM 1292 (dry), Missouri Key, Monroe Co., Florida (24 40 N, 81 10 W)
7 JUL 1957.
allynsmithi, Aperiovula Cate, 1978b: Venus 37(4):197-198, pl. 2, fig. 7.
HOLOTYPE: LACM 1834 (dry) 80 m, off Wakayama, Kii-suido, Wakayama Pref., Honshu,
Japan (31 10 N, 135 15 E).
Remarks: Currently in the genus Cuspivolva fide Fehse (2001). Incorrect coordinates cited by
Cate, correct latitude = ca. 34 10 N, 135 05 E.
aurantiomaculata, Pseudosimnia (Diminovula) Cate & Azuma in Cate, 1973: The Veliger, v.
15 supplement, p. 27.
PARATYPE: LACM 2228 (1 dry), 73 m, off Hinomi-saki, Kii-hanto, Wakayama Pref., Honshu,
Japan, Leg.: Shiago Habo, 17 FEB 1973.
Remarks: Currently in the genus Diminovula fide Fehse (2001).
azumai, Primovula (Primovula) Cate, 1970: The Veliger 13(2):181.
PARATYPE: LACM 1872 (1 dry), 30-40 m, 2-3 km off Kirime-zaki, Kii-suido, Wakayama
Pref., Honshu, Japan, Leg.: Eizo Azuma, 3 JAN 1971
Remarks: Currently in the genus Dentiovula fide Lorenz & Fehse (2009).
basilia, Galeravolva Cate, 1978b: Venus 37(4):191-192, pl. 1, fig. 1.
HOLOTYPE: LACM 1829 (dry), 80 m, off Wakayama, Kii-suido, Wakayama Pref., Honshu,
Japan (31 10 N, 135 15 E).
Remarks: Incorrect latitude cited by Cate, correct coordinates = ca. 34 10 N, 135 05 E.
Currently in the genus Habuprionovolva fide Lorenz & Fehse (2009).
bellica, Primovula (Primovula) Cate, 1973: The Veliger, v. 15 supplement, p. 41, fig. 83.
HOLOTYPE: LACM 1189 (dry), 4 m, Bataan Peninsula, Manilla Bay, Bataan Prov., Luzon Id.,
Philippines (14 40 N, 120 25 E), 31 DEC 1964; PARATYPE: LACM 1863 (1 dry), Palawan
Prov., Philippines, 4 MAR 1969.
Remarks: Currently in the genus Cuspivolva fide Lorenz & Fehse (2009). Incorrect longitude
cited by Cate, correct longitude = ca. 120 35 E.
bijuri, Spiculata Cate, 1976a: The Veliger 18(4):384, fig. 1.
HOLOTYPE: LACM 1708 (dry), 48 m, SE of Key West, Monroe Co., Florida (24 33 N,
81 47 W), JAN 1970.
Remarks: Currently in the genus Neosimnia fide Lorenz & Fehse (2009).
bratcherae, Cymbula Cate, 1973: The Veliger, v. 15 supplement, p. 81-82, fig. 180.
HOLOTYPE: LACM 1610 (dry), Pulmo Reef, Golfo de California, Baja California Sur, México
(23 12 N, 109 32 W), Leg.: Twila Bratcher, APR 1961.
Remarks: Junior synonym of Simnialena rufa (Sowerby, 1832) fide Lorenz & Fehse (2009).
cardini, Dentiovula Cate, 1978b: Venus 37(4):193-194 pl. 1, fig. 3.
HOLOTYPE: LACM 1832 (dry), 80 m, off Wakayama, Kii-suido, Wakayama Pref., Honshu,
Japan (31 10 N, 135 15 E).
Remarks: Incorrect coordinates cited by Cate, correct coordinates = ca. 34 10 N, 135 05 E.
Junior synonym of Dentiovula colobica (Azuma & Cate, 1971) fide Lorenz & Fehse (2009).
castanea, Prionovolva Cate, 1978a: The Nautilus 92(4):160-161, fig. 1.
HOLOTYPE: LACM 1789 (dry), Gulf of Oran, Ouahran Dept., Algeria (35 45 N, 0 38 W).
conspicua, Crenavolva (Crenavolva) Cate, 1975: The Veliger 17(3):258, fig. 9.
HOLOTYPE: LACM 1705 (dry), 91 m, off Nada, Kii-suido, Wakayama Pref., Honshu, Japan
(34 00 N, 134 48 E).
Remarks: Junior synonym of Rotaovula septemacula (Azuma, 1974) fide Lorenz & Fehse
(2009).
coroniola, Pseudosimnia (Inflatovula) Cate, 1973: The Veliger, v. 15 supplement, p. 31.
PARATYPE: LACM 1867 (1 dry), 64 m, off Tonda, Wakayama Pref., Honshu, Japan, Leg.: S.
Habei, OCT 1971.
Remarks: Currently in the genus Diminovula fide Lorenz & Fehse (2009).
culmen, Pseudosimnia (Inflatovula) Cate, 1973: The Veliger, v. 15 supplement, p. 33.
PARATYPE: LACM 1866 (1 dry), Kii-suido, off Kii-hanto, Wakayama Pref., Honshu, Japan
(34 00 N, 134 48 E),13 MAY 1972.
Remarks: Currently in the genus Diminovula fide Lorenz & Fehse (2009).
diadema, Dentiovula Cate, 1978b: Venus 37(4):194-195, pl. 1, fig. 4.
HOLOTYPE: LACM 1835 (dry), 80 m, off Wakayama, Kii-suido, Wakayama Pref., Honshu,
Japan (31 10 N, 135 15 E)
Remarks: Incorrect coordinates cited by Cate, correct coordinates = ca. 34 10 N, 135 05 E.
Junior synonym of Dentiovula colobica(Azuma & Cate, 1971) fide Lorenz & Fehse (2009).
dolabra, Delonovolva Cate, 1974: The Veliger 16(4):383-384, fig. 5.
HOLOTYPE: LACM 1557 (dry), 91 m, off Hinomi-saki, Kii-suido, Wakayama Pref., Honshu,
Japan (34 00 N, 134 48 E), Leg.: S. Habu, MAR 1971
Remarks: Currently in the genus Dissona fide Fehse (2001).
eizoi, Dentiovula Cate & Azuma in Cate, 1973: The Veliger, v. 15 supplement, p. 16.
PARATYPE: LACM 1870 (1 dry), 55-91 m, 2-3 km off Kirime-zaki, Kii-suido, Wakayama
Pref., Honshu, Japan (34 00 N, 134 48 E), Leg.: E. Azuma, 3 JAN 1971.
emersoni, Aperiovula Cate, 1973: The Veliger, v. 15 supplement, p. 39-40.
PARATYPES: LACM 1890 (2 dry), Melilla, Spain (on Moroccan mainland), 5 NOV 1958.
Remarks: Junior synonym of Pseudosimnia adriatica (Sowerby, 1828) fide Lorenz & Fehse
(2009).
figgisae, Xandarovula Cate, 1973: The Veliger, v. 15 supplement, p. 36.
PARATYPE: LACM 1869 (1 dry), Geralia [= Giralia], Exmouth Gulf, North West Cape,
Western Australia,1967.
Remarks:Junior synonym of Ovula ovum (Linné, 1758) fide Lorenz & Fehse (2009).
gibbulum, Pseudocyphoma Cate, 1978a: The Nautilus 92(4):165-166, fig. 9.
HOLOTYPE: LACM 1793 (dry), 18 m, off the Dry Tortugas Ids., Monroe Co., Florida (24 40 N,
82 55 W).
Remarks: Junior synonym of Neosimnia arcuata (Reeve, 1865) fide Lorenz & Fehse (2009).
greenbergae, Phenacovolva (Turbovula) Cate, 1974: The Veliger 16(4):382-383, fig. 3.
HOLOTYPE: LACM 1556 (dry), 80 m, off Kao-hsiung, Kao Hsiung Co., Taiwan (22 44 N,
120 21 E),13 SEP 1973.
Remarks: Junior synonym of Quasisimnia hirasei (Pilsbry, 1913) fide Lorenz & Fehse (2009).
howlandae, Primovula (Primovula) Cate, 1974: The Veliger 16(4):381-382.
PARATYPE: LACM 1864 (1 dry), Clairview (= MacKay), Queensland, Australia (22 07 S,
149 32 E), 8 AUG 1973.
Remarks: Incorrect coordinates cited by Cate, correct coordinates = 21 06 S, 149 10 W.
Currently in the genus Cuspivolva fide Lorenz & Fehse (2009).
iberia, Aperiovula adriatica Cate, 1973: The Veliger, v. 15 supplement, p. 37, fig. 72.
HOLOTYPE: LACM 1607 (dry), Golfo de Cadiz, Cadiz Prov., Spain (36 33 N, 6 12 W);
PARATYPE: LACM 1897 (1 dry), Neapolis, Peloponnessos, Greece, 1965.
Remarks: Junior synonym of Pseudosimnia adriatica (Sowerby, 1828) fide Lorenz & Fehse
(2009).
imitabilis, Crenavolva (Serratovolva) Cate, 1973: The Veliger, v. 15 supplement, p. 54, fig. 112.
HOLOTYPE: LACM 1608 (dry), Ogokuda Beach, Shio-no-misaki, Kii-suido, Wakayama Pref.,
Honshu, Japan (34 00 N, 134 48 E).
Remarks: Junior synonym of Serratovolva dondani (Cate, 1964) fide Lorenz & Fehse (2009).
jeanae, Aperiovula Cate, 1973: The Veliger, v. 15 supplement, p. 37, figs. 73, 73c.
HOLOTYPE: LACM 1187 (dry), Enshu-nada, between Izu-hanto & Ise-wan, Honshu, Japan
(34 00 N, 139 30 E),10 DEC 1969; PARATYPE: LACM 1898 (1 dry),146-183 m, off Tosa,
Tosa-wan, Kochi Pref., Shikoku, Japan, 19 JUL 1970.
Remarks: Currently in the genus Pseudosimnia fide Fehse (2001).
kosugei, Margovula Cate, 1973: The Veliger, v. 15 supplement, p. 17-18.
PARATYPE: LACM 1892 (1 dry), 65-67 m, off Joga-shima, Sagami-wan, Chiba Pref., Honshu,
Japan, 13 MAY 1973.
Remarks: Currently in the genus Diminovula fide Lorenz & Fehse (2009).
kurziana, Cymbovula Cate, 1976b: The Veliger 19(2):161-162, fig. 3.
HOLOTYPE: LACM 1764 (dry), Malaita Id., Solomon Ids. (9 00 S, 161 00 E).
Remarks: Currently in the genus Naviculavolva fide Lorenz & Fehse (2009).
macleani, Delonovolva Cate, 1976b: The Veliger 19(2):159, fig. 5.
HOLOTYPE: LACM 1772 (dry), 54-137 m, small islets off Punta Quepos, Puntarenas Prov.,
Costa Rica (9 22.72 N, 84 09.68 W), Leg.: J.H. McLean, R/V Searcher, 11-13 MAR 1972.
Remarks: Currently in the genus Neosimnia fide Fehse (2001).
malaita, Cymbovula Cate, 1976b: The Veliger 19(2):161, fig. 2.
HOLOTYPE: LACM 1771 (dry); PARATYPE: LACM 1874 (1 dry), Malaita Id., Solomon Ids.
(9 00 S, 161 00 E).
Remarks: Currently in the genus Naviculavolva fide Lorenz & Fehse (2009).
marferula, Simnialena Cate, 1973: The Veliger, v. 15 supplement, p. 75, fig. 165.
HOLOTYPE: LACM 1293 (dry), Gulf Beach, Port Aransas, Mustang Id., Nueces Co., Texas
(24 45 N, 97 50 W).
Remarks: Incorrect coordinates cited by Cate, correct coordinates = ca. 27 45 N, 97 05 W.
masaoi, Dentiovula Cate, 1973: The Veliger, v. 15 supplement, p. 15.
PARATYPE: LACM 1871 (1 dry), 73-91 m, 2-3 km off Kirime-zaki, Kii-suido, Wakayama
Pref., Honshu, Japan (34 00 N, 134 48 E).
meyeriana, Aperiovula Cate, 1973: The Veliger, v. 15 supplement, p. 38, fig. 75.
HOLOTYPE: LACM 1186 (dry); PARATYPE: LACM 1891 (1 dry), Mikawa-wan, Aichi Pref.,
Honshu, Japan (34 40 N, 137 00 E), 8 JAN 1969.
Remarks: Currently in the genus Sandalia fide Lorenz & Fehse (2009).
nielseni, Pseudosimnia (Diminovula) Cate, 1976a: The Veliger 18(4):383-384, fig. 2.
HOLOTYPE: LACM 1707 (dry), 8 m, Keppel Bay, Queensland, Australia (23 10 S, 150 49 E),
Leg.: T. Nielsen, 20 MAY 1975.
Remarks: Currently in the genus Diminovula fide Fehse (2001).
nimbosa, Galeravolva Cate, 1978: Venus 37(4):192-193, pl. 1, fig. 2.
HOLOTYPE: LACM 1830 (dry), 70-90 m, off Wakayama, Kii Strait, Wakayama Pref., Honshu,
Japan (31 10 N, 135 15 E).
Remarks: Currently in the genus Habuprionovolva fide Fehse (2001). Incorrect coordinates
cited by Cate; correct coordinates = 34 10 N, 135°05’E.
ostheimerae, Crenavolva (Cuspivolva) Cate, 1973: The Veliger, v. 15 supplement, p. 58.
PARATYPE: LACM 1868 (1 dry), 18-29 m, Geelvink Bay, ca. 1 mi. SE of Cape
Dgaroewawoffi, Yapen Id., Irian Jaya, Papua-New Guinea (2 30 S, 135 30 E), Leg.: Osteimer
Orr, Powell Exped., 13 FEB 1956.
Remarks: Currently in the genus Cuspivolva fide Fehse (2001).
periopsis, Crenavolva (Crenavolva) Cate, 1978a: The Nautilus 92(4):163-164, fig. 5.
HOLOTYPE: LACM 1791 (dry), Soerabaja, Java, Indonesia (7 22 S, 112 40 E).
Remarks: Junior synonym of Crenavolva virgo (Azuma & Cate, 1971) fide Lorenz & Fehse
(2009).
platysia, Primovula (Primovula) Cate, 1973: The Veliger, v. 15 supplement, p. 42-43.
PARATYPE: LACM 1862 (dry), Mackay, Queensland, Australia, Leg.: J. Howland, FEB 1973.
Remarks: Currently in the genus Cuspivolva fide Lorenz & Fehse (2009).
pontia, Aperiovula Cate, 1975: The Veliger 17(3):255-256, fig. 2.
HOLOTYPE: LACM 1701 (dry); PARATYPE: LACM 1889 (1 dry), 137 m, off Hinomi-saki,
Kii-suido, Wakayama Pref., Honshu, Japan (34 00 N, 134 48 E), Leg.: S. Habu, SEP 1973.
Remarks: Junior synonym of Sandalia tritcea (Lamarck, 1810) fide Lorenz & Fehse (2009).
pustulata, Cypraea Lightfoot, 1786: Cat. Portland Mus., p. 106.
NEOTYPE: LACM 1422 (1 dry), designated by Cate (1973:5, fig. 2), The Veliger, v. 15
supplement. Puertecitos, Golfo de California, Baja California, México, MAR 1960.
Remarks: Currently in the genus Jenneria.
quaylei, Simnia Lowe, 1935: Trans. San Diego Soc. Nat. Hist. 8(6):22, pl, 3, fig. 5.
HOLOTYPE: LACM 2163 (dry); PARATYPES: LACM 2164 (2 dry), 0 m, intertidal, San
Felipe, Golfo de California, Baja California, México, Leg.: H.N. Lowe, 1933.
Remarks: Junior synonym of Neosimnia arcuata (Reeve, 1865) fide Lorenz & Fehse (2009).
Holotype ex UCLA 3388, paratypes ex UCLA 3389.
rhomba, Cyphoma Cate, 1978a: The Nautilus 92(4):164-165, figs. 7-8.
HOLOTYPE: LACM 1792 (dry), 18 m, Fort Lauderdale Reef, Broward Co., Florida (27 00 N,
80 00 W), Leg.: W. Chapman, 20 FEB 1971.
robertsoni, Aperiovula Cate, 1973: The Veliger, v. 15 supplement, p. 39.
PARATYPE: LACM 1895 (1 dry), 80 m, off Kaohsiung, Kao Hsiung Co., Taiwan.
Remarks: Currently in the genus Quasisimnia fide Lorenz & Fehse (2009).
rutherfordiana, Primovula (Adamantia) Cate, 1973: The Veliger, v. 15 supplement, p. 47.
PARATYPE: LACM 1865 (1 dry), Townsville, Queensland, Australia (19 14 S, 146 45 E),
Leg.: T. Neilsen, MAY 1969.
Remarks: Currently in the genus Dentiovula fide Lorenz & Fehse (2009).
segaliana, Cymbovula Cate, 1976b: The Veliger 19(2):160-161, fig. 1.
HOLOTYPE: LACM 1770 (dry), Pinang Id., Pinang, Malaysia (5 25 N, 100 15 E).
Remarks: Junior synonym of Cuspivolva queenslandica (Cate, 1974) fide Lorenz & Fehse
(2009).
shikamai, Aperiovula Cate, 1973: The Veliger, v. 15 supplement, p. 37-38.
PARATYPE: LACM 1894 (1 dry), Between Ise-wan, Aichi Pref. and Izu-hanto, Shizuoka Pref.,
Honshu, Japan (34 44 N, 136 44 E), 1968.
Remarks: Currently in the genus Pseudosimnia fide Lorenz & Fehse (2009).
spectabilis, Dentiovula Cate, 1975: The Veliger 17(3):257-258, fig. 8.
HOLOTYPE: LACM 1702 (dry), 18 m, off Hinomi-saki, Kii Strait, Wakayama Pref., Honshu,
Japan (34 00 N, 143 48 E).
Remarks: Junior synonym of Dentiovula eizoi Cate & Azuma in Cate, 1973 fide Lorenz &
Fehse (2009).
sphaera, Globovula Cate, 1973: The Veliger, v. 15 supplement, p. 22-23.
PARATYPE: LACM 1888 (1 dry), 4 m, Singapore.
stigma, Pseudosimnia (Inflatovula) Cate, 1978b: Venus 37(4):196-197, pl. 2, fig. 6.
HOLOTYPE: LACM 1831 (dry), 80 m, off Wakayama, Kii-suido, Wakayama Pref., Honshu,
Japan (31 10 N, 135 15 E).
Remarks: Currently in the genus Diminovula fide Lorenz & Fehse (2009). Incorect coordinates
cited by Cate, correct coordinates = 34 10 N, 135°05’E.
stohleri, Stohleroma Cate, 1973: The Veliger, v. 15 supplement, p. 47-48, fig. 97.
HOLOTYPE: LACM 1188 (dry), Ogokuda Beach, Shio-no-misaki, Wakayama Pref., Honshu,
Japan (34 00 N, 134 48 E), DEC 1969.
Remarks: Considered a nomen dubium fide Lorenz & Fehse (2009:78).
striola, Pseudosimnia (Diminovula) Cate, 1973: The Veliger, v. 15 supplement, p. 28-29.
PARATYPE: LACM 1861 (1 dry), Preparis Id., SW of mouths of the Irrawaddy River, Preparis
North Channel, Myanmar (= Burma).
Remarks: Currently in the genus Diminovula fide Fehse (2001).
takae, Aperiovula Cate, 1973: The Veliger, v. 15 supplement, p. 38-39.
PARATYPE: LACM 1896 (1 dry), 73-91 m, 2-3 km off Kiri-misaki, Kii Peninsula, Wakayama
Pref., Honshu, Japan.
Remarks: Junior synonym of Sandalia meyeriana (Cate, 1973) fide Lorenz & Fehse (2009).
takaeopsis, Aperiovula Cate, 1978b: Venus 37(4):198, pl. 2, fig. 8.
HOLOTYPE: LACM 1833 (dry), 80 m, off Wakayama, Kii-suido, Wakayama Pref., Honshu,
Japan (31 10 N, 135 15 E).
Remarks: Junior synonym of Sandalia meyeriana (Cate, 1973) fide Lorenz & Fehse (2009).
Incorrect coordinates cited by Cate, correct coordinates = 34 10 N, 135°05’E.
tokioi, Phenacovolva (Calcaria) Cate, 1973: The Veliger, v. 15 supplement, p. 108, fig. 242.
HOLOTYPE: LACM 1190 (dry), Cooktown, Queensland, Australia (15 38 S, 145 25 E), 13
JAN 1958.
Remarks: Junior synonym of Phenacovolva nectarea Iredale, 1930 fide Lorenz & Fehse (2009).
uvula, Primovula (Primovula) Cate, 1978a: The Nautilus 92(4):163, fig. 4.
HOLOTYPE: LACM 1790 (dry), Moreton Bay, Queensland, Australia (27 12 S, 153 12 E),
Leg.: E. Grigg.
virgo, Primovula Azuma & Cate, 1971: The Veliger 13(3):262-263.
PARATYPE: LACM 1873 (1 dry), 91 m, off Hino-misaki, Kii-suido, Wakayama Pref., Honshu,
Japan, Leg.: S. Habu, 20 MAR 1970.
Remarks: Currently in the genus Crenavolva fide Fehse (2001).
whitworthi, Pseudosimnia (Diminovula) Cate, 1973: The Veliger, v. 15 supplement, p. 30.
PARATYPE: LACM 1860 (1 dry), Ogokuda Beach, Shio-no-misaki, Kii-suido, Wakayama Pref.,
Honshu, Japan (34 00 N, 134 48 E).
Remarks: Currently in the genus Diminovula fide Fehse (2001).
Superfamily VELUTINOIDEA Gray, 1840
Family Triviidae Troschel, 1863
akroterion, Pusula (Dolichupis) Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 108, fig. 80.
HOLOTYPE: LACM 1837 (dry), Coron, Palawan Prov., Philippines (12 10 N, 120 13 E), Leg.:
F. Steiner.
Remarks: Currently in the genus Trivirostra fide Fehse (1998).
aquatanica, Niveria (Cleotrivia) Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 67, fig. 78.
HOLOTYPE: LACM 1797 (dry), Bulalacao, Mindoro Oriental Prov., Mindoro Id., Philippines
(12 31 N, 121 26 E).
Remarks: Considered a junior synonym of Niveria nix by Fehse & Grego (2004).
artema, Decoriatrivia Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 96, fig. 88.
HOLOTYPE: LACM 1801 (dry), 59 m, Bahía Stephen, Isla San Cristobal (= Chatham Id.), Islas
Galápagos (0 47 S, 89 31 W), Leg.: R/V Velero III, 21 JAN 1934.
Remarks: Currently in the genus/subgenus Trivia (Decoriatrivia) by Fehse (2002).
aussiorum, Trivirostra Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 93-95, fig. 150.
HOLOTYPE: LACM 1799 (dry), Lighthouse Beach, Vlaming Head, North West Cape, Western
Australia (21 50 S, 114 10 E).
Remarks: Currently in the genus Austrotrivia fide Fehse (2002).
austrafricana, Triviella Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 23, fig. 14.
HOLOTYPE: LACM 1827 (dry), St. Francis Bay, Cape Prov., South Africa (34 02 S, 24 58 E),
Leg.: H. Boswell, DEC 1951.
boswellae, Trivirostra Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 73, fig. 116.
HOLOTYPE: LACM 1805 (dry), Port Shepstone, Natal, South Africa (30 46 S, 30 25 E),
Leg.: H. Boswell.
burius, Decoriatrivia Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 96-97, fig. 90.
HOLOTYPE: LACM 1823 (dry), Bahía de Panama, Panamá Prov., Panamá (8 50 N, 79 15 W).
Remarks: Currently in the genus/subgenus Trivia (Decoriatrivia) fide Fehse (2002).
campus, Pusula (Pusula) Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 101-102, fig. 101.
HOLOTYPE: LACM 1808 (dry), Bahía Cholla, Sonora, México (27 88 N, 111 00 W), Leg.: A.
Sorensen, 1945.
Remarks: Currently in the genus Niveria fide Fehse (2002). Incorrect coordinates cited by Cate,
correct coordinates = 31 19.6'N, 113 37.8 W.
carabus, Pusula (Pusula) Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 100-101, fig. 99.
HOLOTYPE: LACM 1838 (dry), Golfo de Guayaquil, Guayas Prov., Ecuador (3 00 S,
80 30 W).
Remarks: Currently in the genus Niveria fide Fehse (2002).
cherobia, Pusula (Pusula) Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 101, fig. 100.
HOLOTYPE: LACM 1803 (dry), 18 m, Bahía Magdalena, Pacific Coast Baja California Sur,
México (24 30 N, 112 00 W), Leg.: H.N. Lowe, 1931.
Remarks: Currently in the genus Niveria fide Fehse (2002).
citeria, Decoriatrivia Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 97-98, fig. 93.
HOLOTYPE: LACM 1802 (dry), 55 m, Bahía James, Isla San Salvador (= James Id.), Islas
Galápagos (0 10 S, 90 52 W), Leg.: R/V Velero III, 24 JAN 1934.
Remarks: Currently in the genus Dolichupis fide Fehse (2002).
clariceae, Trivirostra Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 74, fig. 119.
HOLOTYPE: LACM 1820 (dry), Jeffreys Bay, Cape Prov., South Africa (34 01 S, 24 55 E).
dumaliensis, Pseudotrivia Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 40, fig. 42.
HOLOTYPE: LACM 1796 (dry), Off Dumali Pt., Mindoro Occidental Prov., Mindoro Id.,
Philippines (13`17'N, 121`33'E).
Remarks: Considered a junior synonym of Niveria nix by Fehse & Grego (2004).
exmouthensis, Trivirostra Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 90, fig. 144.
HOLOTYPE: LACM 2230 (dry, Near Geralia (= Giralia), Exmouth Gulf, Western Australia
(22 00 S, 114 15 E), Leg.: Leslie Figgis.
Remarks: Currently in the genus Austrotrivia fide Fehse (2002). Originally published as LACM
1797, preoccupied. Incorrect latitude cited by Cate, correct latitude = 22°19'S.
geralia, Proerato (Sulcerato) Cate, 1977: Veliger 19(3):348-349, fig. 13.
HOLOTYPE: LACM 1763 (dry); PARATYPE: LACM 1875 (1 dry), Geralia (= Giralia),
Exmouth Gulf, Western Australia (22 00 S, 114 15 E), Leg.: M. Gedling & L. Figgis.
Remarks: Incorrect latitude cited by Cate, correct latitude = 22°19'S.
halians, Decoriatrivia Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 96, fig. 89.
HOLOTYPE: LACM 1822 (dry), San Jose, Escuintla Dept., Guatemala (13 57 N, 90 49 W).
Remarks: Currently in the genus Dolichupis fide Fehse (2002).
hispania, Trivia (Trivia) Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 30, fig. 25.
HOLOTYPE: LACM 1824 (dry), At the Rio Mino (= Mimbo) mouth, S of Pontevedra,
Pontevedra Prov., Spain (42 27 N, 8 39 W).
iota, Circumscapula Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 110, fig. 154.
HOLOTYPE: LACM 1839 (dry), 18-21 m, Bahía San Carlos, Guaymas, Sonora, México
(111 00 N, 27 20 W), Leg.: L. Shy.
Remarks: Currently in the genus Dolichupis fide Fehse (2002). Coordinates reversed by Cate,
correct coordinates = 27 20 N, 111 00 W.
keehiensis, Trivirostra Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 85, fig. 137.
HOLOTYPE: LACM 1825 (dry), 45-109 m, off Ke‘ehi Lagoon entrance, Honolulu Dist., O‘ahu,
Hawaiian Ids. (21 17 N, 157 54 W), Leg.: T. Bratcher, 11 JUL 1975.
koto, Lamellaria (Lamellaria) Schwengel, 1944: The Nautilus 58(1):17-18.
PARATYPE: LACM 1216 (1 dry), Near Bradenton Beach, Manatee Co., Florida, Leg.: A.S.
Koto, 11 JAN-23 FEB 1944.
loochooensis, Pusula (Pusula) california Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p.
104, fig. 106c.
HOLOTYPE: LACM 1840 (dry), Ryukyu-shoto, Japan (24 40 N, 126 10 E).
Remarks: Junior synonym of Pusula californica (Sowerby, 1832).
millardi, Galeatrivia Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 15-16, fig. 173.
HOLOTYPE: LACM 1843 (dry), Sea Pt., Cape Prov., South Africa (34 38 S, 19 17.60 E),
Leg.: V. Millard, 2 AUG 1977.
Remarks: Currently in the genus Triviella fide Fehse (2002).
oshimaensis, Trivirostra Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 73-74, fig. 117.
HOLOTYPE: LACM 1819 (dry), Oshima Id., Amami-o-shima, Amami-shoto, Japan (28 15 N,
129 15 E).
padreserrai, Pusula (Pusula) Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 105-106, fig.
109.
HOLOTYPE: LACM 1828 (dry), Bluff Cove, Palos Verdes Peninsula, Los Angeles Co.,
California (33 45 N, 118 25 W), Leg.: B.C. Draper.
Remarks: Junior synonym of Pusula solandri (Sowerby, 1832) but considered a valid species by
Fehse (2002).
polynesiae, Trivirostra Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 85, fig. 136.
HOLOTYPE: LACM 1121 (dry), Club Mediterranean resort, W side Moorea, Society Ids.,
French Polynesia (17 30 S, 149 46 W), Leg.: D.E. Koontz, 28 JAN-1 FEB 1974.
procella, Trivia (Trivia) Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 32, fig. 28.
HOLOTYPE: LACM 1815 (dry), Barbados (13 15 N, 59 30 W).
rehderi, Proerato (Sulcerato) Raines, 2002a: La Conchiglia 34(304):22-23, fig. 19.
HOLOTYPE: LACM 2940 (dry); PARATYPE LACM 2941 (1 dry), 10-20 m, off Punta Rosalia,
E of Anakena, Isla de Pascua (= Easter Id.), Leg.: B.K. Raines.
rubra, Trivia Shaw, 1909: Proc. Malac. Soc. London 8(5):312.
SYNTYPE [as "COTYPE"]: LACM 1321 (1 dry), Jeffreys Bay, Cape Prov., South Africa.
Remarks: Currently in the genus Triviella.
samarensis, Pseudotrivia Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 40, fig. 41.
HOLOTYPE: LACM 1795 (dry), Borongan, Samar Oriental Prov., Samar Id., Philippines
(11 36 N, 125 27 E).
Remarks: Currently in the genus Trivellona fide Fehse (2002).
rubra, Velutina Willett, 1919: The Nautilus 33(1):25-26.
PARATYPES: LACM 2158 (4 dry), Forrester Id., Prince of Wales-Outer Ketchikan Co., Alaska,
Leg.: George Willett. Ex UCLA 21465.
septentrionalis, Hesperato Cate, 1977: The Veliger 19(3):365, fig. 51.
HOLOTYPE: LACM 1762 (dry), Lighthouse Beach, Vlaming Head, North West Cape, Western
Australia (21 47 S, 114 10 E).
sharoni, Lamellaria (Lamellaria) Willett, 1939b: The Nautilus 52(4):123-124, pl. 9, figs. 1-1b.
HOLOTYPE: LACM 1059 (dry); PARATYPES: LACM 1092 (2 dry), Anaheim Bay, Orange
Co., California, Leg.: R.E. Sharon.
spioinsula, Trivirostra Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 81, fig. 129.
HOLOTYPE: LACM 1807 (dry), Ableta, ca. 35 km from Jingo, Rossel Id., Louisiade Arch.,
Papua New Guinea (11 30 S, 154 00 E), Leg.: J.M. Chow, 26 OCT 1963.
stalagmia, Proerato (Sulcerato) Cate, 1975: The Veliger 17(3):261, fig. 6.
HOLOTYPE: LACM 1703 (dry), Maqueda Bay, Samar Occidental Prov., Philippines (11 30 N,
125 00 E).
thaanumi, Trivirostra Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 70, fig. 113.
HOLOTYPE: LACM 1826 (dry), Lihue, Lihue Dist., Kaua‘i, Hawaiian Ids. (21 59 N,
159 24 W), Leg.: D. Thaanum.
tortuga, Trivia (Trivia) Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 30-32, fig. 27.
HOLOTYPE: LACM 1817 (dry), Playa Veracruz, Bahía de Panamá, Panamá Prov., Panamá,
Leg.: B.D. Harris.
vitrina, Trivirostra Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 79, fig. 126.
HOLOTYPE: LACM 1809 (dry), Malampaya Sound, Palawan Id., Palawan Prov., Philippines
(10 57 N, 119 15 E).
zzyzyxia, Trivirostra Cate, 1979: San Diego Soc. Nat. Hist. Mem. 10, p. 93, fig. 148.
HOLOTYPE: LACM 1800 (dry), Lighthouse Beach, Vlaming Head, North West Cape, Western
Australia (21 47 S, 114 10 E), Leg.: Leslie Figgis.
Remarks: Currently in the genus Austrotrivia fide Fehse & Grego (2004).
Superfamily TONNOIDEA Suter, 1913
Family Ranellidae Gray, 1854
longicaudatum, Gyrenium Beu, 1998: Mém. Mus. Nat. Hist. Nat. 178:54-57.
PARATYPE: LACM 2789 (1 dry), 400 m, off Tudela, Pacijan Id., between Leyte and Cebu,
Philippines (10 40 N, 124 35 E), Leg.: F.J. Springsteen, 29 OCT 1987.
Superfamily NATICOIDEA Guilding, 1834
Family Naticidae Guilding, 1834
brunneolinea, Natica (Natica) McLean, 1970b: The Veliger 12(3):313, pl. 46, fig. 9.
HOLOTYPE: LACM 1282 (dry); PARATYPE: LACM 1283 (1 dry), 91-183 m, Bahía Academy,
Isla Santa Cruz (= Indefatigable Id.), Islas Galápagos (0 45 S, 90 20 W), Leg.: A. & J. DeRoy,
26 APR 1967.
coani, Choristes Marincovich, 1975: The Veliger 18(2):169-171, fig. 7.
PARATYPES: LACM 1733 (2 dry), 2760 m, 68.7 mi. 274.8 degrees T from Government Pt.,
near Depoe Bay, Lincoln Co., Oregon (44 55.4 N, 125 40.6 W), Leg.: R/V Yaquina, 17 MAR
1970.
Remarks: Species excluded from Choristes (McLean, 1992b:293) and is in need of generic
reassignment.
cortezi, Sinum J.Q. Burch & R.L. Burch, 1964: The Nautilus 77(4):109-110.
PARATYPES: LACM 2223 (6 dry), 27 m, between Altata and Mazatlán, Sinaloa, México.
Remarks: Junior synonym of Sinum grayi (Deshayes, 1843) fide Marincovich (197).
lamonae, Neverita (Neverita) Marincovich, 1975: The Veliger 18(2):168-169.
PARATYPE: LACM 1732 (1 dry), 2816 m, abyssal plain off central, Oregon (44 34.8 N,
125 33.6 W), Leg.: R/V Yaquina, 16 MAR 1970.
Remarks: Radula mounted on glass slide.
sigillata, Natica (Natica) McLean, 1970b: The Veliger 12(3):313-314, pl. 46, fig. 11.
HOLOTYPE: LACM 1284 (dry), 91 m, Caleta Tagus, Isla Isabela (= Albemarle Id.), Islas
Galápagos (0 16 N, 91 22 W), Leg.: A. & J. DeRoy, 25-29 JAN 1968; PARATYPE: LACM
1285 (1 dry),150 m, Bahía Academy del Sur, Isla Santa Cruz (= Indefatigable Id.), Islas
Galápagos, Leg.: A. & J. DeRoy, 11 JUN 1968.
Family Tonnidae Suter, 1913
harrisonae, Xenophalium Powell, 1928: Trans. Proc. New Zealand Inst. 59(3):640.
PARATYPE: LACM 1318 (1 dry), Mason's Bay, Stewart Id., New Zealand.
Remarks: Junior synonym of Semicassis pyrum (Lamarck, 1822) fide Kreipl (1997).
“Group” Ptenoglossa
Superfamily TRIPHOROIDEA Gray, 1847
Family Triphoridae Gray, 1847
marshi, Triphora Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):214-215.
PARATYPE: LACM 1554 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
palmeri, Triphora Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):209, 214.
PARATYPE: LACM 1553 (1 dry), 5-16 m, off Isla Taboga, Panamá Prov., Panamá, Leg.: L.G.
Hertlein.
Family Cerithiopsidae H. Adams & A. Adams, 1853
eiseni, Cerithiopsis Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):216.
PARATYPE: LACM 1555 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Herrtlein.
gissleri, Cerithiopsis Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):216-217.
PARATYPES: LACM 1709 (2 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
montezumai, Cerithiopsis Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):217-218.
PARATYPES: LACM 1710 (2 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
pulmoensis, Seila DuShane & Draper, 1975: The Veliger 17(4):342-343, fig. 26.
HOLOTYPE: LACM 1728 (dry), 0 m, intertidal, S side of Cabo Pulmo, Golfo de California,
Baja California Sur, México (23 25 N, 109 25 W), Leg.: H. DuShane, 26 JAN 1967;
PARATYPES: LACM 1729 (7 dry), 0.0-3 m, intertidal, S side of Cabo Pulmo, Golfo de
California, Baja California Sur, México (23 25 N, 109 25 W).
Superfamily EPITONIOIDEA Berry, 1910
Family Epitoniidae Berry, 1910
arcana, Epitonium (Nitidiscala) DuShane, 1979: The Veliger 21(3):379-380, fig. 1.
HOLOTYPE: LACM 1264 (dry); PARATYPES: LACM 1265 (4 dry), 0 m, intertidal,
Puertecitos, Golfo de California, Baja California, México (30 20.03 N, 114 38.13 W), Leg.: H.
DuShane, OCT 1970.
Remarks: Currently in the subgenus Hyaloscala fide Weil & others (1999).
billeeana, Scalina DuShane & Bratcher, 1965: The Veliger 8(2):160-161.
PARATYPE: LACM 1095 (1 dry)
2.4-3 m, SW end of Isla Cerralvo, Golfo California, Baja California Sur, México, Leg.: B.
Dillworth, N. Currin, & T. Bratcher, APR 1965.
Remarks: Questionably in the subgenus Limiscala? fide Weil & others (1999).
bormanni, Epitonium (Nitidoscala) tinctum var. Strong, 1941: The Nautilus 55(2):47.
HOLOTYPE: LACM 1064 (dry); PARATYPES: LACM 1842 (5 dry); PARATYPES: LACM
1859 (25 dry), 0 m, intertidal, Mission Bay, San Diego, San Diego Co., California, Leg.: R. & M.
Bormann, 1939.
Remarks: Junior synonym of Epitonium tincta (Carpenter, 1864) fide Weil & others (1999).
burchorum, Opalia DuShane, 1988: The Veliger 31(3/4):270.
PARATYPE: LACM 2303 (1 dry), 540 m, off Kahe Pt., ‘Ewa Dist., O‘ahu, Hawaiian Ids.
(21 21.2 N, 158 09.1 W), Leg.: T.A. & B.L. Burch, MAR 1978.
chacei, Opalia Strong, 1937: The Nautilus 51(1):5-6, pl. 2, fig. 9.
HOLOTYPE: LACM 1045 (dry), Crescent City, Del Norte Co., California, Leg.: E.P. Chace.
Remarks: Junior synonym of Opalia borealis Keep, 1881 fide Weil & others (1999).
cerralvoensis, Epitonium DuShane, 1970b: L.A. Co. Mus. Cont. Sci. 185:3-4, fig. 3.
HOLOTYPE: LACM 1200 (dry); PARATYPE: LACM 1201 (1 dry), 13 m, W side of Isla
Cerralvo, Golfo de California, Baja California Sur, México (24 11 N, 109 55 W), Leg.: J.H.
McLean, P. Oringer & L. Marincovich, 09 APR 1966.
Remarks: Currently in the genus Acirsa fide Weil & others (1999).
contexta, Amaea (Amaea) DuShane, 1970b: L.A. Co. Mus. Cont. Sci. 185:5-6, fig. 5.
HOLOTYPE: LACM 1199 (dry), 9-18 m, Bahía Petatlán, Guerrero, México (17 31.7 N,
101 27.57 W), Leg.: R/V Velero III, 03 MAR 1934.
cooperi, Epitonium Strong, 1930: Trans. San Diego Soc. Nat. Hist. 6(7):194.
PARATYPES: LACM 2171 (6 dry), San Pedro, Los Angeles Co., California.
Remarks: Junior synonym of Epitonium hindsii (Carpenter, 1856) fide Weil & others (1999). Ex
UCLA 4777.
deroyae, Amaea (Scalina) DuShane, 1970a: The Veliger 12(3):330-331, pl. 51, fig. 2.
HOLOTYPE: LACM 1236 (dry); PARATYPES: LACM 1239 (4 dry), 55 m, Caleta Tagus, Isla
Isabela (= Albemarle Id.), Islas Galápagos (0 16.63 S, 91 22.73 W), Leg.: R/V Velero III, 13
JAN 1934.
Remarks: Holotype and paratype published as LACM-AHF 147-34.
goldsmithi, Asperiscala DuShane, 1988: The Veliger 31(3/4):269, fig. 4.
HOLOTYPE: LACM 2304 (dry); PARATYPES: LACM 2305 (2 dry), 0 m, intertidal, Puhi Bay,
Hilo, So. Hilo Dist., Hawai‘i, Hawaiian Ids. (19 44 N,155 03 W), Leg.: M. Goldsmith, 1983.
Remarks: Currently in the genus/subgenus Epitonium (Asperiscala) fide Weil & others (1999).
hancocki, Epitonium (Nitidiscala) DuShane, 1970a: The Veliger 12(3):332, pl. 51, fig. 1.
HOLOTYPE: LACM 1235 (dry), 37-73 m, N of Isla Espanola (= Hood Id.), Islas Galápagos
(1 21.92 S, 90 40.08 W), Leg.: R/V Velero III, 28 JAN 1938; PARATYPE: LACM 1238 (1 dry),
457 m, NW of Isla Santa Maria (Charles), Islas Galápagos (1 09 S, 90 35 W), Leg.: R/V Velero
III, 23 JAN 1938.
Remarks: Holotype published as LACM-AHF 814-38, paratype published as LACM-AHF 80238.
hemmesi, Epitonium (Epitonium) DuShane, 1988: The Veliger 31(3/4):267.
PARATYPES: LACM 2306 (2 dry), 20-27 m, Lahilahi Pt., near Makaha Beach, Wai‘anae Dist.,
O‘ahu, Hawaiian Ids. (21 28 N, 158 14.2 W), Leg.: D. Hemmes & S. Jazwinski, 1985-1986.
huffmani, Epitonium DuShane & McLean, 1968: L.A. Co. Mus. Cont. Sci. 145:1-2, fig. 1.
HOLOTYPE: LACM 1159 (dry), Intertidal, Bahía Cholla, Bahía Adair, Sonora, México
(31 21 N, 113 40 W), Leg.: Al Huffman, 11 APR 1937.
Remarks: Questionably in the subgenus Pupiscala? fide Weil & others (1999).
longinosanum, Epitonium DuShane, 1970b: L.A. Co. Mus. Cont. Sci. 185:1-2, fig. 1.
HOLOTYPE: LACM 1196 (dry), Rancho Inocentes, Pacific Coast Baja California Sur, México
(23 45 N, 110 40 W), Leg.: Fred Lewis.
Remarks: Junior synonym of Epitonium aciculinum (Hinds, 1844) fide Weil & others (1999).
luceo, Laeviscala DuShane, 1988: Veliger 31(3/4):270.
PARATYPE: LACM 2325 (1 dry), 17 m, off Sand Id., Honolulu Dist., O‘ahu, Hawaiian Ids.
(21 16.3 N, 157 50.7 W), Leg.: S. Jazwinski.
Remarks: Currently in the genus/subgenus Epitonium (Laeviscala) fide Weil & others (1999).
macleani, Epitonium DuShane, 1970b: L.A. Co. Mus. Cont. Sci. 185:3, fig. 2.
HOLOTYPE: LACM 1197 (dry), 27-37 m, Punta Arena de la Ventana, Golfo de California, Baja
California Sur, México, Leg.: J.H. McLean, P. Oringer & L. Marincovich, 08 APR 1966;
PARATYPES: LACM 1198 (8 dry), 37-283 m, off Punta Arena de la Ventana, Golfo de
California, Baja California Sur, México (24 04 N, 109 49 W), Leg.: J.H. McLean, P. Oringer &
L. Marincovich, 08 APR 1966.
Remarks: Currently in the subgenus Limiscala fide Weil & others (1999).
murrha, Epitonium DuShane, 1970b: L.A. Co. Mus. Cont. Sci. 185:4-5, fig. 4.
HOLOTYPE: LACM 1232 (dry), 55 m, 10 mi. SW of Isla Secas, Chiriqui Prov., Panamá,
Leg.: R/V Velero III, 27 MAR 1939.
Remarks: Currently in the subgenus Acirsa fide Weil & others (1999).
paulula, Opalia DuShane, 1974: The Veliger, v. 16 supplement, p. 72, fig. 152.
HOLOTYPE: LACM 1611 (dry), 11 m, Bahía Cuastecomate, Jalisco, México (19 13 N,
104 44 W), Leg.: Helen DuShane, 5 FEB 1969.
Remarks: Currently in the subgenus Dentiscala fide Weil & others (1999).
pedroanum, Epitonium Willett, 1932c: Trans. San Diego Soc. Nat. Hist. 7(9):88-89, pl. 5, fig. 3.
HOLOTYPE: LACM 1036 (dry); PARATYPES: LACM 2150 (3 dry), 37 m, San Pedro, Los
Angeles Co., California, Leg.: G. Willett.
Remarks: Junior synonym of Epitonium politum (Sowerby, 1844) fide Weil & others (1999).
Paratypes ex UCLA 21272.
pigrum, Epitonium García, 2011: Novapex 12(3-4):102-104, fig. 38.
PARATYPE: LACM 3189 (dry), 63-65 m, Bahía de Campeche, NE of Campeche, Campeche,
México (20 52.40 N, 92 24.83 W), 10 JUN 2005.
shyorum, Epitonium DuShane & McLean, 1968: L.A. Co. Mus. Cont. Sci. 145:2-4, fig. 2.
HOLOTYPE: LACM 1160 (dry), 22-24 m, Manzanillo, Colima, México, Leg.: L. & C. Shy,
NOV 1965.
Remarks: Currently in the subgenus Hirtoscala fide Weil & others (1999).
skoglundae, Epitonium DuShane, 1974: The Veliger, v. 16 supplement, p. 72, figs., 150-151,
153.
HOLOTYPE: LACM 1612 (dry); PARATYPES: LACM 1613 (5 dry), Ft. Amador, Pacific Coast
of Panama Canal Zone, Panamá (8 58 N, 79 33 W), Leg.: C. Skoglund, MAR 1972.
Remarks: Questionably in the subgenus Nitidiscala? fide Weil & others (1999).
tehuanarum, Amaea (Scalina) DuShane & McLean, 1968: L.A. Co. Mus. Cont. Sci. 145:4-6,
fig. 4.
HOLOTYPE: LACM 1162 (dry); PARATYPE: LACM 1163 (1 dry), 59-68 m, Golfo de
Tehuantepec, Oaxaca, México (15 58 N, 95 00 W), Leg.: D.R. Shasky & X. Mendoza, JUL
1963.
Remarks: Holotype erroneously published as LACM 1161, paratype erroneously published
as LACM 1162.
textimatta, Epitonium DuShane, 1977: The Nautilus 91(3):89-91, fig. 1.
HOLOTYPE: LACM 1846 (dry), Playa los Angeles, Bahía Tenacatita, Jalisco, Mexico (19 18 N,
104 50 W), Leg.: C. & P. Skoglund, AUG 1975.
Remarks: Currently in the subgenus Sodaliscala fide Weil & others (1999).
thorssoni, Epitonium (Epitonium) DuShane, 1988: The Veliger 31(3/4):269.
PARATYPES: LACM 2302 (2 dry), 62-80 m, off Diamond Head, Honolulu Dist., O‘ahu,
Hawaiian Ids. (21 15.8 N, 157 50 W), Leg.: W.M. Thorsson, 1980-1981.
Superfamily EULIMOIDEA Philippi, 1853
Family Eulimidae Philippi, 1853
callista, Thyca (Bessomia) Berry, 1959: Leaflets in Malacology 1(18):110-111.
HOLOTYPE: LACM 1120 (dry), 5-7 m, Bahía San Carlos, Sonora, México, Leg.: L.C. Bessom,
15 JUL 1959.
clarimaculosa, “Eulima” Raines, 2002b: La Conchiglia 34(305):47-48, fig. 6.
HOLOTYPE: LACM 2959 (dry); PARATYPE: LACM 2960 (1 dry), 10-20 m, off Punta Rosalia,
E of Anakena, Isla de Pascua (= Easter Id.), Leg.: B.K. Raines.
delmontensis, Balcis Smith & Gordon, 1948: Proc. Calif. Acad. Sci., 4th ser., 26(8):219-220.
PARATYPE: LACM 1005 (1 dry), 18 m, off Del Monte, Monterey Co., California, Leg.: A.G.
Smith & C.S. Fackenthall, 16 AUG 1913.
Remarks: Currently in the genus Melanella.
emersoni, Niso (Niso) McLean, 1970a: Malac. Review 2:128, figs. 39-40.
HOLOTYPE: LACM 1403 (dry); PARATYPE: LACM 1404 (1 dry), 13-20 m, off Puerto de San
Jose, Escuintla Dept., Guatemala (13 52.5 N, 91 10.5 W), Leg.: R/V Velero III, 11 JAN 1938.
healeyi, Strombiformis Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):195.
PARATYPE: LACM 2243 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
Remarks: Currently in the genus Eulima fide Keen (1971).
macleani, Eulimostraca Warén, 1992: The Veliger 35(3):187, figs. 11, 32, 34, 40.
HOLOTYPE: LACM 2370 (dry); PARATYPE: LACM 2371 (1 gold coated), 21.3 m, Bahía
Herradura, Puntarenas Prov., Costa Rica (9 38'N, 84 40.5 W), Leg.: J.H. McLean, 9 MAR 1972.
mulata, Eulima Rios & Absalao, 1990: Rev. Bras. Biol. 50(1):61-63.
PARATYPE: LACM 2208 (1 dry), 24 m, off Salinopolis, Para State, Brazil, Leg.: N.O.C.
Saldanha, 21 NOV 1986.
osorio, Subniso Raines, 2002b: La Conchiglia 34(305):46-47, fig. 5.
HOLOTYPE: LACM 2957 (dry); PARATYPE: LACM 2958 (1 dry), 10-20 m, off Punta Rosalia,
E of Anakena, Isla de Pascua (= Easter Id.), Leg.: B.K. Raines.
pagoda, Eulimetta Warén, 1992: The Veliger 35(3):187-189, figs. 14, 36-39, 41.
HOLOTYPE: LACM 2372 (dry); PARATYPES: LACM 2373 (1 dry, 1 gold coated), 18.3-36.6
m, Bahía Cuastecomate, Jalisco, México (19 13.8 N, 104 44.9 W), Leg.: J.H. McLean, P.
Oringer, & G. Radwin, 21 OCT 1968.
pallida, Turveria Warén, 1992: The Veliger 35(3):179, figs. 7-8, 13.
HOLOTYPE: LACM 2425 (dry); PARATYPES: LACM 2426 (1 dry, 1 gold coated), Living on
Encope grandis on sand flats at Isla Willard, Bahía San Luis Gonzaga, Baja California, México
(29 57 N, 114 17 W), Leg.: T. Schowalter, 28 May 1961.
plenicolora, Sticteulima Raines, 2002b: La Conchiglia 34(305):43-46, fig. 3.
HOLOTYPE: LACM 2955 (dry); PARATYPE: LACM 2956 (1 dry), Hanga Nui, Isla de Pascua
(= Easter Id.), Leg.: B.K. Raines.
randolphi, Eulima Vanatta, 1899: Proc. Acad. Nat. Sci. Philadelphia 51:256.
PARATYPES: LACM 1336 (4 dry); PARATYPES: LACM 2157 (2 dry), Dutch Harbor,
Unalaska Bay, Unalaska Id., Aleutian Ids. Co., Alaska, Leg.: P.B. Randolph.
Remarks: Currently in the genus Melanella. Paratypes 2157 ex UCLA 21202.
rosa, Melanella Willett, 1944: Bull. So. Calif. Acad. Sci. 43(2):72, pl. 14, fig. 2.
HOLOTYPE: LACM 1075 (dry), Off Redondo Beach, Los Angeles Co., California, Leg.: J.Q. &
T.A. Burch.
shaskyi, Sabinella Warén, 1992: The Veliger 35(3):189-193, figs. 42-46, 48-52.
HOLOTYPE: LACM 2374 (gold coated); PARATYPES: LACM 2375 (7 dry), 18.3-36.6 m,
Bahía Cuastesomate, Jalisco, México (19 13.7 N, 104 44.8 W), Leg.: J.H. McLean, P. Oringer,
& G. Radwin, 21 OCT 1968.
Clade Neogastropoda Thiele, 1929
Superfamily BUCCINOIDEA Rafinesque, 1815
Family Buccinidae Rafinesque, 1815
berryi, Cantharus (Gemophos) McLean, 1970b: The Veliger 12(3):314, pl. 46, figs. 13-14.
HOLOTYPE: LACM 1286 (dry); PARATYPES: LACM 1287 (119 dry), 18-27 m, off La Cruz,
Bahía de las Banderas, Jalisco, México (20 45 N, 105 30 W), Leg.: J.H. McLean & A. M. Keen,
20-24 MAR 1965.
castillai, Aenator McLean & Andrade, 1982: Nat. Hist. Mus. L.A. Co. Cont. Sci. 342:14-15.
PARATYPES: LACM 1987 (2 dry), 300 m, off Papudo, Aconcagua Prov., Chile (32 31 S,
71 54 W), Leg.: H. Andrade; R/V Goden Wind, 29 MAR 1977.
clarki, Bathybuccinum (Ovulatibuccinum) Kantor & Harasewych, 1998: Venus 57(2):77-83.
PARATYPE: LACM 2869 (1 dry), 169 m, W of Agattu Id., Aleutian Ids. Co., Alaska
(52 13.50 N, 173 27.80 W), Leg.: R.N. Clark, F/V Dominator, 6 AUG 1997.
insculpta, Tritonoidea Sowerby, 1900: Proc. Malac. Soc. London 4(2):2.
SYNTYPES [as "COTYPES"]: LACM 1323 (2 dry), Port Alfred (= The Kowie), Cape Prov.,
South Africa.
Remarks: Currently in the genus Cantharus fide Steyn & Lussi (1998).
major, Buccinum pemphigus Dall, 1919b: Proc. USNM 56(2295):325-326.
PARATYPES: LACM 1206 (2 dry), 183 m, SE of Kamchatka Oblast, western Bering Sea,
Russia, Leg.: R/V Albatross.
obsoletus, Plicifusus Talmadge, 1971: The Veliger 14(1):43-44.
PARATYPE: LACM 1603 (1 dry), 823 m, between Trinidad & the Klamath River mouth,
Humboldt Co., California (41 05 N, 41 25 W), Leg.: R/V City Of Eureka, 1970.
pribiloffensis, Chrysodomus Dall, 1919b: Proc. USNM 56(2295):323.
PARATYPE: LACM 1289 (1 dry), 113 m, W of Pribilof Ids., Bering Sea, Alaska, Leg.: R/V
Albatross.
Remarks: Currently in the genus Neptunea.
stilesi, Neptunea (Neptunea) A.G. Smith, 1968: The Veliger 11(2):117-120.
PARATYPE: LACM 1152 (1 dry), 155-161 m, British Columbia, Canada, Leg.: I.M. Cowan &
J.H. McLean, 22 MAY 1963; PARATYPE: LACM 1211 (1 dry), 155-161 m, ca. 3 mi. N of Cape
James, Hope Id., Rupert Dist., Vancouver Id., British Columbia, Canada.
thermophilum, Buccinum (Viridibuccinum) Harasewych & Kantor, 2002: Jour. Moll. Stud.
68(1):40-44.
PARATYPES: LACM 2893 (2 dry shells, 2 wet animals), 2194 m, “Smoke and mirrors” site,
Endeavor Ridge segement, Juan de Fuca Ridge, off Washington (47°56.89'N, 129°5.91'W).
Family Nassariidae Iredale, 1916
caelolineatus, Nassarius (Nassarius) Nesbitt & Pitt, 1986: The Veliger 28(3):294-295.
PARATYPES: LACM 2173 (6 dry), 9-18 m, Bahía Academy, Isla Santa Cruz (= Indefatigable
Id.), Islas Galápagos (0 44.6 S, 90 18.08 W), Leg.: A.G. Smith & J. DeRoy, FEB 1964.
elongata, Nassa mendicus Williamson, 1892: Proc. USNM 15(898):213.
SYNTYPES: LACM 2207 (16 dry), Santa Catalina Id., California Channel Ids., California.
Remarks: Junior synonym of Nassarius mendicus (Gould, 1849).
harryleei, Nassarius Garcia, 2001: Novapex 2(1):25-26, figs. 1-2
HOLOTYPE: LACM 2895 (dry); PARATYPE: LACM 2896 (1 dry), 270-271 m, Golfo de
Chiriquí, Panamá.
limacina, Alectrion Dall, 1917a: Proc. USNM 51(2166):577.
PARATYPES: LACM 1441 (2 dry), 33 m, near La Paz, Golfo de California, Baja California Sur,
México.
Remarks: Currently in the genus Nassarius fide Keen (1971).
leucops, Nassa Pilsbry & Lowe, 1932b: The Nautilus 46(2):51-52.
PARATYPES: LACM 2181 (3 dry), Estuary back of Bahía Kino, Sonora, México, Leg.: H.N.
Lowe.
Remarks: Junior synonym of Nassarius moestus (Hinds, 1844) fide Keen (1971). Ex UCLA
3114.
osculata, Bullia (Bullia) Sowerby, 1900: Proc. Malac. Soc. London 4(1):3.
SYNTYPES [as "COTYPES"]: LACM 1325 (2 dry), Port Alfred (= The Kowie), Cape Prov.,
South Africa.
shaskyi, Nassarius (Nassarius) McLean, 1970a: Malac. Review 2:128-129, fig. 41.
HOLOTYPE: LACM 1405 (dry); PARATYPES: LACM 1406 (2 dry), 37 m, off Isla de
Gorgona, Cauca Dept., Colombia (3 01.42 N, 78 10 W), Leg.: R/V Velero III, 12 FEB 1934.
smitsorum, Nassarius Kool, 1990: Basteria 54:171-173.
PARATYPES: LACM 2470 (2 dry), Manokwari, at the Uriami River mouth, Dore Bay, Irian
Jaya, Papua New Guinea, Leg.: J.H. & D. Smits, 1960.
Family Melongenidae Gill, 1871
sprucecreekensis, Melongena Tucker, 1994: Bull. Florida Mus. Nat. Hist. 36(7):197-202.
PARATYPES: LACM 2033 (10 dry), North of New Smyrna Beach where U.S. Route 1 crosses
Strickland Bay of the Spruce Creek Estuary, Volusia Co., Florida.
Remarks: Junior synonym of Melongena corona (Gmelin, 1791).
Family Fasciolariidae Gray, 1853
allyni, Fusinus (Fusinus) McLean, 1970b: The Veliger 12(3):314-315, pl. 46, figs. 15-17.
HOLOTYPE: LACM 1290 (dry); PARATYPES: LACM 1291 (3 dry), 128-146 m, off Isla
Daphne, Islas Galápagos (0 24.5 S, 22 40 W), Leg.: R/V Velero III, 20 JAN 1938.
Remarks: Holotype published as LACM-AHF 1290, paratypes published as LACM-AHF 1291.
consagensis, Fusinus (Fusinus) Poorman, 1981: The Veliger 23(4):339-340, figs. 1, 6, 10.
HOLOTYPE: LACM 1934 (1 dry), 22 m, 3 km S of Roca Consag, Golfo de California, Baja
California, México (31 07 N, 114 30 W), Leg.: Chamizal Exped., 28 JUN 1968.
humboldti, Fusinus (Fusinus) Poorman, 1981: The Veliger 23(4):340-341, figs 2, 7, 11.
HOLOTYPE: LACM 1935 (dry); PARATYPES: LACM 1943 (2 dry), 400 m, off Isla Pinzon
(Duncan), Islas Galápagos (0 35 S, 90 40 W), Leg.: J. DeRoy, MAR 1979.
magnapex, Fusinus (Fusinus) Poorman, 1981: The Veliger 23(4):341-342, figs. 3, 8, 12.
HOLOTYPE: LACM 1936 (dry); PARATYPES: LACM 1945 (3 dry), 160-220 m, WSW of
Todos Santos, Pacific Coast Baja California Sur, México (23 20 N, 110 22 W), Leg.: J.H.
McLean; R/V Searcher, 1 FEB 1971.
michaelrogersi, Fusinus Goodwin, 2001: Bull. Inst. Malac. Tokyo 8(3):115-117.
PARATYPE: LACM 2899 (dry), 82-229 m, between Laysan Id. and Nihoa Id., NW Hawaiian
Ids., D.R. Goodwin, 1983-1989.
paulus, Fusinus (Fusinus) Poorman, 1981: The Veliger 23(4):342, figs. 4, 13.
HOLOTYPE: LACM 1938 (dry), 150 m, San Jaime Bank, SW of Cabo San Lucas, Pacific Coast
Baja California Sur, México (22 50 N, 10 15 W); PARATYPES: LACM 1939 (4 dry), 137 m,
San Jaime Bank, SW of Cabo San Lucas, Pacific Coast Baja California Sur, México (22 50.5 N,
110 15 W), Leg.: R/V Velero III, 3 MAR 1937.
sonorae, Fusinus (Fusinus) Poorman, 1981: The Veliger 23(4):343, figs. 5, 9.
HOLOTYPE: LACM 1937 (dry),100-120 m, 5 km SE of Punta San Antonio, near Guaymas,
Sonora, México (27 54 N, 111 05 W), Leg.: C. & L. Shy, OCT 1978.
Family Columbellidae Swainson, 1840
aliciae, Aesopus Marincovich, 1973: Nat. Hist. Mus. L.A. Co. Sci. Bull. 16:35, fig. 74.
HOLOTYPE: LACM 1593 (dry), PARATYPES: LACM 1594 (59 dry), 0 m, intertidal, Iquique,
Tarapaca Prov., Chile (20 13 S, 70 10 W), Leg.: L. Marincovich, AUG 1964.
anitae, Nassarina (Zanassarina) Campbell, 1961: The Veliger 4(1):26.
PARATYPE: LACM 1096 (1 dry), 18-46 m, trawled off Cabo Haro, Sonora, México, Leg.: Ariel
Exped., 28 AUG 1960.
aureola, Pyrene Howard, 1963: Santa Barbara Mus. Nat. Hist. Occas. Pap. 7:2-11.
PARATYPES: LACM 1209 (6 dry); PARATYPES: LACM 2155 (2 dry), 0 m, intertidal, Norse
Beach, Punta Peñasco, Sonora, México, Leg.: F.B. Howard, FEB 1957.
Remarks: Junior synonym of Columbella aureomexicana (Howard, 1963) fide Keen (1971).
Paratype LACM 2155 ex UCLA 35317.
berryi, Anachis (Costoanachis) Shasky, 1970: The Veliger 13(2):190, fig. 5.
HOLOTYPE: LACM 1413 (dry); PARATYPES: LACM 1414 (2 dry), 1-3 m, Pulmo Reef, Golfo
de California, Baja California Sur, México (23 26 N, 109 25 W), Leg.: D.R. Shasky, 23-25 APR
1965.
crassiparva, Cotonopsis (Cotonopsis) Jung, 1989: Schweiz. Palaont. Abhand. 111:160-162, figs.
56.2, 208.1-12, 209.1.3.
HOLOTYPE: LACM 2192 (dry); PARATYPES: LACM 2193 (4 dry), 82 m, Caleta Tagus, Isla
Isabela (= Albemarle Id.), Islas Galápagos (0 16.70 S, 91 22.8 W), Leg.: R/V Velero III, 10
DEC 1934; PARATYPES: LACM 2592 (1 dry, 1 gold coated), 146 m, off Caleta Tagus, Isla
Isabela (= Albermarle Id.), Islas Galápagos (0 16.78 S, 91 23.02 W), Leg.: R/V Velero III, 10
DEC 1934; PARATYPES: LACM 2593 (5 dry), 55 m, Caleta Tagus, Isla Isabela (= Albermarle
Id.), Islas Galápagos (0 16.63 S, 91 22.73 W), Leg.: R/V Velero III, 13 JAN 1934;
PARATYPES: LACM 2594 (2 dry), 91-183 m, N of Isla Espanola (= Hood Id.), Islas Galápagos
(1 19.3 S, 89 39.8 W), Leg.: R/V Velero III, 29 JAN 1938; PARATYPES: LACM 2595 (2 dry),
37-73 m, N of Isla Espanola (= Hood Id.), Islas Galápagos (1 21.92 S, 90 40.08 W), Leg.: R/V
Velero III, 28 JAN 1938; PARATYPE: LACM 2596 (1 dry), 55 m, between Isla Daphne & Isla
Seymour, Islas Galápagos (0 24.8 S, 90 21.7 W), Leg.: R/V Velero III, 13 DEC 1934;
PARATYPES: LACM 2597 (2 dry), 91-128 m, between Isla Isabela (= Albermarle Id.) & Isla
San Salvador (= James Id.), Islas Galápagos (0 10.8 S, 90 52.13 W), Leg.: R/V Velero III, 24
JAN 1934; PARATYPES: LACM 2598 (2 dry), 46-64 m, off Bahía Gardner, Isla Espanola
(Hood), Islas Galápagos (1 21.92 S, 90°40.08 W), Leg.: R/V Velero III, 31 JAN 1934;
PARATYPE: LACM 2599 (1 dry), 78 m, NE of Cabo Barrington, Isla Isabela (= Albermarle Id.),
Islas Galápagos (0 36 S, 90 52 W), Leg.: R/V Anton Bruun, 24 SEP 1966; PARATYPES;
LACM 2600 (3 dry), 106-110 m, E of S end Isla Isabela (= Albermarle Id.), Islas Galápagos
(0 55 S, 90 30 W), Leg.: R/V Velero III, 26 JAN 1934; PARATYPES: LACM 2601 (2 dry), 59
m, Bahía Stephen, Isla San Cristobal (= Chatham Id.), Islas Galápagos (0 47 S, 89 31 W), Leg.:
R/V Velero III, 21 JAN 1934; PARATYPES: LACM 2602 (1 dry, 1 gold coated), 91-110 m, off
Caleta Tagus, Isla Isabela (= Albermarle Id.), Islas Galápagos (0 16.8 S, 91 22.87 W), Leg.: R/V
Velero III, 15 JAN 1934.
ecuadoriana, Ruthia Shasky, 1970: The Veliger 13(2):193, figs. 8, 12.
HOLOTYPE: LACM 1417 (dry), 4 m, off Cabo San Francisco, Esmeraldas Prov., Ecuador
(0 39.5 N, 80 06.5 W), Leg.: R/V Velero III, 11 FEB 1934.
fayae, Anachis (Costoanachis) Keen, 1971: Sea Shells of Trop. W. Amer., p. 579.
PARATYPES: LACM 1634 (2 dry), 0 m, intertidal, Punta Colorado, near Guaymas, Sonora,
México, Leg.: F.B. Howard & Gale Sphon, 4 FEB 1962; PARATYPES: LACM 2180 (2 dry),
0 m, intertidal, Playa Caracol, Nuevo Guaymas, Sonora, México, Leg.: F.B. Howard, FEB-MAR
1966.
Remarks: Paratype lot LACM 2180 ex UCLA 48322 & 48323.
hilli, Anachis (Glyptanachis) Pilsbry & Lowe, 1932a: Proc. Acad. Nat. Sci. Philadelphia 84:73.
PARATYPE: LACM 1009 (1 dry); PARATYPE: LACM 2156 (1 dry), Mazatlán, Sinaloa,
México, Leg.: H.N. Lowe, 8 SEP 1931. Paratype 2156 ex UCLA 4641.
jaliscana, Cotonopsis (Cotonopsis) Jung, 1989: Schweiz. Palaont. Abhand. 111:171-173, figs.
56.6, 223.1-9, 224.1-3.
HOLOTYPE: LACM 2196 (dry); PARATYPES: LACM 2197 (5 dry, 2 gold coated), 37-73 m,
Bahía Banderas, Jalisco, México (20 40 N, 105 25 W), Leg.: G. Willett, 14 FEB 1938;
PARATYPE: LACM 2603 (1 dry), Bahía Banderas, 2-5 mi. east of Punta Mita and off La Cruz,
Jalisco, México (20 45 N, 105 25 W), Leg.: J.H. McLean, 20-24 MAR 1965; PARATYPE:
LACM 2604 (1 dry, missing), 46 m, Bahía Tenacatita, Jalisco (19 16 38"N, 104 50 35"W), 4
MAR 1934; PARATYPE: LACM 2605 (1 dry, missing), 37-73 m, Bahía Tenacatita, Jalisco
(19 17 N, 104 50 W), Leg.: G. Willett, 18 FEB 1938; PARATYPE: LACM 2606 (1 dry),
Intertidal, La Cruz de Juanacaxtl, Bahía Banderas, Nayarit, México (20 43.5 N, 105 25 W),
Leg.: H. DuShane, MAR 1974; PARATYPE: LACM 2607 (1 dry), 27-73 m, Bahía Chamela,
Jalisco, México (19 33 N, 105 07 W), Leg.: G. Willett, 14 FEB 1938.
lalage, Mitrella Pilsbry & Lowe, 1932a: Proc. Acad. Nat. Sci. Philadelphia 84:70-71.
PARATYPE: LACM 1955 (1 dry), Mazatlán, Sinaloa, México, Leg.: H.N. Lowe.
lillianae, Anachis (Anachis) Whitney, 1978: The Veliger 20(3):295-296, fig. 1.
HOLOTYPE: LACM 1852 (dry); PARATYPES: LACM 1847 (2 dry), 0 m, intertidal, Playa
Alicia, near San Felipe, Golfo de California, Baja California, México, MAR 1976.
loisae, Mitrilla Pitt & Kohl, 1979: The Veliger 21(4):467-468.
PARATYPES: LACM 1899 (6 dry), N side of Punta Coralillo, Bahía de Caldera, Puntarenas
Prov., Costa Rica (9 54 N, 84 44 W), Leg.: W. & L. Pitt, 29-30 JAN 1975.
macleani, Decipifus Keen, 1971: Sea Shells of Trop. W. Amer., p. 588, fig. 1224.
HOLOTYPE: LACM 1266 (dry); PARATYPES: LACM 1267 (90 dry), Puertecitos, Golfo de
California, Baja California, México, Leg.: J.H. McLean, 1962.
marrae, Columbella Garcia, 1999: Apex 14(3-4):60-64.
PARATYPE: LACM 2870, 24-37 m, Islas Secas, Golfo de Chiriqui, Chiriqui Prov., Panamá.
mazatlanica, Ruthia Shasky, 1970: The Veliger 13(2):192-193.
PARATYPE: LACM 1416 (1 dry), E side of Isla Chivos, Mazatlán, Sinaloa, México, Leg.: R.,
M. & D.R. Shasky, 26 DEC 1962.
mendozana, Cotonopsis (Cotonopsis) Shasky, 1970: The Veliger 13(2):194-195, fig. 10.
HOLOTYPE: LACM 1419 (dry); PARATYPE: LACM 1420 (1 dry), Golfo de Fonseca, La
Union Dept., El Salvador (15 57 N, 95 32 W), Leg.: X. Mendoza, OCT 1960.
paenoblita, Strombina (Recurvina) Jung, 1989: Schweiz. Palaont. Abhand. 111:141-144, figs.
186.1-6.
HOLOTYPE: LACM 2190 (dry); PARATYPES: LACM 2191 (9 dry), 9-37 m, between Caleta
La Cruz & Puerto Pizzaro, off Tumbes, Tumbes Dept., Perú (3 28 S, 80 36 W), Leg.: J.H.
McLean & D.R. Shasky; R/V Maria Elana, 13-14 APR 1972; PARATYPES: LACM 2586 (2
dry), 18 m, Bahía Independencia, Ica Dept., Perú (14 14.7 S, 76 09.12 W), Leg.: R/V Velero III,
9 FEB 1938; PARATYPE: LACM 2587 (1 dry), Nunura, Piura Dept., Perú, 23 MAY 1958;
PARATYPES: LACM 2588 (4 dry), S of Punta Parinas, N of Paita, Piura Dept., Perú (4 48 S,
81 17 W), Leg.: R/V Anton Bruun, 2 JUN 1966; PARATYPES: LACM 2589 (4 dry), 17 m,
Bahía Sechura, Piura Dept., Perú (5 39.5 S, 81 01 W), Leg.: R/V Velero III, 15 FEB 1938;
PARATYPE: LACM 2590 (1 dry), 20 m, SE of Isla de Santa Clara, Golfo de Guayaquail,
Tumbes Dept., Perú (3 15 S, 80 19 W), Leg.: R/V Anton Bruun, SEP 1966; PARATYPE:
LACM 2591 (1 dry), Panama.
perata, Nassarina (Cigclirina) Keen, 1971: Sea Shells of Trop. W. Amer., p. 594-595, fig. 1248.
HOLOTYPE: LACM 1464 (dry), 37-45 m, Puerto Videra, Chiapas, México, Leg.: D.R. Shasky.
radwini, Cotonopsis (Turrina) Jung, 1989: Schweiz. Palaont. Abhand. 111:188, figs. 57.2,
245.1-6.
HOLOTYPE: LACM 2189 (dry), 16-22 m, off the coast of southeasternmost Chiapas, México
(14 42 N, 92 32 W), Leg.: D.R. Shasky; PARATYPES: LACM 2610 (2 dry), 13-20 m, off
Puerto San Jose, Escuintla Dept., Guatemala (13 52.5 N, 91 10.5 W), Leg.: R/V Velero III, 11
JAN 1938.
radwini, Salitra Marincovich, 1973: Nat. Hist. Mus. L.A. Co. Sci. Bull. 16:36-37, figs. 77-78.
HOLOTYPE: LACM 1595 (dry); PARATYPES: LACM 1596 (3 dry), 0 m, intertidal, Iquique,
Tarapaca Prov., Chile (20 13 S, 70 10 W), Leg.: L. Marincovich, AUG 1964.
rapanuiensis, Zafra Raines, 2002a: La Conchiglia 34(304):24-25, fig. 23.
HOLOTYPE: LACM 2942 (dry); PARATYPE: LACM 2943 (1 dry), 10-20 m, off Punta Rosalia,
E of Anakena, Isla de Pascua (= Easter Id.), Leg.: B.K. Raines.
skoglundae, Cotonopsis (Cotonopsis) Jung, 1989: Schweiz. Palaont. Abhand. 111:178-180, fig.
235.8.
PARATYPE: LACM 2195 (1 dry), 122-183 m, off N end of Isla Danzante, Golfo de California,
Baja California Sur, México, Leg.: C. Skoglund.
socorroensis, Columbella Shasky, 1970: The Veliger 13(2):194, fig. 9.
HOLOTYPE: LACM 1418 (dry), 2 m, Bahía Braithwaite, Isla Socorro, Islas Revillagigedo,
México (18 42.7 N, 110 56.8 W), Leg.: D.R. Shasky, 11 AUG 1965.
suteri, Cotonopsis (Cotonopsis) Jung, 1989: Schweiz. Palaont. Abhand. 111:180-182, figs.
236.16-18.
PARATYPES: LACM 2194 (3 dry), ca. 12 mi. SSW of Rancho Inocentes, Pacific Coast Baja
California Sur, México (23 38 N, 110 43 W), Leg.: J.H. McLean; R/V Searcher, 1 FEB 1971;
PARATYPES: LACM 2467 (2 dry), Off Salina, Oaxaca, México; PARATYPE: LACM 2608 (1
dry), Bahía Agua Verde, Golfo de California, Baja California Sur, México, Leg.: Elanor Dwyer,
SPRING 1965; PARATYPE: LACM 2609 (1 dry), 93-101 m, off Punta Gorda, Golfo de
California, Baja California Sur, México (23 03.63 N, 109 34.3 W), Leg.: R/V Velero III, 16 FEB
1940.
taylorae, Zafra Raines, 2002a: La Conchiglia 34(304):25-26, fig. 24.
HOLOTYPE: LACM 2944 (dry); PARATYPE: LACM 2945 (1 dry), 20-30 m, off Hanga Nui,
Isla de Pascua (= Easter Id.), Leg.: B.K. Raines.
tehuantepecensis, Radwinia Shasky, 1970: The Veliger 13(2):191, figs. 6, 11.
HOLOTYPE: LACM 1415 (dry), 30-35 m, between San Simeon & Puerto Madero, Golfo de
Tehuantepec, Chiapas, México, Leg.: Carlos Carballo, Jr., JUL 1961.
Remarks: Currently in the genus/subgenus Nassarina (Radwinia) fide Keen (1971).
Superfamily PSEUDOLIVOIDEA de Gregorio, 1880
Family Pseudolividae de Gregorio, 1880
albus, Fusulculus Bouchet & Vermeij, 1998: Nautilus 111(3):49-51.
PARATYPE: LACM 3062 (1 dry), 668 m, NW of Cook Ref., New Caledonia Group, New
Caledonia (18°42'S, 163°13'S), Leg.: P. Bouchet & B. Richer, R/V Vauban, 15 SEP 1985.
crenatus, Fusulculus Bouchet & Vermeij, 1998: Nautilus 111(3):49.
PARATYPE: LACM 3061 (1 dry), NE of Matthew Id., New Caledonia Outliers, New Caledonia
(22°17'S, 171°18'E), Leg.: B. Richer, R/V Alis, 5 JUN. 1989.
mcleani, Macron Vermeij, 1998: The Nautilus 111(2):72-73, figs. 16-17.
HOLOTYPE: LACM 2812 (dry), 4-5 m, off Punta Willard, Bahía Gonzaga, Golfo de California,
Baja California, México (29 48 N, 114 23.1 W), Leg.: R/V Velero III, 23 MAR 1937.
Superfamily MURICOIDEA Rafinesque, 1815
Family Muricidae Rafinesque, 1815
advenus, Pazinotus Poorman, 1980: The Veliger 22(4):362-363, figs. 3-4.
HOLOTYPE: LACM 1914 (dry), 100m, 5 km S of Tetas de Cabra, Estero San Carlos, Sonora,
México (27 54 N, 111 05 W), Leg.: F. & R. Poorman, NOV 1978.
Remarks: Currently in the genus Poirieria fide Houart (1994).
albospinosus, Trophon (Boreotrophon) Willett, 1931b: The Nautilus 45(2):66, pl. 4, fig. 3.
HOLOTYPE: LACM 1028 (dry), PARATYPES: LACM 2149 (8 dry), 46 m, off the N shore of
Santa Catalina Id., California Channel Ids., California, Leg.: G. Willett, JUL 1920.
Remarks: Currently in the genus Boreotrophon and currently a junior synonym of Boreotrophon
triangularis (Carpenter, 1864). Patratypes ex UCLA 21821.
arcana, Pterotyphis (Tripterotyphis) DuShane, 1969: The Veliger 11(4):344, pl. 54, figs. 4-6.
HOLOTYPE: LACM 1195 (dry), Mazatlán, Sinaloa, México, Leg.: H. DuShane, 24 FEB 1968.
Remarks: Currently in the genus Tripterotyphis fide Radwin & D’Attilio (1976).
bahamondei, Trophon McLean & Andrade, 1982: Nat. Hist. Mus. L.A. Co. Cont. Sci. 342:10.
PARATYPES: LACM 1983 (17 dry), 340 m, off Pichilemu, Colchagua Prov., Chile (34 27 S,
71 54 W), Leg.: H. Andrade, 25 MAY 1976.
carolskoglundae, Typhisopsis Houart & Hertz, 2006. Nautilus 120(2):56-58.
PARATYPES: LACM 3066 (2 dry), 24-30 m, Playas del Coco, Guanacaste Prov., Costa Rica,
Leg.: C. & P. Skoglund, APR 1986.
clarki, Scabrotrophon McLean, 1995: Tax. Atlas Santa Maria Basin 9(2)(1):97-98, figs. 1.19E
HOLOTYPE: LACM 2774 (dry), 210 m, off Monterey, Monterey Co., California.
cocosensis, Favartia Myers & D'Attilio, 1990: Venus 49(4):282-285.
PARATYPES: LACM 2200 (2 dry), 12-24 m, Roca Sucia, Isla del Coco, East Pacific, Leg.: D.R.
Shasky, APR 1987.
eversoni, Phyllonotus (Phyllonotus) D'Attilio, Myers, & Shasky, 1987: The Nautilus
101(4):162-164.
PARATYPE: LACM 2204 (1 dry), 90 m, off Bahía Chatham, Isla del Coco, East Pacific
(5 33 N, 87 03 W), Leg.: K. Kaiser & D.R. Shasky, 26 APR 1987.
fayae, Pterotyphis (Tripterotyphis) Keen & Campbell, 1964: The Veliger 7(1):54-56.
PARATYPE: LACM 1133 (1 dry); PARATYPE: LACM 2086 (1 dry), 0 m, intertidal, Barra de
Navidad, Jalisco, México, Leg.: F.B. Howard & G. Sphon, 7-11 JAN 1962.
floridana, Thais haysae Clench, 1927: The Nautilus 41(1):6-8.
PARATYPE: LACM 1219 (1 dry), Grand Bayou, Mississippi River Delta, Plaquemines Parish,
Louisiana.
galapaganus, “Murex” Emerson & D'Attilio, 1970a: The Veliger 12(3):271-272.
PARATYPE: LACM 1233 (1 dry), 91-183 m, N of Isla Espanola (= Hood Id.), Islas Galápagos
(1 19.2 N, 89 39.7 W), Leg.: R/V Velero III, 29 JAN 1938.
Remarks: Currently in the genus Paziella fide Keen (1971).
grovesi, Scabrotrophon McLean, 1995: Tax. Atlas Santa Maria Basin 9(2)(1):96-97, fig. 1.19D.
PARATYPE: LACM 2781 (1 dry), 79-140 m, NE of Santa Catalina Id., California Channel Ids.,
California (33 21.5 N, 118 21.5 W), Leg.: C.A. Kofoid & W.J. Raymond, 20 JUN 1901.
hazardi, Boreotrophon McLean, 1995: Tax. Atlas Santa Maria Basin 9(2)(1):91, fig. 1.18C
HOLOTYPE: LACM 2783 (dry), 220 m, off Naples Pt., Isla Vista, Ellwood Oil Field, Santa
Barbara Co., California (ca. 34 23 N, 120 00 W), Leg.: R. Hazard, R/V Kildee, JAN 1974.
hubbsi, Trophonopsis Rokop, 1972: The Veliger, 15(1):16.
PARATYPE: LACM 1549 (1 dry), 3599-3676 m, abyssal plain off Patton Escarpment, 225 mi.
W of, Cabo Colonett, Pacific Coast, Baja California, México (31 19.7 N, 119 32.9 W), Leg.:
R/V Melville, 18 DEC 1969.
Remarks: Currently in the genus Abyssotrophon fide Houart, 1994.
jacobsoni, Muricodrupa Emerson & D'Attilio, 1981: The Nautilus 95(2):78-81, fig. 1.
HOLOTYPE: LACM 1798 (dry), Yanu-yanu-i-loma Id., Great Astrolabe Reef, Kadavu Group,
Fiji (18 55 S, 178 35 E), Leg.: Kanter & Williamson, 16 OCT-2 NOV 1971.
kabati, Boreotrophon McLean, 1995: Tax. Atlas Santa Maria Basin 9(2)(1):89-90, fig. 1.18A.
HOLOTYPE: LACM 2782 (dry), 440-488 m, 8 mi. E of Long Pt., Santa Catalina Id., California
Channel Ids., California (33 24.7 N, 118 14.08 W), Leg.: R/V Velero III, 13 SEP 1941.
keenae, Murexiella E.H. Vokes, 1970a: The Veliger 12(3):328, pl. 50, figs. 8-10.
HOLOTYPE: LACM 1259 (dry), Playa Venado, Pacific Coast of Panama Canal Zone, Panamá,
Leg.: Mrs. R. H. Stewart
keenae, Ocenebra Bormann, 1946: The Nautilus 60(2):40-41.
PARATYPES: LACM 1001 (15 dry), Whites Pt., Palos Verdes Peninsula, Los Angeles Co.,
California, Leg.: Mr. & Mrs. R. Bormann, OCT 1946.
Remarks: Junior synonym of Ocnebra beta (Dall, 1919).
kowieensis, Afritrophon Sowerby, 1901: Proc. Malac. Soc. London 4(5):213.
SYNTYPES [as "COTYPES"]: LACM 1326 (2 dry), Port Alfred (= The Kowie), Cape Prov.,
South Africa.
laurae, Murexiella (Murexiella) E.H. Vokes, 1970a: The Veliger 12(3):328-329, pl. 50, figs.
4-5.
HOLOTYPE: LACM 1260 (dry), PARATYPE: LACM 1959 (1 dry), 31 m, Punta de Juluapan,
Manzanillo, Colima, México, Leg.: L. Shy.
Remarks: Junior synonym of Murexiella humilis (Broderip, 1833) fide Radwin & D’Attilio
(1976).
lowei, Typhis Pilsbry, 1931: The Nautilus 45(2):72.
PARATYPE: LACM 1851 (dry), Bahía Montijo, Panama.
Remarks: Currently in the genus Tripterotyphis.
mcleani, Muricopsis (Muricopsis) Wiedrick, 2009: Novapex 10(2):50-51, figs. 1-2, 7-8.
HOLOTYPE: LACM 3097 (dry), Bahia San Gabriel, Isla Espirito Santo, Golfo de California,
México (24 29 N, 110 27 W), Leg.: G. Sphon, 31 MAR 1974.
mildredae, Murexiella Poorman, 1980: The Veliger 22(4):361-362, figs. 1-2.
HOLOTYPE: LACM 1913 (dry), 100 m, 5 km S of Tetas de Cabra, Estero San Carlos, Sonora,
México (27 54 N, 111 05 W), Leg.: F. & R. Poorman, NOV 1978.
Remarks: Currently in the genus/subgenus Muricopsis (Murexsul) fide Houart (1994).
macleani, Phyllonotus (Purpurellus) Emerson & D'Attilio, 1969: The Veliger 12(2):147-148,
pl. 26, figs. 1-4.
HOLOTYPE: LACM 1230 (dry), 46 m, Loreto Channel, Golfo de California, Baja California
Sur, México, Leg.: Ariel Exped., 29 AUG 1960; PARATYPE: LACM 1273 (1 dry), 22 m, Islas
Secas, Chiriqui Prov., Panamá (7 57.7 N, 82 00.7 W), Leg.: R/V Velero III, 5 FEB 1935.
Remarks: Currently in the genus Purpurellus fide Houart (1994).
myrae, Cinclidotyphis DuShane, 1969: The Veliger 11(4):343-344, pl. 54, figs. 1-3.
HOLOTYPE: LACM 1194 (dry), Bahía Tenacatita, Jalisco, México, Leg.: H. DuShane, 25 JAN
1968.
myrakeenae, Aspella (?Dermomurex) Emerson & D'Attilio, 1970b: The Nautilus 83(3):89-95.
PARATYPE: LACM 1237 (1 dry), 0 m, intertidal, Bahía de las Banderas, Nayarit, México
(20 44 N, 105 29 W). Leg.: H. & J. DuShane, 12-18 JAN 1969.
Remarks: Currently in the genus Dermomurex fide Radwin & D’Attilio (1976).
painei, Murex Dall, 1903b: Proc. Biol. Soc. Washington 16:174.
PARATYPE: LACM 1957 (1 dry), 91 m, off Avalon, Santa Catalina Id., California Channel Ids.,
California.
Remarks: Currently in the genus Ocenotrophon fide McLean, 1995.
poormani, Favartia Radwin & D'Attilio, 1976: Murex Shells of the World, p. 231-232.
PARATYPE: LACM 1694 (1 dry), 31 m, off Bahía San Carlos, near Guaymas, Sonora, México,
Leg.: R. & F. Poorman, MAR 1964-MAY 1970.
Remarks: Currently in the subgenus Pygmaepterys fide Houart (1994).
rosamiae, Favartia D'Attilio & Myers, 1985: The Nautilus 99(2/3):58-60.
PARATYPES: LACM 2118 (2 dry), Balut, Davao Prov., Mindanao Id., Philippines.
Remarks: Currently in the genus Murexiella fide Houart (1994).
ruthae, Murex (?Haustellum) E.H. Vokes, 1988: Tulane Stud. Geol. Paleo. 21(1-2):18-21.
PARATYPE: LACM 2274 (1 dry), 27-46 m, channel between Loreto & Isla Carmen, Golfo de
California, Baja California Sur, México, Leg.: Ariel. Exped., 29 AUG 1960.
Remarks: Currently in the genus Haustellem fide Houart (1994). Erroneously published as
LACM 2247.
seftoni, Ocenebra Chace, 1958: Trans. San Diego Soc. Nat. Hist. 12(20):331-332.
PARATYPES: LACM 1960 (2 dry), Isla Guadalupe, Pacific Coast Baja California, México,
17 DEC 1957.
sykesi, Murex Preston, 1904: Jour. Malac. 11(4):76-77.
PARATYPE: LACM 1965 (1 dry), Sri Lanka.
Remarks: Currently in the genus Ocenebra. Considered a “species of uncertain identity” fide
Radwin & D’Attilio (1976).
tyrianthina, Acanthina Berry, 1957: Leaflets in Malacology 1(14):78-79.
PARATYPE: LACM 1007 (1 dry), Caleta Man of War, Bahía Magdalena, Pacific Coast Baja
California Sur, México, Leg.: H.B. Clemens, 10 JAN 1955.
velero, Poirieria (Panamurex) E.H. Vokes, 1970b: Tulane Stud. Geol. Paleo. 8(1/2):47-48, fig.
1.
HOLOTYPE: LACM 1407 (dry); PARATYPES: LACM 1408 (7 dry), 38-40 m, 2 mi. SW of
Cabo de Vela, Peninsula de Guajira, La Guajira Dept., Colombia (12 11 N, 72 11 W), Leg.: R/V
Velero III, 8 APR 1939.
Remarks: Currently in the genus Calotrophon fide Radwin & D’Attilio, 1976.
venustula, Murexiella Poorman, 1983: The Veliger 26(1):5-7.
PARATYPE: LACM 2038 (1 dry), 150-200 m, off the S coast of Isla Santa Cruz (= Indefatigable
Id.), Isla Galápagos (0 47 S, 90 21 'W).
wendita, Haustellotyphis Hertz, 1995: The Veliger 38(2):158-161.
PARATYPES: LACM 2744 (2 dry), 6-15 m, off Playa Tamarindo, Bahia Tamarindo, Guanacaste
Prov., Costa Rica (ca. 11 `42 N, 85 50 W), Leg.: R. Koch, 6 & 8 MAR 1993.
westonensis, Muricopsis Myers & D'Attilio, 1990: Venus 49(4):285-287.
PARATYPE: LACM 2201 (1 dry), 23-27 m, Isla Iglesias (= Cascara Id.), Isla del Coco, East
Pacific, Leg.: D.R. Shasky; PARATYPE: LACM 2229 (1 dry), 34 m, Islas Dos Amigos, Isla del
Coco, East Pacific, Leg.: D.R. Shasky.
zamboi, Murex anatomica J.Q. Burch & R.L. Burch, 1960: Hawaiian Shell News 9(5):1, 7.
PARATYPE: LACM 1695 (1 dry), Baring Pt., Olango Id., Cebu Prov., Philippines.
Remarks: Currently in the genus Homalocantha fide Radwin & D’Attilio (1976).
Family Coralliophilidae Chenu, 1859
macleani, Coralliophila (Coralliophila) Shasky, 1970: The Veliger 13(2):189-190, fig. 4.
HOLOTYPE: LACM 1411 (dry); PARATYPE: LACM 1412 (1 dry), 3-4 m, Bahía Saladita,
Guaymas, Sonora, México, Leg.: D.R. Shasky, DEC 1958-DEC 1959.
rocasuciae, Coralliophila Myers & D'Attilio, 1990: Venus 49(4):287-290.
PARATYPES: LACM 2202 (2 dry), 18-24 m, Roca Sucia, Isla del Coco, East Pacific., Leg.:
D.R. Shasky, APR 1986.
santacruzensis, Latiaxis Emerson & D'Attilio, 1970a: The Veliger 12(3):272-273.
PARATYPE: LACM 1234 (1 dry), 88-134 m, off Isla Santa Fe (= Barrington Id.), Islas
Galápagos (0 43 S, 90 01 W), Leg.: R/V Velero III, 26 JAN 1938.
Remarks: Currently in the genus Babelomurex.
Family Turbinellidae Swainson, 1835
tomicici, Columbarium McLean & Andrade, 1982: Nat. Hist. Mus. L.A. Co. Cont. Sci.
342:10-12.
PARATYPES: LACM 1985 (6 dry), 950 m, W of Junquillar, Antofagasta Prov., Chile (25 00 S,
70 40 W), Leg.: R/V Anton Bruun, 16 AUG 1966.
Family Volutidae Rafinesque, 1815
jaculoides, Alcithoe (Alcithoe) Powell, 1924: Proc. Malac. Soc. London 16(1):108-109.
PARATYPE: LACM 1320 (1 dry), 73-91 m, off Whakatane, South Auckland, North Id., New
Zealand.
Remarks: Junior synonym of Alcithoe arabica (Gmelin, 1791) fide Weaver & DuPont (1970);
considered a valid species fide Poppe & Goto (1992).
leucostoma, Livonia mammilla Maybloom, 1951: Australian Zool. 11(4):283.
PARATYPES: LACM 2376 (2 dry), 183 m, Green Cape, New South Wales, Australia.
Remarks: Junior synonym of Livonia mammilla (Sowerby, 1844) fide Weaver & DuPont
(1970); considered a form of Livonia mammilla (Sowerby, 1844) by Poppe & Goto (1992)..
lutosa, Volutocorbis Koch, 1948: Jour. Conch. 23(1):5-6.
PARATYPE: LACM 1630 (1 dry), 256 m, off Port Nolloth, Cape Prov., South Africa;
PARATYPES: LACM 2115 (2 dry), 73-110 m, Port Nolloth, Cape Prov., South Africa.
Remarks: Currently in the genus Athleta fide Poppe & Goto (1992).
Family Harpidae Bronn, 1849
goodwini, Harpa Rehder, 1993: The Nautilus 106(4):127-129.
PARATYPE: LACM 2701 (1 dry), 82-228 m, between French Frigate Shoals and Maro Reef,
NW Hawaiian Ids. (approx. 23 45 N, 166 10 E).
veleroae, Morum (Cancellomorum) Emerson, 1968: Jour. de Conchyl. 107(1):53-57, pl. 1, figs.
1-5.
HOLOTYPE: LACM 1170 (dry); PARATYPE: LACM 1171 (1 dry), 86 m, Bahía Chatham, Isla
del Coco, East Pacific (5 33.8 N, 86 58.7 W), Leg.: R/V Velero III, 14 JAN 1938;
PARATYPE: LACM 1172 (1 dry), 57-91 m, off Isla Nuez, Isla del Coco, East Pacific (5 34 N,
86 59.3 W), Leg.: R/V Velero III, 13 JAN 1938.
Family Cystiscidae Stimpson, 1865
achenea, Granula Roth & Coan, 1971: Proc. Calif. Acad. Sci., 4th ser., 37(23):578-580.
PARATYPE: LACM 1601 (1 dry), 24 m, Bahía Chatham, Isla del Coco, East Pacific (5 35 N,
87 02 W), Leg.: W.L. Schmitt, 24 APR 1941.
Remarks: Currently in the genus Gibberula.
bandera, Persicula Coan & Roth, 1965: The Veliger 8(2):67-69, pl. 12, figs. 3a-c, 4a-5b.
HOLOTYPE: LACM 1141 (dry); PARATYPES: LACM 1142 (18 dry), Bahía Banderas, Jalisco,
México, Leg.: Mr. & Mrs. C. Willis, 4 APR 1963.
insularum, Granula Roth & Coan, 1971: Proc. Calif. Acad. Sci., 4th ser., 37(23):580-583.
PARATYPE: LACM 1602 (1 dry), Caleta Negro, ca. 1 mi. W of Caleta Tagus, Isla Isabela (=
Albemarle Id.), Islas Galápagos (0 15 S, 91 23 W), Leg.: L.G. Hertlein, 5 JAN 1932.
Remarks: Currently in the genus Gibberula.
Family Marginellidae Fleming, 1828
aletes, Prunum Roth, 1978: Nat. Hist. Mus. L.A. Co. Cont. Sci. 292:8-11, figs. 12-13.
HOLOTYPE: LACM 1779 (dry); PARATYPES: LACM 1780 (2 dry), 3-11 m, in cove between
Isla San Jose and Isla Cocinero, Islas Murcielagos, Guanacaste Prov., Costa Rica (10 51.8 N,
86 55.5 W), Leg.: P.I. LaFollette & others; R/V Searcher, 17 FEB 1972.
amphorale, Prunum de Souza, 1992, Jour. Conch., London 34:239-241.
PARATYPE: LACM 2259 (1 dry), 75 m, off Para State, Brazil (1 55.5 N, 42 45 W), Leg.:
Norte-Nordeste I Oceanographic Comission, 31 OCT 1967.
epacrodonta, Dentimargo Roth, 1978: Nat. Hist. Mus. L.A. Co. Cont. Sci. 292:14-15, fig. 15.
HOLOTYPE: LACM 1783 (dry); PARATYPES: LACM 1784 (2 dry), 55-64 m, off Bahía
Honda, Isla Coiba, Veraguas Prov., Panamá (7 44 N, 81 35 W), Leg.: R/V Velero III, 4 FEB
1935.
gorgonense, Prunum Roth, 1978: Nat. Hist. Mus. L.A. Co. Cont. Sci. 292:2-3, fig. 1.
HOLOTYPE: LACM 1777 (1 dry), 37 m, off Isla de Gorgona, Cauca Dept., Colombia
(3 01.42 N, 78 10 W), Leg.: R/V Velero III, 12 FEB 1934.
innexa, Volvarina Roth, 1978: Nat. Hist. Mus. L.A. Co. Cont. Sci. 292:16-17, fig. 16.
HOLOTYPE: LACM 1785 (1 dry), 9-37 m, Bahía Sullivan, Isla San Salvador (= James Id.),
Galápagos Ids. (0 16.5 S, 90 35.2 W), Leg.: R/V Velero III, 23 JAN 1934.
macleani, Prunum Roth, 1978: Nat. Hist. Mus. L.A. Co. Cont. Sci. 292:4.
PARATYPES: LACM 1778 (7 dry; 2 wet), 93 m, W of Cabo Pasado, Manabi Prov., Ecuador
(0 21 S, 80 41 W), Leg.: R/V Anton Bruun, 12 SEP 1966.
Remarks: One radula on glass slide.
zetetes, Dentimargo Roth, 1978: Nat. Hist. Mus. L.A. Co. Cont. Sci. 292:13-14, fig. 14.
HOLOTYPE: LACM 1781 (dry); PARATYPE: LACM 1782 (1 dry), 1-11 m, Punta Isla to 500
m S of point, Bahía Elena, Guanacaste Prov., Costa Rica (10 56 N, 85 48.92 W), Leg.: P.I.
LaFollette & D. Cadien, R/V Searcher, 13 FEB 1972.
Family Mitridae Swainson, 1829
edithrexae, Subcancilla Sphon, 1976: The Nautilus 90(2):64, fig. 1.
HOLOTYPE: LACM 1735 (dry), 3-6 m, Punta Alfaro, Isla Isabela (= Albemarle Id.), Islas
Galápagos (0 25.3 N, 90 57.1 W), Leg.: Ameripagos Exped., 25 MAR 1971.
Remarks: Currently in the genus Ziba fide Robin & Martin (2004).
phorminx, Mitra Berry, 1960: Leaflets in Malacology 1(26):162-163.
PARATYPES: LACM 1954 (3 dry), 64-82 m, Rio Bolsa, Acapulco, Guerrero, México, Leg.: A.
Luna, MAY 1969.
Remarks: Currently in the genus Ziba fide Robin & Martin (2004).
welkerorum, Subcancilla Whitney, 1977: The Veliger 20(1):52-53, fig. 1.
HOLOTYPE: LACM 1814 (dry), Off Guaymas, Sonora, México, 1974.
Remarks: Junior synonym of Ziba erythrogramma (Tomlin, 1931) fide Robin & Martin (2004).
Family Costellariidae MacDonald, 1860
filiareginae, Vexillum (Pusia) regina J. Cate, 1961: The Veliger 4(2):80-81.
PARATYPE: LACM 1605 (1 dry), Balabac, Balabac Id., Palawan Prov., Philippines.
Remarks: Considered a junior synonym of Vexilium (Vexilium) regina Sowerby, 1828 by
Cernohorsky (1970); considered a junior synonym of V. (V.) citrinum (Gmelin, 1791) by Turner
(2001) and Robin & Martin (2004).
hansenae, Vexillum (Pusia) Cernohorsky, 1973: Rec. Auckland Inst. Mus. 10:138.
PARTYPE: LACM 1698 (1 dry), 90 mi. E of Albany, Western Australia, Leg.: M.P. Marrow, 26
DEC 1971.
jeancateae, Thala Sphon, 1969: The Veliger 12 (1):85-88, pl. 6, fig 4.
HOLOTYPE: LACM 1202 (dry), 91-110 m, off Caleta Tagus, Isla Isabela (= Albemarle Id.),
Islas Galápagos (0 16.7 N, 91 22.87 W), Leg.: R/V Velero III, 15 JAN 1934.
marrowi, Vexillum (Pusia) Cernohorsky, 1973: Rec. Auckland Inst. Mus. 10:140-142.
PARATYPE: LACM 1699 (1 dry), Sorrento Beach, N of Perth, Western Australia, Leg.: M.P.
Marrow, DEC 1971.
Superfamily OLIVOIDEA Latreille, 1825
Family Olividae Latreille, 1825
cymatilis, Olivella (Dactylidia) Berry, 1963b: Leaflets in Malacology 1(23):146.
PARATYPE: LACM 1961 (1 dry), Bahía Magdalena, Pacific Coast Baja California Sur, México,
Leg.: C.R. Orcutt, MAR 1917.
inusta, Olivella (Olivella) Sowerby, 1915: Ann. Mag. Nat. Hist., 8th ser., 16:165.
SYNTYPES [as "COTYPES"]: LACM 1335 (2 dry), Florida.
jeusitarum, Agaronia Lopez, Montoya, & Lopez, 1988: The Veliger 30(3):302-303, figs. 21-23.
HOLOTYPE: LACM 2271 (dry); PARATYPES: LACM 2272 (5 dry), At river mouth, Playa
Poneloya, Leon Dept., Nicaragua.
matthewsi, Ancilla J.Q. Burch & R.L. Burch, 1967: The Nautilus 80(3):81-82.
PARATYPE: LACM 1248 (1 dry), 29 m, off Fortaleza, Ceara State, Brazil, Leg.: H.R.
Matthews, FEB 1966.
murrha, Agaronia Berry, 1953c: Trans. San Diego Soc. Nat. Hist. 11(16):417-419.
PARATYPE: LACM 1105 (1 dry), Corinto, Chinandega Dept., Nicaragua, Leg.: H.N. Lowe,
1931.
Remarks: Junior synonym of Agaronia (Agaronia) griseoalba (von Martens, 1897) fide Sterba
(2003).
nica, Agaronia Lopez, Montoya, & Lopez, 1988: The Veliger 30(3):301-302, fig. 18.
HOLOTYPE: LACM 2269 (dry); PARATYPES: LACM 2270 (3 dry), San Juan del Sur, Rivas
Dept., Nicaragua.
rejecta, Oliva (Oliva) J.Q. Burch & R.L. Burch, 1962: The Nautilus 75(4):166.
PARATYPE: LACM 1006 (1 dry); PARATYPE: LACM 2154 (1 dry), La Paz, Golfo de
California, Baja California Sur, México.
Remarks: Junior synonym of Oliva spicata (Roding, 1798) fide Tursch & Greifender (2001);
fide Sterba (2003). Paratype LACM 2154 ex UCLA 34976.
sphoni, Olivella (Olivella) J.Q. Burch & Campbell, 1963: The Nautilus 76(4):124-125.
PARATYPE: LACM 1124 (1 dry), 12 m, between Islas de Dos Hermanos, Mazatlán, Sinaloa,
México, Leg.: B. Campbell & V. Bohn, 18 MAY 1961.
Superfamily CANCELLARIOIDEA Forbes & Hanley, 1852
Family Cancellariidae Forbes & Hanley, 1851
mercadoi, Scalptia Old, 1968: The Veliger 10(3):286-289, pl. 43, fig. 3a-c.
PARATYPE: LACM 1606 (1 dry), Tayabas Bay, Quezon Prov., Luzon Id., Philippines.
laurettae, Cancellaria (Euclia) Petit & Harasewych, 1998: The Nautilus 112(4):115-116.
PARATYPE: LACM 2815 (1 dry), 270-360 m, off Isla Montuosa, W of Punta Hermosa, Isla de
Coiba, Golfo de Chiriquí, Panamá.
milleri, Trigonostoma (Extractrix) J.Q. Burch, 1949: Min. Conch. Club So. Calif. 94:3-4.
PARATYPES: LACM 1086 (2 dry), Tambor, Puntarenas Prov., Costa Rica, Leg.: H. Miller,
1947.
Superfamily CONOIDEA Fleming, 1822
Family Conidae Fleming, 1822
aratus, Conus orbignyi Kilburn, 1973: Ann. Natal Mus. 21(3):575-576.
PARATYPE: LACM 2071 (1 dry), 274 m, Tongaat-Umhlanga Rocks area, Natal, South Africa.
Remarks: Name preoccupied, renamed C. orbignyi elokismenos Kilburn, 1975, The Nautilus
89(2):50.
belizeanus, Purpuriconus Petuch & Sargent, 2011a: Visaya 3(3):41-42, pl. 2, figs. 2a-3b + 1
unnumbered fig.
HOLOTYPE: LACM 3145 (dry), 15 m, Southeast Cay, Glover’s Atoll, Belizean Ids., Toledo
Dist., Belize.
boschorum, Conus Moolenbeek & Coomans, 1993: Apex 8(1-2):20-21.
PARATYPES: LACM 2726 (2 wet), Umm Rasas (Rusays), E coast of Masirah Id., Oman, Leg.:
R.G. Moolenbeek & H. Dekker, 21 NOV 1991.
fenzani, Dauciconus Petuch & Sargent, 2011a: Visaya 3(3):40, pl. 2, figs. 1a-1b + 1
unnumbered fig.
HOLOTYPE: LACM 3142 (dry), 150 m, NW of Isla Coiba, Golfo de Chiriqui, Veraguas Prov.,
Panama.
fluviamaris, Jaspidiconus Petuch & Sargent, 2011a: Visaya 3(3):45-46, pl. 5, figs. 1a-5b + 1
unnumbered fig.
HOLOTYPE: LACM 3146 (dry), 10 m, second reef off Hillsboro Inlet, Pompano Beach,
Broward Co., Florida, Leg.: T. & P. Honker.
jeffreyi, Calamiconus Petuch & Sargent, 2011a: Visaya 3(3):39, pl. 1, figs. 2a-2b + 1
unnumbered fig.
HOLOTYPE: LACM 3144 (dry), 20 m, Balabac Id., Palawan Prov., Philippines .
kahiko, Conus Kohn, 1980: Jour. Paleo. 54(3):536-540.
PARATYPE: LACM 1915 (1 dry), Waimanalo Formation (Pleistocene), SE side of Ulupa‘u
Head, Mokapu Peninsula, Ko‘olau Poko Dist., O‘ahu, Hawaiian Ids. (21 27 N, 157 45.8 W),
Leg.: W.B. Wells.
Remarks: Fossil species.
kohni, Conus McLean & Nybakken, 1979: The Veliger 22(2):140-142, figs. 4, 24.
HOLOTYPE: LACM 1885 (shell dry, animal wet), 18-37 m, Caleta Tagus, Isla Isabela (=
Albemarle Id.), Islas Galápagos (0 24.5 S, 90 23 W), Leg.: G. Wellington, 15 JAN 1975.
maya, Gradiconus Petuch & Sargent, 2011a: Visaya 3(3):42, pl. 3, figs. 3a-3b + 1 unnumbered
fig.
HOLOTYPE: LACM 3143 (dry): 1 m, Laguna de Yalahua, near Isla Holbox, Quintana Roo,
Mexico.
mazzolii, Gradiconus burryae Petuch & Sargent, 2011a: Visaya 3(4):98-101, pl. 2, figs. 1a-5a
[actually 5b as there are two figure 5a’s on plate] + 1 unnumbered fig.
HOLOTYPE: LACM 3205 (dry); PARATYPES: LACM 3206 (4 dry), 1-1.5 m, northern end of
Little Torch Key, Little Torch-Big Pine Channel, Monroe Co., Florida, Leg.: N.E. Mazzolii
poormani, Conus Berry, 1968: Leaflets in Malacology 1 (25):156-157.
PARATYPES: LACM 1261 (2 dry), 44-48 m, off Moro Colorado, Sonora, México, Leg.: A.
Luna, NOV 1967.
roatanensis, Jaspidiconus Petuch & Sargent, 2011a: Visaya 3(3):44-45, pl. 4, figs. 1ª-5b + 1
unnumbered fig.
HOLOTYPE: LACM 3147 (dry): 10 m, Coxon’s Hole, Isla Roatan, Islas de la Bahia, Honduras.
tethys, Virgiconus Petuch & Sargent, 2011b: Visaya 3(3):38, pl. 1, figs. 1a-1b + 1unnumbered
fig.
HOLOTYPE: LACM 3141 (dry), 30 m, Laminusa Id., Tapul Group, Sulu Archipelago, Sulu
Prov., Philippines.
tortuganus, Gradiconus Petuch & Sargent, 2011a: Visaya 3(3):43-44, pl. 3, figs. 5a-5b + 1
unnumbered fig.
HOLOTYPE: LACM 3209 [erroneously published as LACM 3143] (dry), 3 m, 6 km due south
of Ft. Jefferson, Dry Tortugas, Florida Keys, Monroe Co., Florida.
Family Turridae H. Adams & A. Adams, 1853
abrupta, Borsonella (Borsonella) McLean & Poorman, 1971: The Veliger 14(1):104-105, fig.
35.
HOLOTYPE: LACM 1524 (dry), 457 m, NW of Isla Santa Maria (= Charles Id.), Isla Galápagos
(1 09 S, 90 35 W), Leg.: R/V Velero III, 23 JAN 1938.
academica, Splendrillia McLean & Poorman, 1971: The Veliger 14(1):98-99, fig. 21.
HOLOTYPE: LACM 1500 (dry), 60 m, Bahía Academy, Isla Santa Cruz (= Indefatigable Id.),
Islas Galápagos (0 45 S, 90 48 W), Leg.: A. & J. DeRoy, 15 OCT 1965.
acclivicallis, Euclathurella McLean & Poorman, 1971: The Veliger 14(1):107-108, fig. 42.
HOLOTYPE: LACM 1532 (dry), 75-100 m, Caleta Tagus, Isla Isabela (= Albemarle Id.), Islas
Galápagos (0 16.13 N, 91 22.63 W), Leg.: A. & J. DeRoy, 21 JAN 1968.
acclivicosta, Bellaspira McLean & Poorman, 1970: L.A. Co. Mus. Cont. Sci. 189:6-8, fig. 9.
HOLOTYPE: LACM 1193 (dry), 27-37 m, 1 km S of the E point at entrance to Bahía San
Carlos, Sonora, México (27 56 N, 111 03 W), Leg.: L. & F. Poorman.
ampla, Acmaturris McLean & Poorman, 1971: The Veliger 14(1):108, fig. 43.
HOLOTYPE: LACM 1533 (dry), 64 m, Golfo de Guayaquil, Guayas Prov., Ecuador (3 08 S,
80 49 W), Leg.: R/V Velero III, 7 FEB 1934.
amplinucis, Elaeocyma McLean & Poorman, 1971: The Veliger 14(1):91-92, fig. 5.
HOLOTYPE: LACM 1472 (dry), 29 m, Bahía James, Isla San Salvador (= James Id.), Islas
Galápagos (0 12 S, 90 52 W), Leg.: A. & J. DeRoy, 4 FEB 1969.
andrei, Fusiturricula McLean & Poorman, 1971: The Veliger 14(1):99-100, fig. 24.
HOLOTYPE: LACM 1504 (dry); PARATYPES: LACM 1505 (4 dry), 100-200 m, off the S
coast of Isla Santa Cruz (= Indefatigable Id.), Islas Galápagos (0 47 S, 90 21 W), Leg.: A. & J.
DeRoy, 23-29 JUL 1969.
angulosa, Pyrgocythara McLean & Poorman, 1971: The Veliger 14(1):109-110, fig. 47.
HOLOTYPE: LACM 1539 (dry); PARATYPES: LACM 1540 (8 dry), 0 m, intertidal, Bahía
Guasimas, ca. 25 mi. S of Guaymas, Sonora, México (27 50 N, 111 33 W), Leg.: L.T. Findley &
P.M. Oringer, 10 APR 1968.
arga, Splendrillia McLean & Poorman, 1971: The Veliger 14(1):99.
PARATYPE: LACM 1501 (1 dry), 13 m, Puerto Guatulco, Oaxaca, México (15 44.47 N,
96 07.85 W), Leg.: W. Bebee & T. Crocker, 5 DEC 1937.
arteaga, Mangelia Dall & Bartsch, 1910: Mem. Canadian Dept. Mines 14-N:11.
PARATYPE: LACM 1424 (1 dry), 11-62 m, Barkley Sound, Alberni-Clayoquot Dist. Vancouver
Id., British Columbia, Canada, Leg.: J. Macoun.
Remarks: Currently in the genus Kurtzia fide McLean (1971).
asymmetrica, Cerodrillia McLean & Poorman, 1971: The Veliger 14(1):98, fig. 20.
HOLOTYPE: LACM 1498 (dry); PARATYPES: LACM 1499 (2 dry), 18-33 m, Caleta Tagus,
Isla Isabela (= Albemarle Id.), Islas Galápagos (0 16.13 N, 91 22.63 W), Leg.: R/V Velero III, 15
JAN 1934.
badia, Agladrillia McLean & Poorman, 1971: The Veliger 14(1):94, fig. 11.
HOLOTYPE: LACM 1481 (dry); PARATYPES: LACM 1482 (2 dry), 170-200 m, off the S
coast of Isla Santa Cruz (= Indefatigable Id.), Islas Galápagos (0 47 S, 90 21 W), Leg.: A. & J.
DeRoy, 26-29 JUL 1969.
baileyi, Cymakra McLean & Poorman, 1971: The Veliger 14(1):105-106, fig. 37.
HOLOTYPE: LACM 1527 (dry), 60 m, off Cabo San Lucas, Golfo de California, Baja California
Sur, México (22 56.60 N, 109 47 W), Leg.: R/V Velero III, 18 FEB 1936.
barbarae, Brachycythara Lyons, 1972: The Nautilus 86(1):4.
PARATYPE: LACM 1552 (1 dry), 55 m, off Egmont Key, Hillsborough Co., Florida (27 37 N,
83 58 W), Leg.: Project Hourglass, 11 AUG 1966.
berryi, Drillia McLean & Poorman, 1971: The Veliger 14(1):97-98, fig. 19.
HOLOTYPE: LACM 1496 (dry); PARATYPE: LACM 1497 (1 dry), 48 m, off Tovari, Sonora,
México (27 22 N, 110 20 W), Leg.: A. Luna, DEC 1968.
Remarks: Currently in the genus Clathrodrillia.
bicarinata, Clathrodrillia Shasky, 1961: The Veliger 4(1):21.
PARATYPE: LACM 1013 (1 dry), 73-165 m, between Islas Partida & Espíritu Santo, Baja
California Sur, Golfo de California, México, Leg.: Ariel Exped., 30 AUG 1960.
Remarks: Currently in the genus Compsodrillia fide McLean (1971).
bisaya, Lophiotoma Olivera, 2004: Sci. Diliman 16:10-11.
PARATYPE: LACM 3076 (dry), 45.7-73.2 m, Aligway Id., Zamboanga del Norte Prov.,
Mindanao Id., Philippines.
brasiliana, Borsonia Tippett, 1983: The Nautilus 97(4):136-138.
PARATYPES: LACM 1737 (2 dry), 274 m, 200 mi. NW of Sao Luis, Amapa Federal Terr.,
Brazil (0 18 N, 44 17 W), Leg.: R/V Oregon, 9 MAR 1963.
bratcherae, Splendrillia McLean & Poorman, 1971: The Veliger 14(1):99, fig. 23.
HOLOTYPE: LACM 1502 (dry); PARATYPES: LACM 1503 (2 dry), 37 m, S of Isla Tiburón,
Golfo de California, México (28 43.7 N, 112 17.8 W), Leg.: R/V Velero III, 11 MAR 1936.
chilensis, Ptychosyrinx Berry, 1968: Leaflets in Malacology 1(25):158.
HOLOTYPE: LACM 1912 (dry), 366 m, off Coquimbo, Coquimbo Prov., Chile, 22 JAN 1962.
Remarks: Currently in the genus Gemmula.
chrysochildosa, Zonulispira Shasky, 1971: The Veliger 14(1):69, fig. 4.
HOLOTYPE: LACM 1569 (dry), 0 m, intertidal, Playa Veracruz, Pacific Coast of Panama Canal
Zone, Panamá (8 53 N, 79 36 W), Leg.: D.R. Shasky & J.H. McLean, 8-11 MAR 1970;
PARATYPES: LACM 1570 (20 dry), 0 m, intertidal, Isla Venado, Pacific Coast of Panama
Canal Zone, Panamá (8 53 N, 79 36 W), Leg.: D.R. Shasky & J.H. McLean, 8-11 MAR 1970.
clarionensis, Bellaspira McLean & Poorman, 1970: L.A. Co. Mus. Cont. Sci. 189:9-11, fig. 16.
HOLOTYPE: LACM 1366 (dry), 51-82 m, Bahía Sulphur, Isla Clarión, Islas Revillagigedo,
México, Leg.: R/V Velero III, 16 MAR 1939; PARATYPE: LACM 1367 (1 dry), 59 m, Bahía
Sulphur, Isla Clarión, Islas Revillagigedo, México (18 20 N, 114 44 W), Leg.: R/V Velero III, 5
JAN 1934.
clendenini, Miraclathurella Garcia, 2008: Novapex 9(1):11-12.
PARATYPE: LACM 3087 (1 dry), 63-65 m, Bahía de Campeche, northwest of Campeche,
Campeche, México (20º51'N, 92º21'W), 10 JUN 2005.
coltrorum, Strictispira Tippett, 2006: Malacologia 48(1-2):45-47.
PARATYPE: LACM 3070 (1 dry), 25-30 m, off Escavalda Id., Guarapari, Espirito State, Brazil,
Leg.: A. Bodart, DEC 1993.
contracta, Kylix McLean & Poorman, 1971: The Veliger 14(1):92-93, fig. 7.
HOLOTYPE: LACM 1474 (dry); PARATYPE: LACM 1475 (1 dry), 73-128 m, Bahía Guatulco,
Oaxaca, México (15 43 N, 96 08 W), Leg.: G. Willett, 7 MAR 1938.
coracina, Crassispira (Striospira) McLean & Poorman, 1971: The Veliger 14(1):100-101, fig.
26.
HOLOTYPE: LACM 1508 (dry); PARATYPES: LACM 1509 (4 dry), 0 m, intertidal, Isla
Venado, Pacific Coast of Panama Canal Zone, Panamá (8 53 N, 79 36 W), Leg.: J.H. McLean &
D.R. Shasky, 8-11 MAR 1970.
cunninghamae, Drillia (Drillia) McLean & Poorman, 1971: The Veliger 14(1):95, fig. 14.
HOLOTYPE: LACM 1487 (dry); PARATYPE: LACM 1488 (1 dry), 18-64 m, near Bahía Kino
and Bahía Agiabampo, Sonora, México, Leg.: A. Luna, NOV 1963.
currani, Crassispira (Monilispira) McLean & Poorman, 1971: The Veliger 14(1):101, fig. 27.
HOLOTYPE: LACM 1510 (dry); PARATYPE: LACM 1511 (1 dry), 0 m, intertidal, Sayulita,
Nayarit, México (20 52 N, 105 28 W), Leg.: G. Sphon & N. Currin, 9-23 MAR 1970.
cyrene, Mangelia Dall, 1919a: Proc. USNM 56(2288):62-63.
PARATYPES: LACM 1440 (3 dry), 48 m, off La Paz, Golfo de California, Baja California Sur,
México.
Remarks: Currently in the genus Kurtziella fide McLean (1971).
deroyae, Rimsodaphnella McLean & Poorman, 1971: The Veliger 14(1):111, fig. 50.
HOLOTYPE: LACM 1543 (dry); PARATYPES: LACM 1544 (4 dry), 201 m, Bahía Academy,
Isla Santa Cruz (= Indefatigable Id.), Islas Galápagos (0 45 S, 90 48 W), Leg.: A. & J. DeRoy, 5
DEC 1969.
deroyorum, Buridrillia Emerson & McLean, 1992: The Nautilus 106(1):39-41, figs. 9-12.
PARATYPES: LACM 2464 (7 dry), 365 m, off Isla Pinzon (= Duncan Id.), Islas Galápagos
(0 25 S, 90 43 W), Leg.: A. & J. Deroy, MAR 1979.
Remarks: Erroneously published as LACM 2461. Radula mounted on glass slide.
diegensis, Borsonia (Borsonella) Dall, 1908: Bull. Mus. Comp. Zool. 43(6):275-276.
PARATYPE: LACM (1 dry), 1503 m, off San Diego, San Diego Co., California,
Leg.: R/V Albatross.
dushanae, Thelecythara McLean & Poorman, 1971: The Veliger 14(1):108-109, fig. 44.
HOLOTYPE: LACM 1534 (dry), 31 m, 1 mi. S of E promontory at entrance to Bahía San Carlos,
Guaymas, Sonora, México (27 56 N, 111 05 W), Leg.: F. & R. Poorman, APR 1962.
electra, Cytharella Dall, 1919a: Proc. USNM 56(2288):77.
PARATYPES: LACM 1429 (2 dry), Head of Bahía Concepción, Golfo de California, Baja
California Sur, México, Leg.: P. Bartsch.
Remarks: Currently in the genus Platycythara fide McLean (1971).
elenensis, Kurtzia McLean & Poorman, 1971: The Veliger 14(1):109, fig. 45.
HOLOTYPE: LACM 1535 (dry); PARATYPE: LACM 1536 (1 dry), 64 m, Golfo de Guayaquil,
Guayas Prov., Ecuador (3 08 N, 80 49 W), Leg.: R/V Velero III, 7 FEB 1934.
emersoni, Pyrgocythara Shasky, 1971: The Veliger 14(1):72, fig. 10.
HOLOTYPE: LACM 1579 (dry); PARATYPE: LACM 1580 (1 dry), 0 m, intertidal, Agua de
Chale, Golfo de California, Baja California, México (30 42 N, 114 41 W), Leg.: H. DuShane,
MAY 1962.
firmichorda, Leptadrillia McLean & Poorman, 1971: The Veliger 14(1):93, fig. 9.
HOLOTYPE: LACM 1478 (dry), 22 m, Bahía Cupica, Choco Dept., Colombia (6 39 N,
77 31 W), Leg.: R/V Velero III, 26 JAN 1935.
friedrichbonhofferi, Lophiotoma Olivera, 2004: Sci. Diliman 16:8-10.
PARATYPE: LACM 3075 (dry), Aligway Id., Zamboanga del Norte Prov., Mindanao Id.,
Philippines, FEB 2003.
flucticulus, Agladrillia McLean & Poorman, 1971: The Veliger 14(1):94-95, fig. 12.
HOLOTYPE: LACM 1483 (dry); PARATYPE: LACM 1484 (1 dry), 64 m, off Puerto Madero,
Golfo de Tehuantepec, Chiapas, México (14 24 N, 92 34 W), Leg.: D.R. Shasky, 11 JUL 1963.
galapagana, Borsonella McLean & Poorman, 1971: The Veliger 14(1):105, fig. 36.
HOLOTYPE: LACM 1525 (dry); PARATYPES: LACM 1526 (3 dry), 170-200 m, off the S
coast Isla Santa Cruz (= Indefatigable Id.), Islas Galápagos (0 47 S, 90 21 W), Leg.: A. & J.
DeRoy, 26-27 JUL 1969.
garciacubasi, Pilsbryspira (Pilsbryspira) Shasky, 1971: The Veliger 14(1):69-70, fig. 5.
HOLOTYPE: LACM 1571 (dry); PARATYPES: LACM 1572 (3 dry), 2-15 m, Cuastecomate
Cove, Jalisco, México (19 13.7 N, 104 44.88 W), Leg.: D.R. Shasky, 8-9 AUG 1965.
gemmulifera, Daphnella McLean & Poorman, 1971: The Veliger 14(1):110, fig. 48.
HOLOTYPE: LACM 1541 (dry), 201 m, off the S coast of Isla Santa Cruz (= Indefatigable Id.),
Islas Galápagos (0 47 S, 90 21 W), Leg.: A. & J.DeRoy, 6 DEC 1969.
girardi, Cerodrillia Lyons, 1972: The Nautilus 86(1):3-4.
PARATYPE: LACM 1551 (1 dry), 55 m, off Egmont Key, Hillsborough Co., Florida (27 37 N,
83 58 W), Leg.: Project Hourglass, 11 AUG 1966.
gorgonensis, Agladrillia McLean & Poorman, 1971: The Veliger 14(1):95, fig. 13.
HOLOTYPE: LACM 1485 (dry); PARATYPES: LACM 1486 (2 dry), 73-110 m, N of Isla
Gorgona, Cauca Dept., Colombia (3 01 N, 78 10.92 W), Leg.: R/V Velero III, 24 FEB 1938.
gracilis, Compsodrillia McLean & Poorman, 1971: The Veliger 14(1):103, fig. 31.
HOLOTYPE: LACM 1517 (dry); PARATYPES: LACM 1518 (3 dry), 100-200 m, off the S
coast of Isla Santa Cruz (= Indefatigable Id.), Islas Galápagos (0 47 S, 90 21 W), Leg.: A. & J.
DeRoy, 23-29 JUL 1969.
granata, Cymakra McLean & Poorman, 1971: The Veliger 14(1):106, fig. 38.
HOLOTYPE: LACM 1528 (dry), 37-73 m, off the SW end of Isla Monserrate, Golfo de
California, Baja California Sur, México (25 34 N, 111 05 W), Leg.: Ariel Exped., 1 SEP 1960.
harryleei, Darrylia Garcia, 2008: Novapex 9(1):10-11.
PARATYPE: LACM 3088 (1 dry), 0.2 m, “The Key,” Oakridge, south-central Roatan, Islas de
La Bahia Dept., Honduras. DEC 1992.
hertleini, Doxospira Shasky, 1971: The Veliger 14(1):67-68.
PARATYPE: LACM 1565 (1 dry), 77-112 m, 14 mi. SSE of Punta Judas, Puntarenas Prov.,
Costa Rica (9 19.53 N, 84 29.5 W), Leg.: W. Bebee, 1 MAR 1938.
humboldti, Kurtzia McLean & Poorman, 1971: The Veliger 14(1):109, fig. 46.
HOLOTYPE: LACM 1537 (dry); PARATYPES: LACM 1538 (2 dry), 18-33 m, Caleta Tagus,
Isla Isabela (= Albemarle Id.), Islas Galápagos (0 16.13 S, 91 22.63 W), Leg.: R/V Velero III, 15
JAN 1934.
informa, Kermia McLean & Poorman, 1971: The Veliger 14(1):112, fig. 52.
HOLOTYPE: LACM 1546 (dry); PARATYPES: LACM 1547 (2 dry), 15-18 m, Bahía Cartago,
Isla Isabela (= Albemarle Id.), Islas Galápagos (0 36 S, 90 57.18 W), Leg.: R/V Velero III, 25
JAN 1934.
inornata, Drillia (Drillia) McLean & Poorman, 1971: The Veliger 14(1):95-96, fig. 15.
HOLOTYPE: LACM 1489 (dry); PARATYPES: LACM 1490 (3 dry), 93-99 m, off Puerto
Refugio, Isla Angel de la Guarda, Golfo de California, México (29 33.7 N, 113 30.78 W),
Leg.: R/V Velero III, 29 JAN 1940.
jaliscoensis, Calliclava McLean & Poorman, 1971: The Veliger 14(1):90, fig. 1.
HOLOTYPE: LACM 1465 (dry); PARATYPE: LACM 1466 (1 dry), 37-73 m, Bahía Tenacatita,
Jalisco, México (19 17 N, 104 50 W), Leg.: G. Willett, 18 FEB 1938.
klasmidia, Agathotoma (Vitricythara) Shasky, 1971: The Veliger 14(1):71, fig. 8.
HOLOTYPE: LACM 1576 (dry); PARATYPES: LACM 1577 (3 dry), 16-20 m, ca. 1 mi. off
Bahía Olas Atlas, Mazatlán, Sinaloa, México (23 11.7 N, 106 27 W), Leg.: D.R. Shasky, 23
DEC 1962.
lachrymosa, Carinodrillia McLean & Poorman, 1971: The Veliger 14(1):102-103, fig. 30.
HOLOTYPE: LACM 1515 (dry); PARATYPE: LACM 1516 (1 dry), 5-20 m, S side of Bahía
Cuastecomate, 3 mi. NW of Barra de Navidad, Jalisco, México (19 13.7 N, 104 44.88 W),
Leg.: J.H. McLean, 13-21 OCT 1968.
levicallis, Daphnella Poorman, 1983: The Veliger 26(1):7-8.
PARATYPE: LACM 2039 (1 dry), 80-100 m, 5 km S of Tetas de Cabra, Estero San Carlos,
Sonora, México (27 54 N, 111 05 W).
lucida, Calliclava McLean & Poorman, 1971: The Veliger 14(1):90-91, fig. 2.
HOLOTYPE: LACM 1467 (dry), 18-37 m, off Punta Gorda, Golfo de California, Baja California
Sur, México (123 08 N, 109 35 W), Leg.: J.H. McLean & P.M. Oringer, 5 APR 1966.
Remarks: Incorrect latitude cited by McLean & Poorman; correct latitude is 23º08’N.
maculata, Drillia moesta Williamson, 1905b: Bull. So. Calif. Acad. Sci. 4(8): 122-123.
HOLOTYPE: LACM 1052 (dry), Pt. Fermin, San Pedro, Los Angeles Co., California.
Remarks: Junior synonym of Pseudomelatoma penicillata (Carpenter, 1865) fide McLean
(1971).
maesae, Maesiella McLean & Poorman, 1971: The Veliger 14(1):101-102, fig. 29.
HOLOTYPE: LACM 1513 (dry); PARATYPES: LACM 1514 (3 dry), 9-18 m, first cove N of
Saladita Cove, near Guaymas, Sonora, México (27 53.2 N, 110 58 W), Leg.: J.H. McLean, 29
MAY-1 JUN 1968.
margaritensis, Bellaspira McLean & Poorman, 1970: L.A. Co. Mus. Cont. Sci. 189:5-6, fig. 6.
HOLOTYPE: LACM 1337 (dry); PARATYPES: LACM 1338 (2 dry), 31-38 m, 4-7 mi. N of Isla
de Margarita, Nueva Esparta State, Venezuela (11 08 N, 64 17 W), Leg.: R/V Velero III, 21
APR 1939.
martiae, Crassispira (Dallspira) McLean & Poorman, 1971: The Veliger 14(1):100, fig. 25.
HOLOTYPE: LACM 1506 (dry); PARATYPES: LACM 1507 (6 dry), 0 m, intertidal, Isla
Venado, Pacific Coast of Panama Canal Zone, Panamá (8 53 N, 79 36 W), Leg.: J.H. McLean,
8-11 MAR 1970.
maryae, Clathurella McLean & Poorman, 1971: The Veliger 14(1):106, fig. 39.
HOLOTYPE: LACM 1529 (dry), 31 m, 1 mi. S of E promontory at entrance to Bahía San Carlos,
Guaymas, Sonora, México (27 56 N, 111 05 W), Leg.: F. & R. Poorman, DEC 1963.
melichroa, Elaeocyma McLean & Poorman, 1971: The Veliger 14(1):92, fig. 6.
HOLOTYPE: LACM 1473 (dry), 37-73 m, N of, Isla Espanola (= Hood Id.), Islas Galápagos
(1 21.92 N, 90 40.08 W), Leg.: R/V Velero III, 28 JAN 1938.
mendozana, Miraclathurella Shasky, 1971: The Veliger 14(1):68, fig. 2.
HOLOTYPE: LACM 1566 (dry), 37-73 m, Golfo de Tehuantepec, Oaxaca, México (15 43 N,
96 07 W), Leg.: D.R. Shasky, San Juan Exped., 14 JUL 1963.
myrae, Glyphostoma (Glyphostoma) Shasky, 1971: The Veliger 14(1):70, fig. 6.
HOLOTYPE: LACM 1573 (dry), 46 m, off Isla Jicarita, Veraguas Prov., Panamá (7 12 N,
81 47 W), Leg.: F. Lewis, 1937.
olssoni, Compsodrillia McLean & Poorman, 1972: The Veliger 14(1):103-104, fig. 32.
HOLOTYPE: LACM 1519 (dry); PARATYPES: LACM 1520 (6 dry), 15-18 m, off Bahía Santa
Elena, Guayas Prov., Ecuador (2 08.3 S, 81 00.2 W), Leg.: R/V Velero III, 9 FEB 1934.
opaca, Compsodrillia McLean & Poorman, 1971: The Veliger 14(1):104, fig. 33.
HOLOTYPE: LACM 1521 (dry); PARATYPE: LACM 1522 (1 dry), 139-141 m, Ranger Bank,
off Isla Cedros, Pacific Coast Baja California, México (28 31.02 N, 115 30.52 W), Leg.: R/V
Velero III, 25 FEB 1941.
pamglaoensis, Lophiotoma Olivera, 2004: Sci. Diliman 16:18-210.
PARATYPE: LACM 3078 (dry), 95.1-122.6 m, east of Talaga, Batangas Bay, Batangas Prov.,
Luzon Id., Philippines, 26 JUL 1966
petiti, Fenimorea Tippett, 1995: The Nautilus 109(4):133.
PARATYPE: LACM 2795 (1 dry). 59 m, Gulf of Mexico, W of Crystal River, Citrus Co.,
Florida, MAY 1963.
powelli, Kurtziella (Rubellatoma) Shasky, 1971: The Veliger 14(1):70-71, fig. 7.
HOLOTYPE: LACM 1574 (dry); PARATYPE: LACM 1575 (1 dry), 0 m, intertidal, Agua de
Chale, Golfo de California, Baja California, México (30 42 N, 114 41 W), Leg.: D.R. Shasky,
3-6 MAY 1962.
queenslandica, Lophiotoma indica Olivera, 2004: Sci. Diliman 16:6-8, fig. 1B6.
PARATYPE: LACM 3074 (dry), 167.4-201.3 m, Swain Reef, Queensland, Australia, Leg. H.
Morrison.
quentinensis, Cryptogemma Dall, 1919a: Proc. USNM 56(2238):32.
PARATYPES: LACM 1423 (2 dry), 1503 m, off San Diego, San Diego Co., California.
rectilabrum, Lioglyphostoma McLean & Poorman, 1971: The Veliger 14(1):101, fig. 28.
HOLOTYPE: LACM 1512 (dry), 73-229 m, off Cabo Haro, Guaymas, Sonora, México
(27 50 N, 111 55 W), Leg.: Ariel Exped., 2 SEP 1960.
redferni, Strictispira Tippett, 2006: Malacologia 48(1-2):54-60.
PARATYPE: LACM 3071 (1 dry), lee side of Great Guana Cay, Great Abaco, Bahamas, Leg.: C.
Redfern, 9 JUL 1992.
redondoensis, Pseudomelatoma semiinflata T. Burch, 1938: The Nautilus 52(1):21-22, fig. 2.
HOLOTYPE: LACM 1751 (dry); PARATYPE: LACM 2151 (1 dry), 46 m, Redondo Beach, Los
Angeles Co., California, Leg.: T.A. Burch, 25 NOV 1937; PARATYPES: LACM 1011 (3 dry),
46 m, off Redondo Beach, Los Angeles Co., California, Leg.: T.A. & J.Q. Burch, & Mackenzie
Gordon, 8 MAY 1938; PARATYPE: LACM 2152 (1 dry), 27 m, Redondo Beach, Los Angeles
Co., California, Leg.: T.A. Burch.
Remarks: Junior synonym of Crassispira (Burchia) semiinflata (Grant & Gale, 1931) fide
McLean (1971). Holotype ex AHF 381, paratype LACM 2151 ex UCLA 8181, paratype LACM
2152 ex UCLA 22141.
retusa, Daphnella McLean & Poorman, 1971: The Veliger 14(1):110-111, fig. 49.
HOLOTYPE: LACM 1542 (dry), 37-73 mm, off Loreto, Golfo de California, Baja California
Sur, México (26 02 N, 116 16 W), Leg.: Ariel Exped., 29 AUG 1960.
rhodina, Calliclava McLean & Poorman, 1971: The Veliger 14(1):91, fig. 3.
HOLOTYPE: LACM 1468 (dry); PARATYPE: LACM 1469 (1 dry), Bahía Isla Grande,
Guerrero, México (17 42 N, 101 41 W), Leg.: W.J. Seaholm, 1940.
ricaudae, Elaeocyma Berry, 1969: Leaflets in Malacology 1(26):164-165.
HOLOTYPE: LACM 1911 (dry); PARATYPES: LACM 1445 (2 dry), 44-48 m, near Boca de
Soledad, Pacific Coast Baja California Sur, México, Leg.: A. Luna, 1966.
scobina, Glyphostoma (Glyphostoma) McLean & Poorman, 1971: The Veliger 14(1):107, fig.
41.
HOLOTYPE: LACM 1530 (dry); PARATYPES: LACM 1531 (3 dry), 170-200 m, off the S
coast of Isla Santa Cruz (= Indefatigable Id.), Islas Galápagos (0 47 S, 90 21 W), Leg.: A. & J.
DeRoy, 25 JUL 1965.
scotti, Crockerella McLean, 1995: Tax. Atlas Santa Maria Basin 9(2)(1):145-146, fig. 1.29B
HOLOTYPE: LACM 2775 (dry); PARATYPES: LACM 2776 (8 dry), 100 m, 2.15 mi. 081
degrees T to oil platform "HOLLY," off Santa Barbara, Santa Barbara Co., California
(34 23.01 N, 119 56.94 W), Leg.: R/V Velero IV, 10 NOV 1975.
sculpturata, Bela Dall, 1887: Proc. USNM 9(571):299.
PARATYPES: LACM 1442 (2 dry), Chiachi Ids., Alaska Peninsula, Aleutian Ids. Co., Alaska,
Leg.: W.H. Dall.
Remarks: Currently in the genus Oenopota fide Shimek (1987).
secalis, Agathotoma (Vitricythara) Shasky, 1971: The Veliger 14(1):71-72, fig. 9.
HOLOTYPE: LACM 1578 (dry), 16-22 m, ca. 1 mi. off Bahía Olas Atlas, Mazatlán, Sinaloa,
México (23 11.7 N, 106 27 W), Leg.: D.R. Shasky, 23 DEC 1962.
shaskyi, Philbertia McLean & Poorman, 1971: The Veliger 14(1):111-112, fig. 51.
HOLOTYPE: LACM 1545 (dry), 37-73 m, off the SW end of Isla Monserrate, Golfo de
California, Baja California Sur, México (25 34 N, 111 05 W), Leg.: Ariel Exped., 1 SEP 1960.
sikatunai, Gemmula Olivera, 2005: Sci. Diliman, 17:3-7.
PARATYPE: LACM 3079 (dry), Tayabas Bay, Quezon Prov., Luzon Id., Philippines, 1960.
sinuosa, Drillia (Drillia) McLean & Poorman, 1971: The Veliger 14(1):96-97, fig. 16.
HOLOTYPE: LACM 1491 (dry), 150 m, off S coast of Isla Santa Cruz (= Indefatigable Id.), Islas
Galápagos, Leg.: A. & J. DeRoy, 10 JUN 1968.
Remarks: Name preoccupied, renamed Drillia macleani Tucker, 1992, The Nautilus 106(2):76.
sogodensis, Gemmula Olivera, 2005: Sci. Diliman 17:8-10.
PARATYPE: LACM 3080 (dry), 91.5-183 m, off Tap’ing Ting Chaio, Taiwan, Leg.: F. Steiner,
AUG 1968, PARATYPE: LACM 3081 (2 dry), Kii-suido, Wakayama Pref., Honshu, Japan.
stillmani, Strictispira Shasky, 1971: The Veliger 14(1):68-69, fig. 3.
HOLOTYPE: LACM 1567 (dry); PARATYPES: LACM 1568 (18 dry), 0 m, intertidal, Isla
Venado, Pacific Coast Panama Canal Zone, Panamá (8 53 N, 79 36 W), Leg.: J.H. McLean &
D.R. Shasky, 8-11 MAR 1970.
strigillata, Clathromangelia(?) Pallary, 1904: Jour. de Conchyl. 52:222-223.
PARATYPES: LACM 2226 (2 dry), Sfax, Tunisia.
subtilis, Calliclava McLean & Poorman, 1971: The Veliger 14(1):91, fig. 4.
HOLOTYPE: LACM 1470 (dry); PARATYPES: LACM 1471 (3 dry), 44 m, Isla Jicarita,
Veraguas Prov., Panamá (7 12.5 N, 81 47.08 W), Leg.: R/V Velero III, 20 FEB 1934.
taprurensis, Ginnania Pallary, 1904: Jour. de Conchyl. 52:218.
PARATYPE: LACM 2227 (1 dry), Sfax, Tunisia.
Remarks: Currently in the genus Bella fide Powell (1966).
tayabasensis, Lophiotoma Olivera, 2004: Sci. Diliman 16:11-13.
PARATYPE: LACM 3077 (dry), 54.9-219.6 m, between Mactan Id., Cebu Prov. and Bohol
Prov., Philippines.
tornipila, Veprecula McLean & Poorman, 1971: The Veliger 14(1):112, fig. 53.
HOLOTYPE: LACM 1548 (1 dry), 146-183 m, Caleta Tagus, Isla Isabela (= Albemarle Id.), Islas
Galápagos (0 16 S, 91 57.18 W), Leg.: R/V Velero III, 15 JAN 1934.
tumida, Drillia (Drillia) McLean & Poorman, 1971: The Veliger 14(1):97, fig. 17.
HOLOTYPE: LACM 1492 (dry); PARATYPES: LACM 1493 (24 dry), 37-73 m, Bahía de las
Banderas, Jalisco, México (20 43 N, 105 25 W), Leg.: G. Willett, 14 FEB 1938.
undatichorda, Compsodrillia McLean & Poorman, 1971: The Veliger 14(1):104-105, fig. 34.
HOLOTYPE: LACM 1523 (dry), 91-128 m, between Albany & Isla San Salvador (= James Id.),
Islas Galápagos (0 10.7 S, 90 52.13 W), Leg.: R/V Velero III, 24 JAN 1934.
valida, Drillia (Drillia) McLean & Poorman, 1971: The Veliger 14(1):97, fig. 18a-b.
HOLOTYPE: LACM 1494 (dry), 90 m, 8.5 mi. S of Dewey Channel, Pacific Coast Baja
California Sur, México (27 42 N, 115 05.13 W), Leg.: R/V Velero III, 27 FEB 1941;
PARATYPE: LACM 1495 (1 dry), 137 m, San Jaime Bank, off Cabo San Lucas, Pacific Coast
Baja California Sur, México (22 50.5 N, 110 15 W), Leg.: R/V Velero III, 3 MAR 1937.
vitrea, Syntomodrillia McLean & Poorman, 1971: The Veliger 14(1);93-94, fig. 10.
HOLOTYPE: LACM 1479 (dry); PARATYPE: LACM 1480 (1 dry), 4-9 m, Bahía Taboga, Isla
Taboga, Panamá Prov., Panamá (8 47 N, 79 33 W), 24 JUL 1951.
wolfei, Drillia (Drillia) Tippett, 1995: The Nautilus 109(4):127, 129.
PARATYPE: LACM 2794 (1 dry), 36 m, 15 km E of Cape Lookout, Carteret Co., North
Carolina, Leg.: D. Wolfe, MAR-MAY 1971.
woodringi, Kylix McLean & Poorman, 1971: The Veliger 14(1):93, fig. 8.
HOLOTYPE: LACM 1476 (dry); PARATYPES: LACM 1477 (3 dry), 44 m, Isla Jicarita,
Veraguas Prov., Panamá (7 12.5 N, 81 47.08 W), Leg.: R/V Velero III, 20 FEB 1934.
Family Terebridae Mörch, 1852
albocancellata, Terebra Bratcher, 1988: The Veliger 30(4):413, 415.
PARATYPE: LACM 2829 (1 dry), Chesterfield Ids., New Caledonia Outliers, New Caledonia
(36 42 S, 158 59 E); PARATYPES: LACM 2830 (3 dry), 15.5 m, off Sogod, Cebu Prov.,
Philippines.
alboflava, Hastula Bratcher, 1988: The Veliger 30(4):415-416, figs. 2, 5.
HOLOTYPE: LACM 2262 (dry); PARATYPES: LACM 2830 (3 dry), 15.5 m, off Sogod, Cebu
Prov., Philippines.
allyni, Terebra Bratcher & R.D. Burch, 1970a: The Veliger 12(3):298-299.
PARATYPE: LACM 1254 (2 dry), 9-18 m, E shore of Isla Maria Madre, Islas Tres Marias,
Nayarit, México (21 35 N, 106 26 W).
apicitincta, Hastula Sowerby, 1900: Proc. Malac. Soc. London 4(1):5.
SYNTYPES [as "COTYPES"]: LACM 1324 (2 dry), Port Alfred (= The Kowie), Cape Prov.,
South Africa.
baileyi, Duplicaria Bratcher & Cernohorsky, 1982: The Nautilus 96(2):64-65, figs, 5, 11.
HOLOTYPE: LACM 1970 (dry); PARATYPE: LACM 2824 (1 dry), 20 m, Guadalcanal Id.,
Solomon Ids. (9 25 S, 159 56 E), Leg.: B. Bailey; PARATYPE: LACM 2823 (1 dry),
Balut Sarangani Ids., Davao Prov., Mindanao Id., Philippines.
betsyae, Hastula R.D. Burch, 1965: The Veliger 7(4):243.
PARATYPES: LACM 1214 (12 dry), 6 m, Honokohau, Lahaina Dist., Maui, Hawaiian Ids.,
Leg.: Alan Tiedman, DEC 1965.
Remarks: Junior synonym of Hastula penicillata (Hinds, 1844) fide Bratcher & Cernohorsky
(1987).
boucheti, Terebra Bratcher, 1981: The Veliger 23(4):329-330.
PARATYPES: LACM 1364 (2 dry), 67-70 m, Batangas Prov., Luzon Id., Philippines (14 16 N,
120 31 E), Leg.: Musorstrum Exped.
brandi, Terebra Bratcher & R.D. Burch, 1970a: The Veliger 12(3):296-297, pl. 44, figs. 5-6.
HOLOTYPE: LACM 1252 (dry); PARATYPE: LACM 1253 (1 dry), 9-18 m, Bahía Petatlan,
Guerrero, México (17 31.7 N, 101 27.57 W), Leg.: R/V Velero III, 3 MAR 1934.
burchi, Terebra Bratcher & Cernohorsky, 1982: The Nautilus 96(2):63.
PARATYPES: LACM 1971 (2 dry), 15-20 m, northern Mozambique Channel, Archpelago de
Comoros (12 45 S, 45 18 E), Leg.: Benthedi Exped.
caddeyi, Terebra Bratcher & Cernohorsky, 1982: The Nautilus 96(2):63, figs, 4, 12.
HOLOTYPE: LACM 1967 (dry); PARATYPE: LACM 2820 (1 dry), 3 m, Korere Village,
Rabaul, New Britain, Bismarck Arch., Papua New Guinea, Leg.: P.T. Caddey.
campbelli, Terebra (Strioterebra) R.D. Burch, 1965: The Veliger 7(4):247-248.
PARATYPE: LACM 1461 (1 dry), 128 m, off Tosa, Tosa-wan, Kochi Pref., Shikoku, Japan,
1963.
carolae, Terebra Bratcher, 1979a: The Veliger 22(1):63-64, figs. 5, 7.
HOLOTYPE: LACM 1178 (dry); PARATYPES: LACM 1184 (12 dry), 54 m, off Punta Hughes,
Bahía Santa Maria, Pacific Coast Baja California Sur, México (24 45.08 N, 112 19 W), Leg.:
R/V Velero III, 7 MAR 1934; PARATYPES: LACM 2816 (9 dry), Laguna San Ignacio, Pacific
Coast Baja California Sur, México, Leg.: T. Bratcher; PARATYPES: LACM 2817 (7 dry), 36-40
m, Bahía San Bartolome (= Turtle Bay), Pacific Coast Baja California Sur, México, Leg.: D.
Brown; PARATYPES: LACM 2818 (5 dry), 15 m, Bahía Santa Maria, Pacific Coast Baja
California Sur, México, Leg.: D. Brown, 1973; PRATYPE: LACM 2995 (1 dry),14.6 m, off Isla
Cedros, Pacific Coast Baja California Sur, México.
cernohorskyi, Terebra (Punctoterebra) R.D. Burch, 1965: The Veliger 7(4):244-245.
PARATYPE: LACM 1462 (1 dry), Natadola Id., Viti Levu Group, Fiji (18 06.6 S, 19 30 E),
Leg.: A. Morse, 1959.
colorata, Terebra Bratcher, 1988: The Veliger 30(4):416.
PARATYPES: LACM 2263 (9 dry), Lighthouse Beach, Augusta, Western Australia (34 20 S,
115 10 E).
crakei, Duplicaria R.D. Burch, 1965: The Veliger 7(4):245-246.
PARATYPES: LACM 1212 (32 dry), Cable Beach, Broome, Western Australia, Leg.: T. Crake;
PARATYPE: LACM 2073 (1 dry), Broome, Western Australia, Leg.: T. Crake; PARATYPE:
LACM 2996 (1 dry), Cable Beach, Broome, Western Australia, Leg.: T. Crake.
danai, Terebra Berry, 1958b: Leaflets in Malacology 1(16):96-97.
PARATYPE: LACM 1463 (1 dry), 0 m, intertidal, San Pedro, Los Angeles Co., California,
Leg.: S.S. Berry, MAY 1959.
Remarks: Junior synonym of Terebra hemphilli Vanatta, 1924 fide Bratcher & Cernohorsky
(1987).
dedonderi, Terebra Terryn, 2003: Gloria Maris 42(4-5):67-69.
PARATYPE: LACM 3031 (1 dry), 36-50 m, Hansa Bay, Madang Prov., Papua New Guinea.
dorothyae, Terebra Bratcher & R.D. Burch, 1970a: The Veliger 12(3):297, pl. 44, figs. 7-8.
HOLOTYPE: LACM 1250 (dry); PARATYPES: LACM 1251 (11 dry), 13-20 m, off Puerto de
San Jose, Escuintla Dept., Guatemala (13 52.58 N, 91 10.58 W), Leg.: R/V Velero III, 11 JAN
1938.
Remarks: Junior synonym of Terebra glauca Hinds, 1844 fide Bratcher & Cernohorsky (1987).
dushaneae, Terebra Campbell, 1964: The Veliger 6(3):135-136.
PARATYPES: LACM 1208 (4 dry), Puertecitos, Golfo de California, Baja California, México,
Leg.: Helen DuShane, APR 1963.
Remarks: Junior synonym of Terebra bridgesi Dall, 1908 fide Bratcher & Cernohorsky (1987).
elliscrossi, Terebra Bratcher, 1979b: The Veliger 22(1):65-66, fig. 1.
HOLOTYPE: LACM 1257 (dry), 300 m, Honolulu side of Pearl Harbor, Honolulu, Honolulu
Dist., O‘ahu, Hawaiian Ids., Leg.: E. Cross, 10 MAY 1960; PARATYPES: LACM 2819 (2 dry),
21-37.5 m, Guadalcanal Id., Solomon Ids., Leg.: B. Bailey.
fitchi, Terebra (Strioterebrum) Berry, 1958a: Leaflets in Malacology 1(15):89.
PARATYPE: LACM 2831, Entrance to Bahía Santa Maria, Pacific Coast Baja California Sur,
México, Leg.: J.E. Fitch, 4 JUN 1955.
Remarks: Junior synonym of Terebra tiarella Deshayes, 1857 fide Bratcher & Cernorhorsky
(1987).
hancocki, Terebra Bratcher & R.D. Burch, 1970a: The Veliger 12(3):299, pl. 44, fig. 11.
HOLOTYPE: LACM 1255 (dry); PARATYPES: LACM 1256 (2 dry), 15-18 m, off La Libertad,
Bahía de Santa Elena, Guayas Prov., Ecuador (2 08.3 N, 81 00.2 W), Leg.: R/V Velero III, 9
FEB 1934.
hemphilli, Terebra Vanatta, 1924: Proc. Acad. Nat. Sci. Philadelphia 76:425.
PARATYPES: LACM 2832 (2 dry), Laguna Ojo de Liebre (= Scammon's Lagoon), Pacific Coast
Baja California Sur, México, Leg.: H. Hemphill.
hertleini, Terebra Bratcher & R.D. Burch, 1970b: L.A. Co. Mus. Cont. Sci. 188:1-2.
PARATYPE: LACM 1288 (1, dry), 27-46 m, Bahía Academy, Isla Santa Cruz (Indefatigable),
Islas Galápagos (0 46.27 S, 90 19.63 W), Leg.: R/V Velero III, 20 JAN 1934.
Remarks: Junior synonym of Terebra jacquelinae Bratcher & R.D. Burch, 1970 fide Bratcher &
Cernohorsky (1987).
hoffmeyeri, Terebra (Strioterebra) Abbott, 1952: The Nautilus 65(3):78-79.
PARATYPES: LACM 1457 (5 dry); PARATYPES: LACM 1697 (4 dry), Pasay Beach, Manila
Bay, Rizal Prov., Luzon Id., Philippines, Leg.: H. Hoffmeyer, MAY-AUG 1939.
Remarks: Junior synonym of Terebra plumbea Quoy & Gaimard, 1833 fide Bratcher &
Cernohorsky (1987).
insalli, Terebra Bratcher & Burch, 1976: The Veliger 10(1):7
PARATYPE: LACM 2814 (1 dry), Eilat, Gulf of Aqaba, Southern Dist., Israel (29 32.3 N,
34 57.0 E), Leg.: D.C. Insall.
jacquelinae, Terebra Bratcher & R.D. Burch, 1970b: L. A. Co. Mus. Cont. Sci. 188:2-5.
PARATYPES: LACM 1179 (28 dry), 18-46 m, Bahía Academy, Isla Santa Cruz (= Indefatigable
Id.), Islas Galápagos (0 46.27 S, 90 19.63 W), Leg.: R/V Velero III, 24 JAN 1938.
jenningsi, Terebra (Triplostephanus) R.D. Burch, 1965: The Veliger 7(4):248-249.
PARATYPE: LACM 1454 (1 dry), Nadi, Viti Levu, Viti Levu Group, Fiji, Leg.: A. Jennings,
1963; PARATYPES: LACM 1455 (2 dry), 6 m, Natadola Id., Viti Levu, Viti Levu Group, Fiji,
Leg.: A. Jennings, MAR 1962.
kilburni, Terebra (Decorihastula) R.D. Burch, 1965: The Veliger 7(4):249-250.
PARATYPES: LACM 1458 (5 dry), 0.5-1.0 m, Wadingi Id. (= Wading Id.), Mamanuca Group,
Fiji, Leg.: A. Jennings, JUL 1962; PARATYPES: LACM 1459 (2 dry), 1.8 m, Lomalagi,
Mamanuca Group, Fiji, Leg.: W.O. Cernohorsky, DEC 1963; PARATYPES: LACM 1460 (4
dry), Madang, Madang Prov., Papua New Guinea, Leg.: I. Pert, MAR 1952.
knudseni, Terebra Bratcher, 1983: Steenstrupia 8(15):301-305.
PARATYPES: LACM 1363 (3 dry), Malawali Id., Kudat District, Sabah, Borneo, Malaysia,
Leg.: M. Saul, 1962.
lillianae , Terebra Whitney, 1976: The Veliger 18(4):381-382, figs. 1, 4.
HOLOTYPE: LACM 1724 (dry), Off Mossuril, Nampula Prov., Mozambique (14 55 S,
40 41 E), 1974.
Remarks: Junior synonym of Terebra mamillata Watson, 1886 fide Bratcher & Cernohorsky
(1987).
mactanensis, Terebra Bratcher & Cernohorsky, 1982: The Nautilus 96(2):63-64, figs. 1, 10.
HOLOTYPE: LACM 1968 (dry); PARATYPE: LACM 2821 (1 dry), 200 m, Punta Egano,
Mactan Id., Cebu Prov., Philippines.
marrowae, Terebra Bratcher & Cernohorsky, 1982: The Nautilus 96(2):64, figs. 2, 9.
HOLOTYPE: LACM 1969 (dry), 0 m, intertidal, Cleaverville, Western Australia, Leg.: L.
Marrow; PARATYPES: LACM 2822 (2 dry), MonteBello Ids., Western Australia.
maryleeae, Hastula (Hastula) R.D. Burch, 1965: The Veliger 7(4):242.
PARATYPES: LACM 1456 (14 dry), Surfside Beach, Freeport, Brazoria Co., Texas,
Leg.: M. Burch, 16 MAR 1961.
murgrideae, Terebra Garcia, 1999: Apex 14(3-4):64.
PARATYPE: LACM 2871 (1 dry), 18 m, Gulf of México, SE of Biloxi, Harrison Co.,
Mississippi (29°53.25'N, 88°40.46'W).
ninfae, Terebra (Strioterebra) Campbell, 1961: The Veliger 4(1):27-28.
PARATYPE: LACM 1106 (1 dry), 27-37 m, trawled, between Puerto Madero and 30 mi. N of
Guatemala border, Chiapas, México (14 55 N, 92 15 W), Leg.: X. Mendoza, JAN 1961.
onslowensis, Strioterebra Petuch, 1974: The Veliger 17(2):205-206
PARATYPES: LACM 2833 (2 dry), 21.6 m, Onslow Bay, 20 km SSE of Bogue Inlet, Carteret
Co., North Carolina (34 25 N, 77 00 W), Leg.: Duke University, R/V Eastward.
Remarks: Junior synonym of Terebra dislocata Say, 1822 fide Bratcher & Cernohorsky (1987).
parkinsoni, Terebra Cernohorsky & Bratcher, 1976: Rec. Auckland Inst. Mus. 13:137, 139
PARATYPE: LACM 2834 (1 dry), 24 m, Nordup, New Britain, Bismark Archipelago, Papua
New Guinea, Leg.: B. Parkinson, 17 SEP 1975.
paucincisa, Terebra Bratcher, 1988: The Veliger 30(4):413.
PARATYPES: LACM 2260 (2 dry), 17 m, Grand Recif South, New Caledonia Group, New
Caledonia (22 37 S, 166 01 E); PARATYPES: LACM 2828 (2 dry), Tailevu, Fiji, Leg.: B.
Parkinson.
poppei,Terebra Terryn, 2003: Gloria Maris 42(4-5):69-71.
PARATYPE: LACM 3032 (1 dry), 240 m, Aliguay, Bohol Prov. Philippines.
pseudopertusa, Terebra Bratcher & Cernohorsky, 1985: The Nautilus 99(4):91.
PARATYPE: LACM 2835 (1 dry), 80-83 m, Reunion, Mascarene Ids., Indian Ocean (21 06 S,
55 01 E), Leg.: R/V Marion-Dufresne, 1982.
purdyae, Terebra Bratcher & R.D. Burch, 1970b: L.A. Co. Mus. Cont. Sci. 188:5, figs. 5-6.
HOLOTYPE: LACM 1182 (dry); PARATYPE: LACM 1183 (1 dry), 128-146 m, N of Isla Santa
Maria (= Charles Id.), Islas Galápagos (0 59 S, 90 25 W), Leg.: R/V Velero III, 29 JAN 1934.
Remarks: Junior synonym of Terebra shyana Bratcher & Burch, 1970 fide Bratcher &
Cernohorsky (1987).
rancheria, Terebra Bratcher, 1988: The Veliger 30(4):413, figs. 6, 8.
HOLOTYPE: LACM 2261 (dry); PARATYPES: LACM 2827 (3 dry), 3.5 m, Isla Rancheria,
Golfo de Chiriqui, Chiriqui Prov., Panamá (7 38 N, 81 44 W).
reunionensis, Terebra Bratcher & Cernohorsky, 1985: The Nautilus 99(4):93.
PARATYPE: LACM 2836 (1 dry), 210 m, Reunion, Mascarene Ids., Indian Ocean (21 05 S,
55 12 E), Leg.: J. Drivas, 1982.
riosi, Terebra Bratcher & Cernohorsky, 1985: The Nautilus 99(4):93-94
HOLOTYPE: LACM 1974 (missing); PARATYPE: LACM 2837 (1 dry), Intertidal, Buzios,
Cabo Frio, Rio de Janeiro State, Brazil, Leg.: E. Rios & L. Tostes.
shyana, Terebra Bratcher & R.D. Burch, 1970a: The Veliger 12(3):295-296, pl. 44, figs. 9-10.
HOLOTYPE: LACM 1249 (dry); PARATYPES: LACM 2813 (2 dry), 31-73 m, Manzanillo,
Colima, México (19 02 N, 104 21 W), Leg.: L. Shy & J. Frisbey.
stohleri, Terebra Bratcher & R.D. Burch, 1970a: L.A. Co. Mus. Cont. Sci. 188:5-6, figs. 7-8.
HOLOTYPE: LACM 1180 (dry), 18-37 m, Bahía Braithwaite, Isla Socorro, Islas Revillagigedo,
México (18 42.3 N, 110 56.27 W), Leg.: R/V Velero III, 18 MAR 1939; PARATYPE: LACM
1181 (1 dry), 7-18 m, off Cabo Rule, Isla Socorro, Islas Revillagigedo, México (18 42.2 N,
110 57.08 W), Leg.: R/V Velero III, 8 JUN 1934.
swobodai, Terebra Bratcher, 1981: The Veliger 23(4):330-331, figs. 5-6.
HOLOTYPE: LACM 1185 (dry); PARATYPE: LACM 2825 (1 dry), Zamboanga, Zamboanga
del Sur Prov., Mindanao Id., Philippines.
troendlei, Terebra Bratcher, 1981: The Veliger 23(4):330.
PARATYPE: LACM 1365 (6 dry), 10 m, Tahuata, Marquesas Ids., French Polynesia.
turschi, Terebra Bratcher, 1981: The Veliger 23(4):331, figs. 7-8.
HOLOTYPE: LACM 1191 (dry); PARATYPES: LACM 2826 (4 dry), 36 m, Hansa Bay, Papua
New Guinea (4 06 S, 144 22 E).
Clade Heterobranchia
Informal Group “Lower Heterobranchia”
Family Orbitestellidae Iredale, 1917
aequicostata, Orbitestellidae, Raines, 2002a: La Conchiglia 34(304):17, fig. 11.
HOLOTYPE: LACM 2936 (dry); PARATYPE: LACM 2937 (1 dry), 30-50 m, of Tahai, Isla de
Pascua (= Easter Id.), Leg.: B.K. Raines.
Superfamily ARCHITECTONICOIDEA Gray, 1850
Family Architectonicidae Gray, 1850
consobrina, Architectonica Bieler, 1993: Abh. Naturwiss. Ver. Hamburg (NF) 30:95-97, fig. 74.
HOLOTYPE: LACM 2742 (dry); PARATYPE: LACM 2743 (1 dry), 259-293 m, S of Gaspar
Id., Marinduque Prov., Philippines.
Remarks: Erroneously published as LACM 2279 (holotype) & 2280 (Paratype).
orba, Solatisonax? Bieler, 1993: Ahb. Naturwiss. Ver. Hamburg (NF) 30:181-183, fig. 151.
HOLOTYPE: LACM 2471 (dry), 108-143 m, Inner Gorda Bank, Golfo de California, Baja
California Sur, México (23 02.45 N, 109 30.70 W), Leg.: R/V Velero III, 17 FEB 1940;
PARATYPE: LACM 2472 (1 dry), intertidal, W beach of Bahía Concepcion, Golfo de
Califorina, Baja California Sur, México (26 44.7 N, 110 54 W), Leg.: R/V Velero III, 14 MAR
1936.
propinqua, Solatisonax Bieler, 1993: Abh. Naturwiss. Ver. Hamburg (NF) 30:176-178.
PARATYPE: LACM 2473 (1 dry), 37-73 m, Bahía Banderas, Sonora, México (20 40 N,
105 25 W), Leg.: G. Willett, 14 FEB 1938; PARATYPE: LACM 2474 (1 dry), 55 m, off Isla del
Cano, Puntarenas Prov., Costa Rica (8 45 N, 83 54 W), Leg.: J.H. McLean, W. Bussing, R/V
Searcher, 16-17 MAR 1972.
turritus, Heliacus Bieler, 1987: Arch Moll. 117(4/6):204-206.
PARATYPE: LACM 2116 (1 dry), 46 m, Bolo Pt., 0.5 km ESE of Zampa-misaki, Okinawa-jima,
Ryukyu-shoto, Japan (26 26.1 N, 127 42.5 E), Leg.: R.F. Bolland, 25 AUG 1978.
Superfamily RISSOELLOIDEA Gray, 1850
Family Rissoellidae Gray, 1850
peruviana, Rissoella (Rissoella) Ponder & Worsfold, 1994: Nat. Hist. Mus. L.A. Co. Contrib.
Sci. 446:49-51, fig. 31C.
HOLOTYPE: LACM 2698 (dry); PARATYPES: LACM 2699 (21 dry), 2-10 m, Isla Lobos de
Afuera (NW & NE of isthmus), Piura Prov., Perú (6 57.1 S, 80 42.3 W), Leg.: J.H. McLean,
J.A. Coyer, & J.M. Engle, 19-20 JAN 1974.
Superfamily PYRAMIDELLOIDEA Gray, 1840
Family Pyramidellidae Gray, 1840
amandi, Turbonilla (Pyrgiscus) Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):
202-203.
PARATYPE: LACM 1345 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
azteca, Odostomia (Miralda) Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):
207-208.
PARATYPE: LACM 1348 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
Remarks: Currently in the genus Miralda.
bartonella, Turbonilla (Pyrgiscus) Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):
203.
PARATYPE: LACM 1346 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
burchi, Turbonilla (Pyrgiscus) Gordon, 1938: The Nautilus 52(2):49-51, pl. 4, figs. 3-5.
HOLOTYPE: LACM 1057 (dry), 46 m, 1 mi. W of pier at Redondo Beach, Los Angeles Co.,
California, Leg.: M. Gordon & J.Q. Burch, JUN 1938.
canadensis, Turbonilla (Strioturbonilla) Bartsch, 1917: Proc. USNM 52(2193):640.
PARATYPES: LACM 2166 (7 dry), Forrester Id., Prince of Wales Co., Alaska, Leg.: G. Willett.
Remarks: Ex UCLA 21047.
cayucosensis, Turbonilla (Strioturbonilla) Willett, 1929b: The Nautilus 43(1):26-27, figs. 2-3.
HOLOTYPE: LACM 1016 (dry); PARATYPE: LACM 1083 (1 dry); PARATYPES: LACM
2148 (3 dry), Cayucos, San Luis Obispo Co., California, Leg.: H.N. Lowe, JUN 1927. Paratypes
ex UCLA 21108.
cowlesi, Turbonilla (Strioturbonilla) Strong & Hertlein, 1939: Allan Hancock Pac. Exped.
2(12):196-197.
PARATYPE: LACM 1342 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
isthmica, Odostomia (Evalea?) Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):
208-209.
PARATYPE: LACM 1156 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
sealei, Turbonilla (Pyrgiscus) Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):
201-202.
PARATYPE: LACM 1344 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
stonei, Turbonilla (Pyrgiscus) Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):
199-200.
PARATYPE: LACM 1343 (2 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
strongi, Odostomia (Evalea) Bartsch, 1927: Proc. USNM 70(2660):19-20.
PARATYPES: LACM 1958 (9 dry), Isthmus Cove, Santa Catalina Id., California Channel Ids.,
California, Leg.: A.M. Strong.
strongi, Turbonilla (Pyrgolampros) Willett, 1931b: The Nautilus 45(2):67, pl. 4, fig. 4.
HOLOTYPE: LACM 1019 (dry); PARATYPE: LACM 2144 (1 dry), 46 m, off the N shore of
Santa Catalina Id., California Channel Ids., California, Leg.: G. Willett, JUL 1920.
Remarks: Paratype ex UCLA 21106.
swetti, Odostomia (Chrysallida) Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):206.
PARATYPES: LACM 1347 (2 dry), 5-16 m, Isla Taboga, Panamá Prov., Panamá, Leg.: L.G.
Hertlein.
Remarks: Currently in the genus Chrysallida.
Family Amathinidae Ponder, 1987
carotica, Iselica Marincovich, 1973: Nat. Hist. Mus. L.A. Co. Sci. Bull. 16:40, fig. 90.
HOLOTYPE: LACM 1599 (dry); PARATYPES: LACM 1600 (5 dry), 0 m, intertidal, Punta
Morro, Iquique, Tarapaca Prov., Chile (20 13 S, 70 10 W), Leg.: L. Marincovich, JUL 1964.
chilensis, Iselica Marincovich, 1973: Nat. Hist. Mus. L.A. Co. Sci. Bull. 16:39, fig. 89.
HOLOTYPE: LACM 1597 (dry); PARATYPES: LACM 1598 (17 dry), 0 m, intertidal, Punta
Morro, Iquique, Tarapaca Prov., Chile (20 13 S, 70 10 W), Leg.: L. Marincovich, JUL 1964.
kochi, Iselica Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):227.
PARATYPE: LACM 1715 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
Superfamily ACTEONOIDEA d’Orbigny, 1843
Family Acteonidae d’Orbigny, 1843
aphyodes, “Acteon” Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:626-627.
PARATYPE: LACM 2984 (dry), 970-983 m, E of Tongatapu, Tongatapu Group, Tonga
(21°12'S, 174°56'W), Leg.: R/V Alis, P. Bouchet & others, 3 JUN 2000.
chauliodous, “Acteon” Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:631-632.
PARATYPES: LACM 2987 (dry), 570-650 m, Pass de Hienghene, E of New Caledonia
(20°34'S, 164°58'E), Leg.: R/V Alis, P. Bouchet & B. Richer, 17 MAR 1993.
chrystomatus, Acteon Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:593-594.
PARATYPES: LACM 2976 (5 dry), 110 m, between French & Cook reefs, NW of Belep Arch.,
New Caledonia Group, New Caledonia (19°7'S, 163°22'E), Leg.: R/V Vauban, P. Bouchet & B.
Richer,14 SEP 1985.
cohibilis, “Acteon” Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:619-621.
PARATYPE: LACM 2981 (dry), 577-780 m, Pass de Dumbes, SW of Noumea, New Caledonia
(22°23'S, 166°13'E), Leg.: R/V Alis, P. Bouchet & B. Richer, 16 MAY 1993.
comptus, “Acteon” Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:629-631.
PARATYPES: LACM 2986 (2 dry), 2365-2470 m, Loyalty Ridge, NE of New Caledonia
(21°24'S, 166°30'E), Leg.: R/V Jean-Charcot, P. Bouchet & others, 7 SEP 1985.
editus, “Acteon” Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:615.
PARATYPES: LACM 2979 (2 dry), 500-614 m, SE of Yasawa, Yasawa Group, Fiji (16°52.5'S,
177°43.7'E), Leg.: R/V Alis, P. Bouchet & B. Richer, 10 AUG 1998.
herosae, “Acteon” Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:627-629.
PARATYPE: LACM 2985 (dry), 105 m, Grand Recif Sud, SE of Ouen, New Caledonia Group,
New Caledonia (22°37'S, 167°4'E), Leg.: B. Richer, 30 NOV 1984.
ionfasciatus, Acteon Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:593.
PARATYPE: LACM 2975 (dry), 448-466 m, S of Mindoro Occidental Prov., Mindoro Id.,
Philippines (11°58'N, 126°6'E), Leg.: R/V Coriolis, P. Bouchet & others, 3 JUN 1985.
longissimus, Japonacteon Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:604-605.
PARATYPE: LACM 2978 (dry), 457-461 m, Arafura Sea, SE of Kepulauan Tanimbar, Indonesia
(8°19'S, 132°2'E), Leg.: R/V Baruna Jaya 1, P. Bouchet & others, 31 OCT 1991.
loyautensis, “Acteon” Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:621.
PARATYPES: LACM 2982 (2 dry), 600 m, N of Vauvilliers, Loyalty Ids., New Caledonia
(21°6'S, 167°33'E), Leg.: R/V Alis, P. Bouchet & B. Richer, 21 FEB 1989.
pascuana, Pupa Raines, 2002c: La Conchiglia 34(305):52-53, fig. 1.
HOLOTYPE: LACM 2954 (dry), 30-50 m, near Tahai, Isla de Pascua (= Easter Id.) (27°7.20'S,
109°26.30'W), Leg.: B.K. Raines.
rektos, “Acteon” Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:621-622.
PARATYPES: LACM 2983 (2 dry), 970-983 m, E of Tongatapu, Tongatapu Group, Tonga
(21°12'N, 174°56'W), Leg.: R/V Alis, P. Bouchet & others, 3 JUN 2000.
ringiculoides, “Acteon” Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:617-618.
PARATYPE: LACM 2980 (dry), 482-498 m, SW of Kelefesia, Ha’apai Group, Tonga (20°38'S,
174°58'W), Leg.: R/V Alis, P. Bouchet & others, 6 JUN 2000.
sanguinea, Rictaxis Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:596.
PARATYPE: LACM 2977 (dry), 447-450 m, Banc Stylaster, S of Isle of Pines, New Caledonia
Group, New Caledonia (23°37'S, 167°42'E), Leg.: R/V Alis, P. Bouchet & B. Richer, 29 JAN
1993.
vancouverensis, Acteon punctocoelata I.S. Oldroyd, 1927: Stanford Univ. Pubs. in Geol. Sci.
2(1):25.
PARATYPE: LACM 1078 (1 dry), 18-27 m, off Brandon Id., Departure Bay, Nanaimo Dist.,
Vancouver Id., British, Columbia, Canada, Leg.: I.S. Oldroyd.
Remarks: Junior synonym of Rictaxis punctocaelatus (Carpenter, 1864).
Family Bullinidae Gray, 1850
rubropunctata, Bullina Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:644.
PARATYPE: LACM 2988 (dry), 444-452 m, Grand Passage, NE of French Reef, New
Caledonia Group, New Caledonia (18°56'S, 163°22'E), Leg.: R/V Alis, B. Metivier & B. Richer,
7 AUG 1994.
Superfamily VALVATOIDEA Gray, 1840
Family Hyalogyrinidae Warén & Bouchet, 1993
depressa, Hyalogyrina Hasegawa, 1997: Nat. Sci. Mus. Tokyo Mem. 12:113-116
PARATYPES: LACM 2845 (2 dry), 400-740 m, off Heda, Suruga-wan, Shizuoka Pref., Honshu,
Japan (35 00.4 N, 138 43.2 E), Leg.: R/V Soro-Maru, 17 NOV 1993.
Informal Group Opisthobranchia
Clade Cephalaspidea
Superfamily PHILINOIDEA Gray, 1850
Family Philinidae Gray, 1850
alba, Philine Mattox, 1958: Bull. So. Calif. Acad. Sci. 57(2):98-104, pl. 33, fig. 1.
HOLOTYPE: LACM 2072 (wet); PARATYPE: LACM 1118 (1 dry); PARATYPES: LACM
2014 (17 wet), 64 m, 1.8 mi. SE of Long Pt., Santa Catalina Id., California Channel Ids.,
California (33 22.9 N, 118 20.7 W), Leg.: R/V Velero IV, 25 JUN 1954.
babai, Philine Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:721-722.
PARATYPE: LACM 3000 (wet), 533-610 m, NE of New Caledonia (20°33'S, 164°57'E), Leg.:
R/V Alis, R. Metivier & B. Richer, 10 AUG 1994.
californica, Philine Willett, 1944: Bull. So. Calif. Acad. Sci. 43(2):72, pl. 14, fig. 4.
HOLOTYPE: LACM 1074 (dry), 91 m, off Redondo Beach, Los Angeles Co., California, Leg.:
J.Q. & T.A. Burch.
habei, Philine Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:717-720.
PARATYPES: LACM 2999 (50 wet), 587-592 m, E of Tongatapu, Tongatapu Group, Tonga
(21°14'S, 174°59'E), Leg.: R/V Alis, P. Bouchet & others, 3 JUN 2000.
Family Cylichnidae H. Adams & A. Adams, 1853
grovesi, Cylichna Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:663-664.
PARATYPE: LACM 2992 (dry), 455 m, Pass de Dumbea, W of Noumea (22°19'S, 166°11'E),
Leg.: R/V Alis, P. Bouchet & B. Richer, 16 MAY 1993.
intermedia, Acteocina culcitella Willett, 1928: The Nautilus 42(2):38.
HOLOTYPE: LACM 1015 (dry), 55 m, off Santa Catalina Id., California Channel Ids.,
California, Leg.: G. Willett, 11 AUG 1928.
Remarks: Junior synonym of Tornastra cerealis (Gould, 1853).
mucronatum, Cylichnium Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:703-704.
PARATYPE: LACM 2997 (dry), 1004-1012 m, E of Rabi, Vanua Levu Group, Fiji (16°28'S,
179°50'W), Leg.: R/V Alis, P. Bouchet & others, 25 FEB 1999.
nanum, Cylichnium Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:704-705.
PARATYPES: LACM 2998 (5 dry), 640-668 m, Calatagan, Batangas Prov., Luzon Id.,
Philippines (13°47'N, 120°3)'E), Leg.: R/V Coriolis, P. Bouchet & others, 2 JUN 1985.
pyriformis, Sabatia Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:692-693.
PARATYPES: LACM 2993 (2 dry), 491-533 m, Pass de Hieghene, NE of New Caledonia
(20°34'S, 167°57'E), Leg.: R/V Alis, P. Bouchet & B. Richer, 17 MAR 1993.
rolleri, Utriculastra (Tornastra) Marcus, 1977: Jour. Moll. Stud., supplement 4, p. 29, figs.
82-84.
HOLOTYPE: LACM 1774 (dry), Flood Control Channel, San Diego, San Diego Co., California,
Leg.: R. Roller, 4 MAY 1970.
smithae, Roxania Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:696-698.
PARATYPES: LACM 2994 (8 dry), 425 m, NE of Lifou Id., Loyalty Ids., New Caledonia
(21°1'S, 167°31'E), Leg.: R/V Alis, P. Bouchet & B. Richer, 20 FEB 1989.
stephensae, Cylichna Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):190.
PARATYPE: LACM 1339 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
tabogaensis, Cylichna (Cylichnella) Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):
191-192.
PARATYPE: LACM 1341 (1 dry), 5-16 m, off Isla Taboga, Panamá Prov., Panamá, Leg.: L.G.
Hertlein.
Remarks: Currently in the genus Cylichnella.
tanyumphalos, Cylichna Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:661-663
PARATYPE: LACM 2991 (dry), 250-350 m, Western Reefs, SE of New Caledonia (22°30'S,
166°24'E).
veleronis, Cylichna Strong & Hertlein, 1939: Allan Hancock Pac. Exped. 2(12):191.
PARATYPE: LACM 1340 (1 dry), 5-16 m, Bahía Honda, Isla Coiba, Veraguas Prov., Panamá,
Leg.: L.G. Hertlein.
Family Aglajidae Pilsbry, 1895
regiscorona, Aglaja Bertsch, 1972: The Veliger 15(2):103-104.
PARATYPES: LACM 1617 (3 wet), 0 m, intertidal, Bahía Las Cruces, Golfo de California, Baja
California Sur, México (24 13 N, 110 05 W), Leg.: H. Bertsch, 19-22 JUL 1969.
Family Retusidae Thiele, 1925
abyssicola, Retusa Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:737.
PARATYPE: LACM 3004 (dry), 1620-1630 m, W of Lifou Id., Loyalty Ids., New Caledonia
(20°48'S, 166°53'E), Leg.: R/V Coriolis, P. Lozouet, 2 MAY 1987.
diaphana, Retusa Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:733-735.
PARATYPES: LACM 3001 (24 dry), 350 m, W of ‘Uvea, Wallis Id., Wallis & Futuna (13°19'S,
176°17'W), Leg.: R/V Alis, P. Bouchet & others, 25 MAY 1992.
hadra, Relichna Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:764-765.
PARATYPE: LACM 3008 (dry), 514-530 m, Norfolk Ridge, S of Isle of Pines, New Caledonia
Group, New Caledonia (24°54'S, 168°22'E), Leg.: R/V Alis, P. Bouchet & B. Richer, 27 JAN
1993.
insolita, Retusa Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:735-736
PARATYPES: LACM 3002 (5 dry), 778-811 m, SW of Espiritu Santo, New Hebrides Group,
Vanuatu (16°2'S, 166°38'E), Leg.: R/V Alis, P. Bouchet & B. Richer, 10 OCT 1994.
lenis, Retusa Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:736-737.
PARATYPES: LACM 3003 (2 dry), 200 m, E of Lifou Id., Loyalty Ids., New Caledonia
(20°54'S, 167°17'E), Leg.: R/V Alis, P. Bouchet & B. Richer, 19 FEB 1989.
multistriata, Volvulella Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:752-753.
PARATYPES: LACM 3007 (2 dry), 497-520 m, Pass de Hienghene, E of New Caledonia
(20°34'S, 164°57'E), Leg.: R/V Alis, P. Bouchet & B. Richer, 17 NOV 1993.
onoae, Volvulella Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:751-752.
PARATYPES: LACM 3006 (4 dry), 250-350 m, SW of Noumea, New Caledonia (22°30'S,
166°24'E).
tenuissima, Volvulella Willett, 1944: Bull. So. Calif. Acad. Sci. 43(2):71-72, pl. 14, fig. 1.
HOLOTYPE: LACM 1073 (dry); PARATYPES: LACM 2147 (67 dry), 137 m, off Redondo
Beach, Los Angeles Co., California, Leg.: J.Q. & T.A. Burch; PARATYPE: LACM 1693 (1 dry),
91 m, off Redondo Beach, Los Angeles Co., California, Leg.: J.Q. Burch, AUG 1940.
Remarks: Paratype lot LACM 2147 ex UCLA 22167.
trunca, Retusa Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:737-740.
PARATYPES: LACM 3005 (11 dry), 735 m, SW of Tongatapu, Tongatapu Group, Tonga
(21°16'S, 175°18'E), Leg.: R/V Alis, P. Bouchet & others, 5 JUN 2000.
Superfamily DIAPHANOIDEA Odhner, 1914
Family Diaphanidae Odhner, 1914
epongensis, Toledonia Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:650-651.
PARATYPES: LACM 2990 (3 dry), 447-450 m, Banc Stylaster, SE of Isle of Pines, New
Caledonia Group, New Caledonia (23°37'S, 167°42'E), Leg.: R/V Alis, P. Bouchet & B. Richer,
23 JAN 1993.
neocaledonica, Toledonia Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:649-650.
PARATYPE: LACM 2989 (dry), 405-410 m, Banc Jumeau Ouest, SE of Isle of Pines, New
Caledonia Group, New Caledonia (23°42'S, 168°16'E), Leg.: R/V Alis, P. Bouchet & others,28
NOV 1993.
pacifica, Diaphana Schiøtte, 1998: Steenstrupia 24(1):128-129, figs. 26A-B.
HOLOTYPE: LACM 2875 (dry), 82 m, 0.4-0.7 mi. 110 to 132 T from Ship Rock, Santa
Catalina Id, California Channel Ids., California (33 27 N, 118 30 W), Leg.: R/V Velero IV, 13
FEB 1965.
Superfamily HAMINOEOIDEA Pilsbtu, 1895
Family Haminoeidae Pilsbry, 1895
wareni, Micratys Valdés, 2008. Mem. Mus. Nat. Hist. Naturelle 196:770-772.
PARATYPES: LACM 3010 (2 dry), 570-650 m, Pass de Hienghene, NE of New Caledonia
(20°34'S, 164°58'E), Leg.: R/V Alis, P. Bouchet & B. Richer, 17 MAR1993.
Clade Aplysiomorpha [= Anaspidea]
Superfamily APLYSIOIDEA Lamarck, 1809
Family Aplysiidae Lamarck, 1809
nettiae, Aplysia (Neaplysia) Winkler, 1959: Bull. So. Calif. Acad. Sci. 58(1):8-10, pl. 5.
HOLOTYPE: LACM 2013 (wet), Mouth of Topanga Canyon, Malibu, Los Angeles Co.,
California, Leg.: E.Y. Dawson & F. Durham, 18 DEC 1945.
Remarks: Ex AHF 991.
reticulopoda, Aplysia Beeman, 1960: Bull. So. Calif. Acad. Sci. 59(3):144-151, pls. 46-48.
HOLOTYPE: LACM 2011 (wet); PARATYPE: LACM 2012 (1 wet), Laguna, Orange Co.,
California
vaccaria, Aplysia Winkler, 1955: Bull. So. Calif. Acad. Sci. 54(1):5-7.
HOLOTYPE: LACM 2010 (wet), PARATYPES: LACM 1231 (2 dry), Pt. Fermin, San Pedro,
Los Angeles Co., California, 14 MAR 1954.
Remarks: Holotype ex AHF 983.
Superfamily AKEROIDEA Mazzarelli, 1891
Family Akeridae Mazzarelli, 1891
juliae, Akera Valdés & Barwick, 2005: Nautilus 119(1):44-49, figs. 1, 2, 3A-C, 4A-C
HOLOTYPE: LACM 3033 (wet), 40.6 m, off SW tip of Santa Catalina Id., California Channel
Ids., California (33°18.24'N, 118°22.5'W), 24 JUL 1998; PARATYPES: LACM 3034 (3 dry),
1.5-10.7 m, SE corner of Bahía Jobo, off beach W of Bahía Salinas, Guanacaste Prov., Costa
Rica (11°2.22'N, 85°45.16'W), 14 FEB 1972.
“G roup“ Acochlidiacea
Superfamlily ACOCHLIDIOIDEA Küthe, 1935
Family Acochlidiidae Küthe, 1935
fijiensis, Acochlidium Haynes & Kenchington, 1991: The Veliger 34(2):166-171, figs. 1-6.
HOLOTYPE: LACM 2457 (wet); PARATYPES: LACM 2458 (2 wet), at the Labasa-Savusavu
Highway bridge, 7 km from the Nasekawa River mouth, Vanua Levu, Vanua Levu Group, Fiji
(16 40 S, 179 16 E), Leg.: A. Haynes, JUL 1989.
Family Limapontiidae Gray, 1847
willowi, Alderia Krug, Ellingson, Burton, & Valdés, 2007: Jour. Moll. Stud. 73:33-34, fig.1C.
HOLOTYPE: LACM 2034 (wet); PARATYPES: LACM 2035 (4 wet), Cabrillo Aquarium
wetlands, San Pedro, Los Angeles Co., California (33º42'48"N, 118º17'2"W), Leg.: P. Krug, 18
JUL 2005.
Clade Umbraculidae
Superfamily UMBRANCULOIDEA Dall, 1889
Family Tylodinidae Gray, 1847
spongotheras, Anidolyta Bertsch, 1980: Sarsia 65(3):234-236, figs. 2b, 3a-b.
HOLOTYPE: LACM 1916 (wet), 25-100 m, Queen's Reach, off Hill Rock, Jervis Inlet, Sunshine
Coast Dist., British Columbia, Canada (50 09.8 N, 123 50.4 W), Leg.: W. Austin, 14 MAR
1976.
Remarks: Radula mounted on glass slide.
Clade Nudipleura
Subclade Pleurobranchomorpha
Superfamily PLEUROBRANCHOIDEA Gray, 1827
Family Pleurobranchidae Gray, 1827
amarillius, Pleurobranchus (Oscanius) Mattox, 1953: The Nautilus 66(4):110-114.
PARATYPES: LACM 2016 (13 dry), Isla de la Gata, Parguera, Southwest Coast, Puerto Rico,
Leg.: N. Mattox, MAR 1950-APR 1952.
antarctica, Bathyberthella Willan & Bertsch, 1987: The Veliger 29(3):292-296.
PARATYPES: LACM 2120 (2 wet), 360-486 m, W of South Sandwich Ids., South Atlantic
(56 40.6 S, 27 00.8 W), Leg.: R/V Islas Orcada, 30 MAY 1975.
kaniae, Berthella Sphon, 1972: The Nautilus 86(2/4):53-55.
PARATYPE: LACM 1453 (1 dry), 0 m, intertidal, Punta Mita (ca. 20 mi. N of Puerto Vallarta),
Nayarit, México (20 46 N, 105 33 W), Leg.: G. Sphon, 21-22 JAN 1970.
Remarks: Jaw plates on glass slide.
Clade Euctenidiacea
Subclade Gnathodoridacea
Superfamily BATHYDORIDOIDEA Bergh, 1891
Family Bathydoridae Bergh, 1891
spiralis, Bathydoris Valdés, 2002: Can. Jour. Zool. 80:1090-1091.
PARATYPE: LACM 2900 (1wet), 305-315 m, Banc Azteque, SE of Isle of Pines, New
Caledonia Group, New Caledonia (2318’S, 1685’E), Leg.: P. Bouchet & B. Richer, 31 JAN
1993.
Subclade Doridacea
Superfamily DORIDOIDEA Rafinesque, 1815
Family Dorididae Rafinesque, 1815
mimeticus, Asteronotus Gosliner & Valdés, 2002: Proc. Calif. Acad. Sci. 53(5):52-57.
PARATYPE: LACM 2901 (1 wet), 15 m, Arthur’s Rock, SW side of Calumpan Peninsula,
Batangas Prov., Luzon Id., Philippines, Leg.: T.M. Gosliner, 22 FEB 1992.
spongicolus, Asteronotus Gosliner & Valdés, 2002: Proc. Calif. Acad. Sci. 53(5):57-61.
PARATYPE: LACM 2902, 2 m, 3 km N of Uroa Beach Motel, Uroa, Zanzibar, Tanzania, Leg.:
T.M. Gosliner, NOV 1994.
tanya, Doris Marcus, 1971: Proc. Malac. Soc. London 39:357-362, figs. 4-8.
HOLOTYPE: LACM 1768 (wet), Newport Bay, Orange Co., California, Leg.: R.L. Miller, SUM
1968.
Family Discodorididae Bergh, 1891
carolynae, Platydoris Mulliner & Sphon, 1974: Trans. San Diego Soc. Nat. Hist. 17(15):
209-213.
PARATYPE: LACM 1619 (1 wet), 0 m, intertidal, Isla Pinzon (= Duncan Id.), Islas Galápagos
(0 35.8 S, 90 39.2 W), Leg.: Ameripagos Exped., 25-26 MAR 1971.
farmersi, Diaulula Valdés, 2004: Nat. Hist. Mus. L.A. Co. Cont. Sci. 501:6-7, figs. 1B, 4-5.
HOLOTYPE: LACM 3016 (wet), 1-2 m, in front of Bayside Resort, mile marker 99.5, Key
Largo, Monroe Co., Florida, Leg.: A. Valdés, 13-22 JUL 2003.
lopezi, Paradoris Hermosillo & Valdés, 2004: Proc. Calif. Acad. Sci., 4th ser. 55(28):551-556,
figs. 1A-B, 2-3.
PARATYPE: LACM 3041 (wet), 12 m, Majahuitas, Bahía de Banderas, Jalisco, México
(20º29.111' N, 105º35.057' W), 10 MAR 2004
pardus, Jorunna Behrens & Henderson, 1981: The Veliger 24(2):125-128.
PARATYPES: LACM 1949 (3 wet), 5 m, Cat Rock, Anacapa Id., California Channel Ids.,
California (34 15 N, 119 25.3 W), Leg.: D. Behrens, 25 OCT 1979.
rubescens, Thordisa Behrens & Henderson, 1981: The Veliger 24(2):120-124.
PARATYPE: LACM 1948 (1 wet), 22 m, Big Kelp Reef, Paradise Cove, Los Angeles Co.,
California, Leg.: R. Henderson, 8 OCT 1978.
Remarks: Radula mounted on glass slide.
Family Chromodorididae Bergh, 1891
andersoni, Hypselodoris Bertsch & Gosliner, 1989: The Veliger 32(3):247-254.
PARATYPES: LACM 2294 (5 wet), 15 m, Pupukea Beach Co. Park, Waialua Dist., O‘ahu,
Hawaiian Ids., Leg.: S. Johnson, 18 SEP 1978.
antonii, Chromodoris Bertsch, 1976: The Veliger 19(2):156-158, figs. 1-8.
HOLOTYPE: LACM 1810 (wet), 9 m, Punta Aguja (near Mulege), Golfo de California, Baja
California Sur, México (26 53 N, 111 56 W), Leg.: A.J. Ferreira, 28 JUL 1974; PARATYPE:
LACM 1811 (1 wet), 10-12 m, Isla San Jose, Golfo de California, Baja California Sur, México
(24 53 N, 11 34 W), Leg.: A.J. Ferreira, 14 AUG 1973; PARATYPE: LACM 1812 (1 wet)
28 m, Los Islotes, Golfo de California, Baja California Sur, México (24 36 N, 110 24 W), Leg.:
A.J. Ferreira, 14 AUG 1973; PARATYPE: LACM 1813 (1 wet), Bahía Santiago, near
Manzanillo, Colima, México (19 06 N, 104 23 W), Leg.: L. Shy, R. & F. Poorman, JAN 1976.
Remarks: Holotype radula mounted on glass slide, paratype radulae LACM 1811 and LACM
1813 mounted on glass slides.
galexorum, Chromodoris Bertsch, 1978a: The Veliger 20(4):310-312, figs. 5, 33-36.
HOLOTYPE: LACM 1848 (wet); PARATYPES: LACM 1908 (2 wet), Isla San Pedro Martir,
Sonora, México (28 22 N, 112 20 W), Leg.: A. Kerstich, 16 JUN 1976.
Remarks: Holotype and paratype radulae mounted on glass slides.
ghiselini, Hypselodoris Bertsch, 1978b: The Veliger 21(2):245-247, figs. 28, 65-68.
HOLOTYPE: LACM 1849 (wet); PARATYPES: LACM 1909 (2 wet), 2-3 m, Bahía de las
Cruces, Golfo de California, Baja California Sur, México (24 13 N, 110 05 W), Leg.: M.
Ghiselin, H. Bertsch, & J. Allen, 1 JUL 1974.
Remarks: Holotype and paratype radulae mounted on glass slides.
lapislazuli, Thorunna Bertsch & Ferreira, 1971: The Veliger 16(4):343-345.
PARATYPES: LACM 1614 (2 wet), 0 m, intertidal, Charles Darwin Research Station, Bahía
Academy, Isla Santa Cruz (= Indefatigable Id.), Islas Galápagos (0 45 S, 90 15 W), Leg.: G.
Sphon, 27 MAR 1971.
lilyevae, Hypselodoris Alejandrino & Valdés, 2006: Jour. Molluscan Stud. 72(2):191-196, figs.
1A-B.
HOLOTYPE: LACM 3017 (wet); PARATYPE: LACM 3018 (1 wet), 12-18 m, Pelican Ghut,
Guana Id., British Virgin Ids., Leg.: T. Zimmerman, 11 AUG 1999.
marislae, Chromodoris Bertsch in Bertsch, Ferreira, Farmer, & Hayes, 1973: The Veliger 15(4):
289-292.
PARATYPES: LACM 1618 (3 wet), La Paz, Golfo de California, Baja California Sur, México,
Leg.: E. Janss, APR 1972.
poliahu, Glossodoris Bertsch & Gosliner, 1989: The Veliger 32(3):256-259.
PARATYPES: LACM 2295 (4 wet), 15 m, Pupukea Beach Co. Park, Waialua Dist., O‘ahu,
Hawaiian Ids., Leg.: S. Johnson, 12 JUN 1978; PARATYPES: LACM 2296 (4 wet), 15 m, Three
Tables, S of Pupukea Beach Co. Park, Waialua Dist., O‘ahu, Hawaiian Ids., Leg.: S. Johnson, 10
JUN 1978.
sphoni, Felimida Marcus, 1971: Proc. Malac. Soc. London 39:356-357.
SYNTYPES: LACM 1616 (2 wet), 0 m, intertidal, Santa Cruz, ca. 15 mi. S of San Blas, Nayarit,
México (21 31'N, 105 16'W), Leg.: G. Sphon, 5-8 JAN 1970.
tibboeli, Glossodoris Valdés & Adams, 2005: Pacific Science 59(4):603-608, figs. 1-3.
HOLOTYPE: LACM 3044 (wet); PARATYPES: LACM 3045 (3 wet);10 m, Para Id., N of
Sulawesi, Kepulauan Sangir, Indonesia (3°2.84'S, 125°30.20'E), Leg.: M.J. Adams, 13 JUN
2004.
tomsmithi, Glossodoris Bertsch & Gosliner, 1989: The Veliger 32(3):259-261.
PARATYPES: LACM 2297 (3 wet), 21 m, off Makaha, Wai`anae Dist., O‘ahu, Hawaiian Ids.,
Leg.: H. Bertsch & J. Young, 17 SEP 1978; PARATYPE: LACM 2326 (1 wet), 15 m, off Halona
Blowhole (near Lana‘i lookout), Honolulu Dist., O‘ahu, Hawaiian Ids., Leg.: H. Bertsch & S.
Johnson, 26 SEP 1978.
Superfamily ONCHIDORIDOIDEA Gray, 1827
Family Goniodorididae Adams & Adams, 1854
goddardi, Trapania Hermosillo & Valdés, 2004: Proc. Calif. Acad. Sci., 4th ser., 55(28):556-559,
figs. 1C, 4-5.
HOLOTYPE: LACM 3039 (wet), 9 m, Islas Marietas, Bahía de Banderas, Jalisco, México
(20º42.042' N, 105º33.878' W), 17 APR 2003; PARATYPE: LACM 3040 (wet), 12 m, Islas
Marietas, Bahía de Banderas, Jalisco, México (20º42.042' N, 105º33.878' W), 17 APR 2003
Superfamily POLYCEROIDEA Alder & Hancock, 1845
Family Polyceridae Alder & Hancock, 1845
kaiseri, Polycera Hermosillo & Valdés, 2007: Proc. CAS 58(23):478-483, figs. 1B, 2B-C, 3.
PARATYPES: LACM 1850 (3 wet), 11 m, Mismaloya, Bahía de Banderas, Jalisco, México
(20º31.937'N, 105º17.700'W). 11 JAN 2006.
Superfamily PHYLLIDIOIDEA Rafinesque, 1814
Family Phyllidiidae Rafinesque, 1814
suzanneae, Reticulidia Valdés & Behrens, 2002: Nat. Hist. Mus. L.A. Co. Contr. Sci. 492:5-6,
figs. 1C, 3.
HOLOTYPE: LACM 2897 (wet), 21.3 m Similan Ids., Andaman Sea, Phuket Changwat,
Thailand (8°40.5'N, 97°38.8'E), Leg.: M. Strickland, FEB 2001.
Family Dendrodorididae O’Donoghue, 1814
azineae, Dendrodoris Behrens & Valdés, 2004: Pr. Calif. Acad. Sci. 55(21):409-413, figs. 1A, 23.
HOLOTYPE: LACM 3035 (wet), 54 m, La Jolla Canyon, San Diego Co., California, Leg.: G.
Spaulding, 14 MAR 2004.
Superfamily POLYCEROIDEA Alder & Hancock, 1845
Family Gymnodorididae Donner, 1941
abdera, Tambja Farmer, 1978: The Veliger 20(4):377-378.
PARATYPE: LACM 1717 (1 wet), La Paz area, Golfo de California, Baja California Sur,
México, Leg.: E. Janss, MAY 1972.
fusca, Tambja Farmer, 1978: The Veliger 20(4):378-379.
PARATYPE: LACM 1719 (1 wet), Isla Monserrate, Golfo de California, Baja California Sur,
México, Leg.: E. Janss, APR 1974.
mullineri, Tambja Farmer, 1978: The Veliger 20(4):379-380.
PARATYPE: LACM 1718 (1 wet), 11 m, Bahía James, Isla San Salvador (= James Id.), Islas
Galápagos, Leg.: Ameripagos Exped., MAR 1971; PARATYPE: LACM 1775 (1 wet), 11 m, Isla
Rabida (= Jervis Id.), Islas Galápagos, Leg.: Ameripagos Exped., MAR 1971.
tigris, Roboastra Farmer, 1978: The Veliger 20(4):381-383.
PARATYPE: LACM 1720 (1 wet), 24-27 m, 20 mi. N of Guaymas, Sonora, México, Leg.: B.
van Zandt, APR 1969.
Clade Cladobranchia
Subclade Euarminida
Superfamily ARMINOIDEA Iredale & O’Donoghue, 1923
Family Arminidae Iredale & O’Donoghue, 1923
juliana, Armina Ardilla & Diaz, 2002: Bol. Inv. Mar. Cos. 31:27-29.
PARATYPE: LACM 2908 (1 wet), 306-312 m, off Palomino, La Guajira Dep., Colombia
(1°26’N, 73°32’W).
Remarks: Jaw plate and radulae on SEM stub.
Subclade Dendronotida
Superfamily TRITONIOIDEA Lamarck, 1810
Family Tritoniidae Lamarck, 1809
aurantia, Tritoniopsis Mattox, 1955: Bull. So. Calif. Acad. Sci. 54(1):8-13, pl. 4.
HOLOTYPE: LACM 2015 (wet), 73 m, 1.3 mi. SE of Long Pt., Santa Catalina Id., California
Channel Ids., California, Leg.: R/V Velero IV, 26 JUN 1954.
Remarks: Ex AHF 985.
myrakeenae, Tritonia Bertsch & Mozqueira, 1986: The Nautilus 100(2):47-49, fig. 1.
HOLOTYPE: LACM 2135 (wet); PARATYPE: LACM 2136 (1 wet), 0 m, intertidal, SE end of,
Isla Cedros, Pacific Coast Baja California, México (28 02.5 N, 115 12 W), Leg.: H. Bertsch & S.
Tsung, 26 DEC 1985.
Family Scyllaeidae Alder & Hancock, 1855
daedali, Crosslandia Poorman & Mulliner, 1981: The Nautilus 95(2):96-99.
PARATYPES: LACM 1910 (4 wet), Tinajas, Bahía de Bacochibampo, Sonora, México
(27 56 N, 110 59 W), Leg.: F. & R. Poorman, 4 DEC 1975.
Subclade Aeolidida
Superfamily FLABELLINOIDEA Bergh, 1889
Family Flabellinidae Bergh, 1889
riwo, Flabellina Gosliner & Willan, 1991: The Veliger 34(2):105-108.
PARATYPE: LACM 2465 (1 wet), 15.2 m, Barracuda Pt., E side of Pig Id. (= Tab Id.), N coast
of Madang Prov., Papua New Guinea, Leg.: T.M. Gosliner, 13 JAN 1988.
Superfamily FIONOIDEA Gray, 1857
Family Eubranchidae Odhner, 1934
yolandae, Eubranchus Hermosillo & Valdés, 2007: Amer. Malac. Bull. 22(1/2):130-133, figs.
1E, 10-11.
HOLOTYPE: LACM 3055 (wet); PARATYPES: LACM 3056 (2 wet), 17 m, Los Arcos, Bahía
de Banderas, Jalisco, México (20º32.86'N, 105º17.34'N), 26 MAY 2004; PARATYPE: LACM
3057 (wet), 17 m, Mismaloya, Bahía de Banderas, Jalisco, México (20º31.94'N, 105º17.70'W),
21 MAY 2004.
Remarks: Holotype radula and jaw mounted on SEM stub.
Family Tergipedidae Bergh, 1889
behrensi, Cuthona Hermosillo & Valdés, 2007: Amer. Malac. Bull. 22(1/2):128-130, figs. 1F, 89.
HOLOTYPE: LACM 3054 (wet), 13 m, Los Frailes, Azuero Peninsula, Panama (7º22.37'N,
8º9.29'W), 2 MAY 2003.
Remarks: Radula and jaws mounted on SEM stub.
destinyae, Cuthona Hermosillo & Valdés, 2007: Amer. Malac. Bull. 22(1/2):121-124, figs. 1C,
4-5.
HOLOTYPE: LACM 3052 (wet), La Godornia, Zihuatanejo, Guerrero, México (17º 37.85'N,
101º33.56'W), Leg.: A. Hermosillo & D. Behrens, 19 MAR 2004.
Remarks: Radula and jaws mounted on SEM stub
lizae, Cuthona Angulo & Valdés, 2003: Veliger 46(2):179-182, figs. 2- 3.
HOLOTYPE: LACM 2951 (wet); PARATYPES: LACM 2952 (4 wet), Ensenada de los
Muertos, Golfo de California, Baja California Sur, México, Leg.: O. Angulo, 16 JAN 2002.
Remarks: Jaw, masticatory border elements, radula, and penal spine mounted on SEM stub
longi, Cuthona Behrens, 1985: The Veliger 27(4):418-421.
PARATYPE: LACM 2085 (1 wet), 3.1 m, Isla Raza, Golfo de California, Baja California,
México (28 48 N, 113 00 W), Leg.: J. Hamann, JUL 1982.
millenae, Cuthona Hermosillo & Valdés, 2007: Amer. Malac. Bull. 22(1/2):124-128. figs. 1D,
6-7.
HOLOTYPE: LACM 3053 (wet), 19 m, Los Arcos, Bahía de Banderas, Jalisco, México
(20º32.86'N, 105º17.34'W), 29 MAY 2003; PARATYPE: LACM 3050 (wet), Islas Maietas,
Bahía de Banderas, Jalisco, México (20º42.042'N, 105º33.878'W), 11 JAN 2005
Remarks: Holotype radula and jaws mounted on SEM stub
rickettsi, Catriona Behrens, 1984: The Veliger 27(1):66-70.
PARATYPES: LACM 2024 (6 wet), Port of Redwood City, San Francisco Bay, San Mateo Co.,
California (37 30.03 N, 112 13.38 W), Leg.: D. Behrens, 24 DEC 1981
Superfamily AEOLIDIOIDEA Gray, 1827
Family Aeolidiidae Gray, 1827
chavezi, Cerberilla Hermosillo & Valdés, 2007: Amer. Malac. Bull. 22(1/2):133-136, figs. 1A,
12-13.
HOLOTYPE: LACM 3058 (wet), 6 m, La Boquita, Bahía de Santiago, Colima, México
(19º6.30'N, 104º23.92'W), 21 FEB 2004; PARATYPE: LACM 3059 (wet), 10 m, La Cruz de
Huanacaxtle, Bahía de Banderas, Nayarit, México (20º44.44'N, 105º23.16'W), 14 JAN 2004;
PARATYPE: LACM 3060 (wet), 6 m, La Boquita, Bahía de Santiago, Colima, México
(19º6.30'N, 104º23.92'W), 26 MAR 2004.
Remarks: Radula and jaw mounted on SEM stub.
mosslandica, Cerberilla McDonald & Nybakken, 1975: The Veliger 17(4):378-381.
PARATYPES: LACM 1725 (2 wet), 63 m, Monterey Bay, Monterey Co., California (36 52.3 N,
121 59.8 W), Leg.: Gary McDonald, 18 NOV 1972; PARATYPE: LACM 1726 (1 wet), 34.5 m,
Monterey Bay, Monterey Co., California (36 53.6 N, 121 57.5 W), Leg.: G. McDonald, 10 MAY
1972.
Family Facelinidae Bergh, 1889
morroensis, Emarcusia Roller, 1972: The Veliger 14(4):420-422.
PARATYPES: LACM 1615 (3 wet), Morro Bay, San Luis Obispo Co., California (35 22 N,
120 51 W), Leg.: R. Roller, 15 NOV 1969.
Informal Group Pulmonata
Informal Group Basommatophora
Superfamily SIPHONARIOIDEA Gray, 1827
Family Siphonariidae Gray, 1827
stewartiana, Kerguelenia Powell, 1939: Rec. Auckland Inst. and Mus. 2:237-238.
PARATYPES: LACM 1215 (4 dry), Aker's Pt., Stewart Id., New Zealand
Remarks: Currently in the genus Siphonaria.
Clade Hygrophila
Superfamily CHILINOIDEA Dall, 1870
FFamily Chilinidae Dall, 1870
oldroydae, Chilina Marshall, 1924: Proc. USNM 66(13):4.
PARATYPE: LACM 1076 (1 dry); PARATYPE: LACM 2050 (1 dry), Lago Fetalafquen,
Chubut Prov., Argentina, Leg.: J. Pemberton.
Superfamily PLANORBOIDEA Rafinesque, 1815
Family Planorbidae Rafinesque, 1815
nevadensis, Parapholyx effuse Henderson, 1934: The Nautilus 47(3):90-91.
PARATYPES: LACM 1080 (3 dry), Lake Winnemucca, Humboldt Co., Nevada, Leg.: H.
Hannibal.
Remarks: Fossil species currently in the genus Vorticifex.
Family Physidae Fitzinger, 1833
natricina, Physa (Haitia) Taylor, 1988: Malacological Review 21:67-72, figs. 6k-n.
HOLOTYPE: LACM 2970 (shell dry, animal wet), Snake River SW¼ SE¼ sec. 21, T6N, R13E,
Gooding Co., Idaho, Leg.: D.W. Taylor, 27 JUL 1980.
Remarks: Originall published as LACM 2256 (preoccupied).
Clade Eupulmonata
Superfamily ELLOBIOIDEA Pfeiffer, 1854
Family Ellobiidae Pfeiffer, 1854
colmani, Pythia Martins, 1995: Moll. Res. 16:60-67.
PARATYPE: LACM 2803 (1 dry), elev 100 m, Medina, New Ireland, Bismark Archipelago,
Papua New Guinea (2 56 S, 151 22.5 E), Leg.: P. Colman et al.
mousleyi, Melampus Berry, 1964: Leaflets in Malacology 1(24):152-153.
PARATYPES: LACM 2122 (4 dry), Bahía Cholla, Sonora, México, Leg.: L.B. Mousley, 29
NOV 1951.
Clade Stylommatophora
Subclade Elasmognatha
Superfamily SUCCINEOIDEA Beck, 1837
Family Succineidae Beck, 1837
utowana, Succinea Clench, 1933: Proc. New England Zool. Club 13:85.
PARATYPES: LACM 1669 (3 dry), back of Victory Hill Settlement, Wattling Id., Bahamas.
Subclade Orthurethra
Superfamily PARTULOIDEA Pilsbry, 1900
Family Partulidae Pilsbry, 1900
variabilis, Partula Pease, 1866: Amer. Jour. Conch. 2(3):203.
PARATYPE: LACM 1877 (1 dry), Raiatea, Society Ids., French Polynesia.
Remarks: Ex W.O. Gregg collection 7238.
Superfamily COCHLICOPOIDEA Pilsbry, 1900
Family Amastridae Pilsbry, 1910
metamorpha, Amastra Pilsbry & Cooke, 1914: Tryon's Manual of Conchology, 2nd ser.,
23:19-20.
PARATYPES: LACM 2060 (4 dry), Olowalu Gulch, Lahaina Dist., Maui, Hawaiian Ids., Leg.:
D. Thaanum, JUL 1913.
Superfamily PUPILLOIDEA Turton, 1831
Family Pupillidae Turton, 1831
sterkiana, Pupa Pilsbry, 1889: Proc. Acad. Nat. Sci. Philadelphia 41:411.
SYNTYPES: LACM 1333 (45 dry); PARATYPES: LACM 1690 (11 dry), Near Bahía de San
Quintín, Pacific Coast Baja California, México, Leg.: C.R. Orcutt, APR 1886.
Remarks: Currently in the genus Pupilla.
Family Vertiginidae Fitzinger, 1833
mariposa, Vertigo ovata Pilsbry, 1919: Tryon's Manual of Conchology, 2nd ser., 25:88-89.
PARATYPES: LACM 1688 (28 dry), Meadow near Wawona, Yosemite National Park, Mariposa
Co., California, Leg.: H.N. Lowe, 1916.
Remarks: Junior synonym of Vertigo ovata Say, 1822 fide Roth & Sadeghian (2003).
microphasma, Vertigo modesta Berry, 1919b: The Nautilus 33(2):48-51.
PARATYPES: LACM 2199 (2 dry), 2286 m, cienaga near Bluff Lake, San Bernardino
Mountains, San Bernardino Co., California, Leg.: N.G. Spaulding, G.E. Dole, & S.S. Berry,
AUG 1910.
Remarks: Junior synonym of Vertigo modesta castanea (Pilsbry & Vanatta, 1900) fide Roth &
Sadeghian (2003).
Informal Group Sigmurethra
Superfamily CLAUSILIOIDEA Gray, 1855
Family Clausiliidae Gray, 1855
aequistriata, Gracilinenia Weyrauch, 1956b: The Nautilus 69(4):111-113.
PARATYPES: LACM 2612 (3 dry), 1300 m, on road from San Ramon to Mina Pichita Caluga,
Chanchamayo Valley, Junin Dept., Perú, Leg.: W. Weyrauch.
boettgeri, Nenia quadrata Pilsbry, 1945: The Nautilus 58(3):81-82.
PARATYPES: LACM 2049 (5 dry), 2300 m, Carpapata, Rio Tarma, Pasco Dept., Perú, Leg.: W.
Weyrauch.
cajamarcana, Steeriana (Steeriana) Weyrauch & Zilch in Zilch, 1954: Arch. Moll.
83(1/3):73-76.
PARATYPES: LACM 1329 (5 dry), 2850 m, Hacienda Cochambul, near Cajamarca, Cajamarca
Dept., Perú, Leg.: W. Weyrauch.
celendinensis, Steeriana (Steeriana) Weyrauch & Zilch in Zilch, 1954: Arch. Moll.
83(1/3):70-72.
PARATYPES: LACM 1330 (5 dry), 2700 m, 3 km S of Celendin, at km 113 on the highway
from Celendin to Balsas, Cajamarca Dept., Perú, Leg.: W. Weyrauch.
huarangoensis, Steeriana (Steeriana) Zilch, 1949: Arch. Moll. 78(1/3):94-95.
SYNTYPES: LACM 2239 (2 dry), 1400 m, Huarango, E of Rio Chinchipe, Cajamarca Dept.,
Perú, Leg.: W. Weyrauch.
isidroensis, Steeriana (Steeriana) celendinensis Weyrauch & Zilch in Zilch, 1954: Arch. Moll.
83(1/3):72-73.
PARATYPES: LACM 1331 (5 dry), 2750-2800 m, Mount San Isidro, near Celendin, Cajamarca
Dept., Perú, Leg.: W. Weyrauch.
omissa, Temesa Weyrauch, 1957: Arch. Moll. 86(1/3):18-21.
PARATYPES: LACM 1332 (5 dry), 3400-3600 m, Tarmatabo, near Tarma, Loreto Dept., Perú,
Leg.: W. Weyrauch.
urubambensis, Nenia angrandi Pilsbry, 1945: The Nautilus 58(3):83-84.
PARATYPES: LACM 2048 (5 dry), 1400 m, Huadquena Vilcanota Valley, Cuzco Dept., Perú,
Leg.: W. Weyrauch.
Superfamily ORTHALICOIDEA Albers, 1860
Family Orthalicidae Albers, 1860
alcaldei, Liguus fasciatus Roig, 1951: Revista Sociedad Malac. 8(1):23-24.
PARATYPES: LACM 1564 (3 dry), Monte de Hernandez, Guanabana, Cienfuegos, Cienfuegos
Prov., Cuba.
multilineatus, Liguus fasciatus Alcalde, 1944: Revista Sociedad Malac. 2(1):13.
PARATYPES: LACM 1563 (2 dry), Monte del Tarabico, Hoyo de Padilla, Cienfuegos,
Cienfuegos Prov., Cuba.
tigrae, Liguus fasciatus Alcalde, 1944: Revista Sociedad Malac. 2(1):12.
PARATYPE: LACM 1562 (1 dry), Monte de Lewis, Gavilan, Cienfuegos, Cienfuegos Prov.,
Cuba.
Family Bulimulidae Tryon, 1867
annectens, Placostylus (Maoristylus) ambagiosus Powell, 1938: Rec. Auckland Inst. Mus.
2(3):148-149.
PARATYPE: LACM 2056 (1 dry), 274 m, Unuwharo, on track between Spirits Bay & Tom
Boling Bay, Northland, North Id., New Zealand.
camachoi, Neopetraeus Weyrauch, 1967: Acta Zool. Lilloana 21:418-420.
PARATYPES: LACM 1629 (4 dry), 1500 m, Rio Chotano, between Chiclayo and Cutervo,
Lambayeque Dept., Perú, Leg.: W. Weyrauch.
cavagnaroi, Naesiotus A.G. Smith, 1972: Proc. Calif. Acad. Sci., 4th ser., 39(2):12-17.
PARATYPE: LACM 1622 (1 dry), 884 m, top of Mt. Crocker, Isla Santa Cruz (Indefatigable),
Islas Galápagos, Leg.: Templeton Crocker, 10-11 MAY 1932; PARATYPES: LACM 1623 (2
dry), 7 km NE of Santa Rosa, Isla Santa Cruz (Indefatigable), Islas Galápagos, Leg.: A. DeRoy,
27 NOV 1966; PARATYPE: LACM 1624 (1 dry), 2 mi. W of Mt. Crocker, Isla Santa Cruz
(Indefatigable), Islas Galápagos, Leg.: A. & J. DeRoy, 10 MAY 1970; PARATYPE: LACM
1625 (1 dry), On top of Mt. Crocker, Isla Santa Cruz (Indefatigable), Islas Galápagos, Leg.:
Templeton Crocker, 10-11 MAY 1932.
Remarks: Currently in the genus Bulimulus.
celendinensis, Drymaeus Weyrauch, 1956a, Arch. Moll. 85(4/6):151-152.
PARATYPES: LACM: 2241 (3 dry), 2700 m, hills near Celendin, Cajamarca Dept., Perú, Leg.:
W. Weyrauch.
chiletensis, Scutalus (Scutalus) Weyrauch, 1967: Acta Zool. Lilloana 21:373-375.
PARATYPES: LACM 1633 (5 dry), 850 m, River Jequetepeque, mountains above Chilete,
Cajamarca Dept., Perú, Leg.: W. Weyrauch.
deroyi, Naesiotus A.G. Smith, 1972: Proc. Calif. Acad. Sci., 4th ser., 39(2):9-12.
PARATYPES: LACM 1621 (3 wet), 264 m, NW side of Isla Santa Cruz (= Indefatigable Id.),
Islas Galápagos, Leg.: A. DeRoy, 18 FEB 1964.
Remarks: Currently in the genus Bulimulus.
gibber, Drymaeus (Drymaeus) Haas, 1949: Fieldiana-Zoology 31(28):238-240.
PARATYPE: LACM 2046 (1 dry), 1650 m, Divisoria, Huanuco Dept., Perú, Leg.: J.M. Schunke,
5 AUG-10 SEP 1947.
haasi, Bostryx (Bostryx) Weyrauch, 1960: Arch. Moll. 89(1/3):33-34.
PARATYPES: LACM 2240 (5 dry), 2400 m, Matucana, Rio Rimac, Lima Dept., Perú, Leg.: W.
Weyrauch.
lachayensis, Scutalus (Scutalus) versicolor Weyrauch, 1967: Acta Zool. Lilloana 21:383-384.
PARATYPES: LACM 1631 (8 dry), 300 m, Lachay, near Chancay, Lima Dept., Perú, Leg.: W.
Weyrauch.
multivolvis, Bostryx (Geoceras) Pilsbry, 1944b: Nautilus 57(4):124.
PARATYPE: LACM 2053 (1 dry), 1900 M, Ninabamba, near Ayachucho, Rio Pampas,
Ayacucho Dept., Perú, Leg.: W. Weyrauch.
perforatus, Bulimulus (Ataxys) Haas, 1951: Fieldiana-Zoology 31(46):518
SYNTYPES: LACM 2238 (5 dry), 1900 m, Ninibamba, near Ayachucho, Rio Pampas, Ayacucho
Dept., Perú, Leg.: W. Weyrauch.
Remarks: Currently in the genus Bostryx.
perhirsutus, Rabdotus (Plicolumna) Miller, Christensen, & Roth in Smith, Miller, Christensen,
& Roth, 1990: Proc. Calif. Acad. Sci., 4th ser., 47(4):101-102.
PARATYPES: LACM 2757 (4 dry), 460 m, along Todos Santos-Cabo San Lucas Rd., 3.5 km N
of El Saucito, Baja California Sur, México, Leg.: C.C. Christensen, J.A. Christensen, & W.B.
Miller, 22 DEC 1973.
scalesiana, Naesiotus A.G. Smith, 1972: Proc. Calif. Acad. Sci., 4th ser., 39(2):17-22.
PARATYPES: LACM 1626 (8 dry), 244 m, Horneman Farm area, Isla Santa Cruz (=
Indefatigable Id.), Islas Galápagos, Leg.: A.G. Smith, 19 FEB 1964.
Remarks: Currently in the genus Bulimulus.
subcostatus, Bulimulus (Protoglyptus) Haas, 1948: Fieldiana-Zoology 31(23):190-192.
SYNTYPES: LACM 2237 (2 dry), 500-700 m, mountains near Jaen, Cajamarca Dept., Perú,
Leg.: W. Weyrauch.
Remarks: Currently in the genus Protoglyptus.
weyrauchi, Bostryx Pilsbry, 1944a: The Nautilus 57(3):87-88.
PARATYPES: LACM 2047 (3 dry), 2100 m, Ninabamba, Rio Pampas, Ayacucho Dept., Perú,
Leg.: W. Weyrauch.
Family Cerionidae Pilsbry, 1901
albolabra, Strophiops Maynard, 1921: Records of walks & talks with nature, App. 10:129.
SYNTYPES: LACM 2235 (5 dry), Great Harbor Cay, Berry Ids., Bahamas
Remarks: Currently in the genus Cerion.
fernandina, Cerion (Strophiops) Clench, 1937: The Nautilus 51(1):21-22.
PARATYPES: LACM 1304 (2 dry), 8 mi. SE of Simms, Long Id., Bahamas, Leg.: J.V. Malone,
OCT 1936.
hughesi, Cerion Clench, 1952: Revista Sociedad Malac. 8(3):107.
PARATYPES: LACM 1295 (4 dry), Sandy Pt., Savannah Sound, Eleuthera, Bahamas, Leg.: W.J.
Clench, MAY 1936.
huntingtoni, Cerion Clench, 1938: Bull. Mus. Comp. Zool. 80:526.
PARATYPES: LACM 1632 (2 dry), Columbus Pt., Cat Id., Bahamas, Leg.: W.J. Clench, Russell
& Huntington.
malonei, Cerion (Strophiops) Clench, 1937: The Nautilus 51(1):20-21.
PARATYPES: LACM 1303 (2 dry), 3.5 mi. SE of Simms, Long Id., Bahamas, Leg.: Russell,
McLean, Huntington & Foster, JUL 1936.
marielinum, Cerion Torre & Pilsbry, 1927: The Nautilus 40(3):74.
PARATYPES: LACM 1296 (6 dry), Mariel, Pinar del Rio Prov., Cuba, Leg.: M.L. Jaume.
mcleani, Cerion (Strophiops) Clench, 1937: The Nautilus 51(1):22.
PARATYPES: LACM 1305 (3 dry), 1 mi. E of O'Neills, Long Id., Bahamas, Leg.: Russell,
McLean, Huntington, & Foster, JUL 1936.
melanostomum, Cerion (Strophiops) Clench, 1934: Proc. Boston Soc. Nat. Hist. 40(2):212-213.
PARATYPES: LACM 2022 (3 dry), S end of Long Id., Bahamas, Leg.: T. Barbour & Mr. & Mrs.
J.C. Greenway, 6 MAR 1934.
obtusa, Strophiops Maynard, 1921: Records of walks & talks with nature, App. 10:132.
SYNTYPES: LACM 2236 (2 dry), Cat Cay, Bimini Group, Bahamas.
Remarks: Currently in the genus Cerion.
salvatori, Cerion Torre & Pilsbry, 1927: The Nautilus 40(3):74.
PARATYPES: LACM 1297 (6 dry), Playa de Santa Fe, La Habana Prov., Cuba, Leg.: M.L.
Jaume.
Family Urocoptidae Pilsbry, 1898
alamellata, Holospira (Sonoraloa) dentaxis Gilbertson, 1993: Amer. Malacological Bull.
10(1):73-75.
PARATYPES: LACM 2246 (4 dry), South side of Highway 15, 7.5 road km W of Sahuaripa,
Sonora, México (29 01.5 N, 109 18.7 W).
alleni, Urocoptis Torre, 1929a: The Nautilus 42(4):141 [pl. 4, fig. 7].
SYNTYPES: LACM 1298 (17 dry), Pena Blanca, Sierra Anafe, Pinar del Rio Prov., Cuba,
Leg.: M.L. Jaume.
Remarks: Currently in the genus Tetrentodon.
animasensis, Holospira Gilbertson & Worthington, 2003, The Veliger 46(3):220-224, figs. 1-4.
HOLOTYPE: LACM 2917 (dry); PARATYPES: LACM 2953 (5 dry), NW-SE trending
limestone ridge, north end of Animas Mountains, 11 air km SE of Animas, Hidalgo Co., New
Mexico (31°53.0'N, 108°47.70'W), Leg.: R.D. Worthington, 5 APR 1998.
barbouri, Urocoptis livida Torre & Clench, 1930: The Nautilus 44(1):15-16.
PARATYPES: LACM 1300 (4 dry), La Portuguesa, Central Soledad, Cienfuegos Prov., Cuba.
barroi, Tetrentodon (Cylindrocoptis) Jaume & Torre, 1972, Circ. Museo Biblio. Zool. Habana,
p. 17.
PARATYPES: LACM 2613 (10 dry), north side of Sitio Rd. near Jaruco, La Habana Prov.,
Cuba, Leg.: M.L. Jaume.
cuestai, Urocoptis Torre, 1929b: Proc. Acad. Nat. Sci. Philadelphia 81:445.
SYNTYPES [as COTYPES]: LACM 1351 (3 dry), 14 km marker on the road from Pinar del Rio
to Vinales, Pinar del Rio Prov., Cuba.
fergusoni, Holospira Gilbertson & Naranjo-Garcia, 2010: Nautilus 124(4):181-184, fig. 1.
HOLOTYPE: LACM 3112 (dry), 3600 m elevation, 0.5 km E of summit of Sierra La Viga,
Municipio Arteaga, Coahuila, México (25 21 35″N, 100 33 15″W), Leg.: G.M. Ferguson, 22
MAY 1986.
fortiuscula, Urocoptis (Gongylostoma) Torre, 1912: The Nautilus 26(4):57.
PARATYPES: LACM 1302 (3 dry), "El Mamey," Sagua la Grande, Santa Clara, Villa Clara
Prov., Cuba, Leg.: M.L. Jaume.
hubbardi, Callocoptis Jaume & Torre, 1972: Circ. Museo Biblio. Zool. Habana, p. 86-87.
PARATYPES: LACM 2614 (3 dry), El Cejanal, Vinales Pinan del Rio Prov., Cuba, Leg.: M.L.
Jaume.
lermondi, Tetrentodon Jaume & Torre, 1972: Circ. Museo Biblio. Zool. Habana, p. 12.
PARATYPES: LACM 2615 (4 dry), Hills near Sierra del Grillo, Madruga, La Habana Prov.,
Cuba, Leg.: M.L. Jaume.
mellita, Urocoptis Torre, 1932: The Nautilus 45(3):88-89.
PARATYPES: LACM 1217 (10 dry), "Finca El Ingles," Sierra del Grillo, Madruga, La Habana
Prov., Cuba, Leg.: M.L. Jaume.
mendezi, Tetrentodon Jaume & Torre, 1972: Circ. Museo Biblio. Zool. Habana, p. 14.
PARATYPES: LACM 2617 (6 dry), Sierra del Grillo, Aguacaste, La Habana Prov., Cuba, Leg.:
M.L. Jaume.
milleri, Holospira (Millerella) Gilbertson & Naranjo-Garcia, 1998: The Veliger 41(4):315-318.
PARATYPES: LACM 2805 (3 dry), 20.5 km E of the Rio Yaqui Bridge along Highway 15 from
La Estrella to Bacanora, Sierra Batamonte (near El Milagro Mine), Sonora, México (28°57.5'N,
109°30.50'W), 3 JAN 1988.
nataliae, Torrecoptis Jaume & Torre, 1972: Circ. Museo Biblio. Zool. Habana, p. 28.
PARATYPES: LACM 2616 (4 dry), Loma Castellanos, near Calabazar de Sagua, Villa Clara
Prov., Cuba, Leg.: M.L. Jaume.
nodulifera, Urocoptis Torre, 1929b: Proc. Acad. Nat. Sci. Philadelphia 81:444-445.
PARATYPES: LACM 1352 (4 dry), Dos Hermanos marker, Vinales, Pinar del Rio Prov., Cuba.
sherbrookei, Holospira (Holospira) Gilbertson, 1989: The Veliger 32(3):308-312.
PARATYPE: LACM 2209 (1 dry), Silver Creek Canyon, 3.4 mi. W of Portal, along road to
Paradise, Chiricahua Mountains, Cochise Co., Arizona, Leg.: L.H. Gilbertson, 16 APR 1987.
sinistra, Urocoptis Torre, 1928: The Nautilus 42(1): [pl. 1, fig. 8].
PARATYPES: LACM 1301 (19 dry), near Playa Marianao, La Habana Prov., Cuba, Leg.: M.L.
Jaume.
Superfamily STREPTAXOIDEA Gray, 1860
Family Streptaxidae Gray, 1860
cookeana, Streptaxis F. Baker, 1914: Proc. Acad. Nat. Sci. Philadelphia 65:628-629.
SYNTYPES: LACM 2234 (2 dry), Ceara Mirim, Rio Grande do Norte State, Brazil.
Superfamily RHYTIDOIDEA PIlsbry, 1893
Family Haplotremidae H.B. Baker, 1925
zopherum, Ancotrema Roth, 1990: Wasmann Jour. Biol. 47(1-2):68-71
PARATYPES: LACM 2430 (3 dry), Sutter Creek Cave, Amador Co., California, Leg.: R.
deSaussure, 25 MAY 1953.
Family Scolodontidae H.B. Baker, 1925
ceronitens, Systrophia Haas, 1951: Fieldiana-Zoology 31(46):533-534.
PARATYPES: LACM 2045 (4 dry), 1200 m, S of San Ignacio and W of Rio Chinchipe,
Cajamarca Dept., Perú, Leg.: W.Weyrauch.
platysma, Systrophia Haas, 1951: Fieldiana-Zoology 31(46):530-531.
PARATYPES: LACM 2044 (4 dry), 1400 m, Huarango, E side of Rio Rinchipe, close to Rio
Maranon, Amazonas Dept., Perú, Leg.: W.Weyrauch.
Superfamily ACAVOIDEA Pilsbry, 1895
Family Megomphicidae H.B. Baker, 1930
allyni, Ammonitella yatesi Chace, 1951: The Nautilus 64(4):122.
PARATYPES: LACM 1644 (14 dry), near Boyden's Cave, Fresno Co., California, Leg.: E.P.
Chace, 5 SEP 1950.
californicus, Megomphix A.G. Smith, 1960: Occas. Pap. Calif. Acad. Sci. 28:1-3.
PARATYPES: LACM 1679 (2 dry), Natural Bridge Cave, Trinity Co., California, Leg.: R. de
Soussure, 23 AUG 1958.
Superfamily PUNCTOIDEA Morse, 1864
Family Punctidae Morse, 1864
hannai, Punctum (Toltecia) Roth, 1985b: Malac. Review 18:52-55.
PARATYPES: LACM 2174 (2 dry), 914 m, W side of stream 0.5-0.75 mi. N of Trinity Alps
Camp, Trinity River, Trinity Co., California, Leg.: G D. Hanna, JUN 1931.
pasadenae, Punctum conspectum Pilsbry, 1896a: The Nautilus 10(2):21-22.
PARATYPES: LACM 1676 (50 dry), Pasadena, Los Angeles Co., California.
Remarks: Junior synonym of Paraloma servilis (Shuttleworth, 1852) fide Roth & Sadeghian
(2003).
Family Helicodiscidae H.B. Baker, 1927
spirellum, Speleodiscoides A.G. Smith, 1957: Proc. Calif. Acad. Sci., 4th ser., 29(2):34-37.
PARATYPES: LACM 1111 (3 dry); PARATYPES: LACM 1668 (3 dry), Violin Cave, Amador
Co., California, Leg.: E. Danehy & A. Lang, 22 NOV 1951.
Family Discidae Thiele, 1931
eyerdami, Anguispira (Zonodiscus) kochi Clench & Banks, 1939: Mem. Sociedad Cubana Hist.
Nat. 13(5):285.
PARATYPES: LACM 1645 (3 dry), ca. 20 mi. W of Santus Pass, Horscleawalls, Yakima Co.,
Washington, Leg.: W.J. Eyerdam, 31 MAY 1938.
Remarks: Junior synonym of Anguispira kochi occidentalis (von Martens, 1882).
Family Oreohelicidae Pilsbry, 1939
almoloya, Oreohelix Drake, 1949: The Nautilus 62(4):110-112.
PARATYPES: LACM 1658 (2 dry), foothills of Sierra de Almoloya, 5 mi. NW of Cueva Diablo,
Chihuahua, México, Leg.: L. Pierson, 18 AUG 1947.
anchana, Oreohelix Gregg, 1953: Bull. So. Calif. Acad. Sci. 52(2):71-74, pl. 14 (upper figs.).
HOLOTYPE: LACM 1450 (dry), 2194 m, NE slope Center Mountain, below cave, Sierra Ancha,
Gila Co., Arizona, Leg.: W.O. Gregg & W.L. Walton, JUN 1942; PARATYPES: LACM 1683
(45 dry), 2194 m, NW slope of Center Mountain, Sierra Ancha, Gila Co., Arizona, Leg.: W.O.
Gregg & W.L. Walton, 10 OCT 1949.
Remarks: Holotype ex W.O. Gregg collection 6142.
californica, Oreohelix Berry, 1931b: Ann. Mag. Nat. Hist., 10th ser., 8:115-117.
PARATYPES: LACM 1659 (4 dry), head of gulch on W slope of Clark Mountain, San
Bernardino Co., California, Leg.: E.C. Jaeger, 14 OCT 1930.
grahamensis, Oreohelix Gregg & Miller, 1974: Bull. So. Calif. Acad. Sci. 73(3):150-151, fig. 4.
HOLOTYPE: LACM 1155 (dry), S slope of Mt. Graham, Pinaleno Mountains, Graham Co.,
Arizona, Leg.: M.L. Walton & W.O. Gregg, 21 APR 1954; PARATYPES: LACM 1691 (9 dry),
2743 m, on S slope of Mt. Graham, 20.7 mi. from junction with U.S. Highway 666, Graham
Mountains, Graham Co., Arizona, Leg.: M.L. Walton & W.O. Gregg, 21 APR 1954.
hachetana, Oreohelix Pilsbry, 1915: Proc. Acad. Nat. Sci. Philadelphia 67:330-332.
PARATYPE: LACM 1680 (1 dry), 2550 m, summit of Hacheta Grande Mountain, Hidalgo Co.,
New Mexico, Leg.: H.A. Pilsbry & Daniels.
Remarks: Currently in the genus Radiocentrum. Ex W.O. Gregg collection 7856.
parawanensis, Oreohelix Gregg, 1940: The Nautilus 54(3):95-96.
PARATYPES: LACM 1660 (19 dry), Brian Head, Parowan Mountains, Iron Co., Utah.
profundorum, Oreohelix yavapai Pilsbry & Ferriss, 1911: Proc. Acad. Nat. Sci. Philadelphia
63:182-184.
PARATYPES: LACM 1677 (3 dry), 1737 m, Spectacle Cave, near Bass Camp, Grand Canyon
National Park, Coconino Co., Arizona, 22 NOV 1909.
“Limacoid clade”
Superfamily HELICOIDEA Rafinesque, 1815
Family Bradybaenidae Pilsbry, 1934
rustica, Cochlostyla metaformis Mousson, 1849: Die land-und Susswasser-Mollusken von Java,
p. 114.
PARATYPE: LACM 2619 (1 dry), Sibal, Luzon Id., Philippines.
Remarks: Currently in the genus/subgenus Helicostyla (Dryochchlias).
tagolandensis, Helicostyla leucopthalma Pilsbry, 1908: The Nautilus 22(4/5):47.
SYNTYPE: LACM 1294 (1 dry); PARATYPE: LACM 2043 (1 dry), Tahulandang Id.,
Kepulauan Sangihe, Celebes, Indonesia.
Family Polygyridae Pilsbry, 1895
animorum, Ashmunella tetrodon Pilsbry & Ferriss, 1917: Proc. Acad. Nat. Sci. Philadelphia
69:89-90.
PARATYPES: LACM 1648 (3 dry), 0.5 mi. S of Reed Cabin, Black Canyon, Black Range,
Sierra Co., New Mexico.
bequaerti, Ashmunella Clench & Miller, 1966: Brevoria, Mus. Comp. Zool. 244:1-6.
PARATYPES: LACM 1646 (11 dry), 1493 m, NE slope of Black Mountain, Davis Mountains,
Jeff Davis Co., Texas, Leg.: J.C. Bequaert & W.B. Miller, 6 MAY 1965.
dispar, Ashmunella danielsi Pilsbry & Ferriss, 1915: The Nautilus 29(4):41-42.
PARATYPES: LACM 1647 (3 dry), 2286 m, W slope of Mogollon Mountains, in Little
Whitewater Canyon, Socorro Co., New Mexico, Leg.: Ferris & Daniels, 2 DEC 1914.
hirsutus, Hochbergellus Roth & Miller, 1992: The Veliger 35(3):223-225
PARATYPE: LACM 2758 (1 dry), Sisters Rocks, ca. 3.8 km N of Euchre Creek at Ophir, Curry
Co., Oregon, Leg.: W.B. Miller, 16 JUL 1991.
karokorum, Vespericola Talmadge, 1962: The Veliger 5(1):28-29.
PARATYPES: LACM 1228 (2 dry), Sawmill Gulch, N side of Klamath River, 2 mi. E of
Orleans Gulch Ranger Station, Humboldt Co., California, Leg.: R. Talmadge, SEP 1957.
lenticula, Ashmunella Gregg, 1953: Bull. So. Calif. Acad. Sci. 52(2):74-75, pl. 14 (lower figs.).
HOLOTYPE: LACM 1452 (dry); PARATYPES: LACM 1091 (87 dry), 1463 m, rockslide at
mouth of N fork of Horseshoe Canyon, Chiricahua Mountains, Cochise Co., Arizona, Leg.: W.O.
Gregg & M.L. Walton, 13 OCT 1949.
Remarks: Holotype ex W.O. Gregg collection.
lombardii, Allogona A.G. Smith, 1943: Proc. Calif. Acad. Sci., 4th ser., 23(36):545-547.
PARATYPES: LACM 1221 (4 dry), 579 m, along meadow creek, 1.5 mi. S of Selway Falls,
Idaho Co., Idaho, Leg.: G D. Hanna, SEP 1911.
pasonis, Polygyra Drake, 1951a: Revista Sociedad Malac. 8(1):44-46.
PARATYPES: LACM 1661 (4 dry), foothills of Franklin Mountains, 5 mi. SE of the federal
penitentiary, La Tuna, El Paso Co., Texas, Leg.: J. & R. Drake, 30 OCT 1949.
Remarks: Currently in the genus Ashmunella.
populi, Polygyra Vanatta, 1924: Proc. Acad. Nat. Sci. Philadelphia 76:26.
PARATYPE: LACM 1662 (1 dry), Cottonwood Tree Canyon, on the Snake River, 50 mi. S of
Lewiston, Idaho Co., Idaho.
Remarks: Junior synonym of Cryptomastix mullani (Bland & Cooper, 1861).
porterae, Ashmunella thomsoniana Pilsbry & Cockerell, 1899: The Nautilus 13(5):49-50.
SYNTYPES: LACM 1229 (2 dry), upper Sapello Canyon, Beulah, San Miguel Co., New
Mexico, Leg.: W. Porter, 1889.
pressleyi, Vespericola Roth, 1985a: Wasmann Jour. Biol. 42(1/2):84-91.
PARATYPE: LACM 2121 (1 dry), 380 m, 0.2 km E of Price Creek mouth, S bank of the Trinity
River, Trinity Co., California, Leg.: B. Roth, 4 AUG 1983.
solida, Polygyra ptychophora Vanatta, 1924: Proc. Acad. Nat. Sci. Philadelphia 76:25.
PARATYPES: LACM 1682 (5 dry), Cottonwood Tree Canyon, 50 mi. S of Lewiston, Idaho Co.,
Idaho, Leg.: R. Jackson.
Remarks: Currently in the genus Allogona. Ex W.O. Gregg collection 4236.
Family Helminthoglyptidae Pilsbry, 1939
amboiana, Micrarionta hutsoni Willett, 1931a: The Nautilus 44(4):123-124, pl. 7, fig. 4.
HOLOTYPE: LACM 1029 (dry), 6 mi. NW of Amboy, San Bernardino Co., California,
Leg.: George Willett, 7 FEB 1931.
Remarks: Currently in the genus Eremarionta and currently a subspecies of E. rowelli
(Newcomb, 1865) fide Roth & Sadeghian (2003).
aguacalientensis, Sonorella walkeri Pilsbry & Ferriss, 1915: Proc. Acad. Nat. Sci. Philadelphia
75:396-397.
PARATYPES: LACM 1681 (3 dry), Agua Caliente Canyon, W slope of Santa Rita Mountains,
Santa Cruz Co., Arizona, Leg.: H.A. Pilsbry & J.H. Ferriss.
Remarks: Junior synonym of Sonorella walkeri Pilsbry & Ferriss, 1923. Ex W.O. Gregg
collection 2855.
allynsmithi, Sonorella Gregg & Miller, 1969: The Nautilus 82(3):90-93.
PARATYPES: LACM 1246 (9 dry), 457 m, Squaw Peak Rd., approx. 0.3 mi. from Lincoln
Drive, Phoenix Mountains, Maricopa Co., Arizona, Leg.: J.C. Bequaert, W.B. Miller & M.L.
Walton, 30 OCT 1965; PARATYPES: LACM 1663 (174 dry), S slope of the Phoenix
Mountains, back of the Arizona Biltmore Hotel, Maricopa Co., Arizona, Leg.: W.O. Gregg &
M.L. Walton, 23 APR 1954; PARATYPES: LACM 1664 (20 dry), S side of Squaw Peak,
Phoenix Mountains, near the reservoir, Maricopa Co., Arizona, Leg.: A.G. Smith, 1-3 APR 1953.
angelus, Sonorelix (Herpeteros) Gregg, 1949: Bull. So. Calif. Acad. Sci. 47(3):100-102.
HOLOTYPE: LACM 1173 (dry), hillsides on N side of W end Soledad Canyon, Los Angeles
Co., California, Leg.: W.O. Gregg, FEB-MAR 1947; PARATYPE: LACM 1085 (1 dry), 2 mi. E
of Solemint, N side of Soledad Canyon, Los Angeles Co., California, Leg.: W.O. Gregg, MAR
1947.
Remarks: Currently in the genus Hespeteros fide Roth & Sadeghian (2003). Holotype ex W.O.
Gregg collection 3692a.
benitoensis, Helminthoglypta Lowe, 1930: The Nautilus 44(2):43, pl. 5, fig. 1.
HOLOTYPE: LACM 1021 (dry), Pinnacles National Monument, San Benito Co., California,
Leg.: H.N. Lowe, JUL 1929.
beryllica, Monadenia fidelis Chace & Chace, 1935: The Nautilus 49(2):48-49.
PARATYPES: LACM 1225 (5 dry), 8 mi. N of Brookings, Curry Co., Oregon, Leg.: E.P. Chace,
1935.
brunnea, Micrarionta Willett, 1935a: The Nautilus 49(1):15.
HOLOTYPE: LACM 1042 (dry); PARATYPES: LACM 1357 (4 dry), Chuckwalla Spring, Little
Chuckwalla Mountains, Riverside Co., California. Leg.: G. & O. Willett, 21 FEB 1935.
Remarks: Currently in the genus Eremarionta fide Roth & Sadeghian (2003).
carrizoensis, Micrarionta harperi Willett, 1937b: The Nautilus 50(4):123.
HOLOTYPE: LACM 1049 (dry), hills above Painted Gorge, Carrizo Mountains, Imperial Co.,
California, Leg.: G. Willett, 1 DEC 1936.
Remarks: Currently a subspecies of Sonorelix harperi (Bryant, 1900) fide Roth & Sadeghian
(2003).
caruthersi, Helminthoglypta Willett, 1934b: Bull. So. Calif. Acad. Sci. 33(2):57-58, pl. 20.
HOLOTYPE: LACM 1039 (1 dry), 2133 m, Morris Canyon, Kern Co., California, Leg.: Mrs.
V.L. Carr.
Remarks: Currently in the subgenus Coyote fide Roth & Sadeghian (2003).
cathedralis, Micrarionta indioensis Willett, 1930b: The Nautilus 43(4):115-116.
HOLOTYPE: LACM 1022 (dry), Cathedral Canyon, Riverside Co., California, Leg.: G. Willett,
8 FEB 1930.
Remarks: Currently in the genus Cahuillus fide Roth & Sadeghian (2003).
chacei, Micrarionta Willett, 1940: Bull. So. Calif. Acad. Sci. 39(1):81-82, pl. 12.
HOLOTYPE: LACM 1062 (dry); PARATYPES: LACM 1093 (4 dry), 9 mi. N of Ensenada,
Pacific Coast Baja California, México, Leg.: G. Willett, 22 FEB 1937.
Remarks: Currently in the genus Sonorelix.
chocolata, Micrarionta Willett, 1935a: The Nautilus 49(1):15-16.
HOLOTYPE: LACM 1041 (dry), Beal's Well, Chocolate Mountains, Imperial Co., California,
Leg.: O.A. Willett, 19 FEB 1935.
Remarks: Currently in the genus Eremarionta and currently a subspecies. of E. rowelli
(Newcomb, 1865) fide Roth & Sadeghian (2003).
chuckwallana, Micrarionta Willett, 1935a: The Nautilus 49(1):15.
HOLOTYPE: LACM 1040 (dry), 1 mi. S of Corn Springs, Chuckwalla Mountains, Riverside
Co., California, Leg.: G. Willett, 6 FEB 1935.
Remarks: Currently in the genus Eremarionta and currently a subspecies of E. rowelli
(Newcomb, 1865) fide Roth & Sadegghian (2003).
concolor, Helminthoglypta (Coyote) Roth & Hochberg, 1988: The Veliger 31(3/4):258-261.
PARATYPE: LACM 2215 (1 dry), 1615 m, 10.3 road mi. E of cemetary, Tejon Canyon, Kern
Co., California, Leg.: W.B. Miller, F.G. Hochberg, & B. Roth, 7 MAR 1987.
cossi, Sonorella baboquivariensis Miller, 1966: The Nautilus 80(2):46-48.
PARATYPES: LACM 1244 (2 dry), Arch Canyon, Ajo Range, Organ Pipe Cactus National
Monument, Pima Co., Arizona, Leg.: W.B. Miller & J.C. Bequaert, 25 JAN 1966.
crotalina, Helminthoglypta Berry, 1928: Ann. Mag. Nat. Hist., 10th ser., 1:276-278.
PARATYPES: LACM 1678 (2 dry), Sidewinder Mine, N end of Granite Mountains, San
Bernardino Co., California, Leg.: Jaeger.
Remarks: Currently in the subgenus Coyote fide Roth & Sadeghian (2003). Ex W.O. Gregg
collection 839.
cuyama, Helminthoglypta Hanna & Smith, 1937: The Nautilus 51(1):15-16.
PARATYPES: LACM 1224 (6 dry), Cuyama River Valley, 23.7 mi. E of Santa Maria, Santa
Barbara Co., California, Leg.: G D. Hanna & C.C. Church.
edwardsi, Helminthoglypta Gregg & Miller, 1976: Bull. So. Calif. Acad. Sci. 75(1):13-16, figs.
2-3 (upper fig.).
HOLOTYPE: LACM 1761 (dry); PARATYPES: LACM 1108 (3 dry), 1128 m, along Manzanita
Trail near Manzanita Creek, Pine Valley, San Diego Co., California (32 49.7 N, 116 31.1 W),
Leg.: W.O. Gregg, 18 JAN 1953.
fontiphila, Helminthoglypta Gregg, 1931: The Nautilus 45(2):49-51, pl. 4, figs. 8-10.
HOLOTYPE: LACM 1032 (dry); PARATYPES: LACM 1650 (21 dry), 945 m, 0.5 mi. below the
dam in Little Rock Creek Canyon, Los Angeles Co., California, 26 MAR 1931.
Remarks: Currently in the subgenus Coyote fide Roth & Sadeghian (2003).
granitensis, Micrarionta Willett, 1935b: Bull. So. Calif. Acad. Sci. 34(1):1-2.
HOLOTYPE: LACM 1043 (dry), NW end of Granite Mountains, Riverside Co., California,
Leg.: O.A. Willett, 5 MAR 1935.
Remarks: Currently in the genus Eremarionta and currently a subspecies of E. rowelli
(Newcomb, 1865) fide Roth & Sadeghian (2003).
greggi, Helminthoglypta Willett, 1931a: The Nautilus 44(4):124-125, pl. 7, fig. 3
HOLOTYPE: LACM 1031 (1 dry), 3.5 mi. S of Mojave, Kern Co., California, Leg.: G. Willett,
23 FEB 1931.
Remarks: Currently in the subgenus Coyote fide Roth & Sadeghian (2003).
greggi, Sonorella Miller, 1967a: The Nautilus 80(4):114-116.
PARATYPES: LACM 1665 (2 dry), 1920 m, Sierra Purica, Sonora, México, Leg.: W.B. Miller,
29 JUL 1965.
hausabasensis, Sonorella mormonum Miller, 1967b: The Nautilus 81(1):2-4.
PARATYPE: LACM 1667 (1 dry), 1097 m, ca. 5 mi. E of Huasabas, Sonora, México, Leg.:
W.B. Miller, 1 SEP 1965.
Remarks: Currently in the genus Monadenia.
hilli, Micrarionta hutsoni Willett, 1930a: The Nautilus 44(1):6-7.
HOLOTYPE: LACM 1024 (dry), Sheep Hole Mountains, San Bernardino Co., California,
Leg.: G. Willett, 8 NOV 1929.
Remarks: Junior synonym of Eremarionta rowelli unifasciata (Willett, 1930) fide Roth &
Sadeghian (2003).
huecoensis, Sonorella Gilbertson & Metcalf, 2005: American Malacological Bulletin 20(1/2):
38-40, fig. 2.
HOLOTYPE: LACM 2965 (dry); PARATYPE: LACM 2966 (anatomy), Rimrock escarpment,
Hueco Mountains, El Paso Co., Texas (31˚58.24’N, 106˚2.54’W), Leg.: A.L. Metcalf, 2 NOV
1987; PARATYPES: LACM 2967 (21 dry), Rimrock escarpment, Hueco Mountains, El Paso
Co., Texas, Leg.: A.L. Metcalf, 17 JAN 1976; PARATYPES: LACM 2968 (8 dry), Rimrock
escarpment, Hueco Mountains, EL Paso Co., Texas, Leg.: T. Dillon & A.L. Metcalf, 12 AUG
1989.
Remarks: Paratype anatomies mounted on glass slides.
imitator, Sonorella Gregg & Miller, 1974: Bull. So. Calif. Acad. Sci. 73(3):148-150, fig. 2.
HOLOTYPE: LACM 1154 (dry), S slope of Graham Mountain, Pinalento Mountains, Graham
Co., Arizona, Leg.: M.L. Walton & W.O. Gregg, 21 APR 1954; PARATYPES: LACM 1692 (5
dry), 2743 m, N of Swift Trail, 20.7 mi. from U.S. Highway 666, on S slope of Graham
Mountain, Graham Co., Arizona, Leg.: M.L. Walton & W.O. Gregg, 21 APR 1954.
immaculata, Micrarionta Willett, 1937a: Bull. So. Calif. Acad. Sci. 36(1):6-7, pl. 1, figs. a-c.
HOLOTYPE: LACM 1051 (dry), 7 mi. S of Vidal, E side of Riverside Mountains, Riverside Co.,
California, Leg.: G. & O. Willett, 24 MAR 1937.
Remarks: Currently in the genus Eremarionta fide Roth & Sadeghian (2003).
intersessa, Helminthoglypta Roth, 1986: Wasmann Jour. Biol. 44(1-2):57-65.
PARATYPE: LACM 2242 (1 dry), 7 m, coastal bluffs immediately N of Pudding Creek mouth,
Mendocino Co., California, Leg.: B. Roth, 15 OCT 1972.
mariposa, Monadenia (Corynadenia) hillebrandi A.G. Smith, 1957: Proc. Calif. Acad. Sci., 4th
ser., 29(2):24-26.
PARATYPES: LACM 1112 (2 dry), McLean Cave, Mariposa Co., California, Leg.: R. de
Saussure, 16 NOV 1953.
Remarks: Currently a valid species fide Roth & Sadeghian (2003).
mattolensis, Helminthoglypta arrosa A.G. Smith, 1938: The Nautilus 51(3):83-85.
PARATYPES: LACM 1222 (3 dry), between Cape Mendocino & the Mattole River mouth,
Humboldt Co., California, Leg.: G D. Hanna, 13 JUN 1929.
Remarks: Currently a subspecies of Helminthoglypta expansilabris A.G. Smith, 1938 fide Roth
& Sadeghian (2003).
mccoiana, Micrarionta Willett, 1935b: Bull So. Calif. Acad. Sci. 34(1):2.
HOLOTYPE: LACM 1044 (dry), McCoy Well, McCoy Mountains, Riverside Co., California,
Leg.: G. Willett, 6 MAR 1935.
Remarks: Currently in the genus Eremarionta and currently a subspecies E. rowelli (Newcomb,
1865) fide Roth & Sadeghian (2003).
meadi, Sonorella Miller, 1966: The Nautilus 80(2):50-52.
PARATYPE: LACM 1666 (1 dry), 488 m, Agua Dulce Mountains, Pima Co., Arizona, Leg.:
W.B. Miller, 29 JAN 1966.
micrometalleoides, Helminthoglypta Miller, 1970: The Veliger 12(3):275-278.
PARATYPES: LACM 1651 (3 dry), 3 mi. from the junction of Iron Canyon Rd. and the
Garlock-Goler Highway, El Paso Mountains, Kern Co., California, Leg.: W.B. Miller.
Remarks: Currently in the subgenus Coyote fide Roth & Sadeghian (2003).
mina, Helminthoglypta (Helminthoglypta) salviae Roth, 1987: The Veliger 30(2):186-187.
PARATYPE: LACM 2221 (1 dry), 1524 m, gully N of Cuddy Canyon Rd., 1.8 mi. (2.9 km) W
of Frazier Park, Ventura Co., California, Leg.: W.O. Gregg, 23 MAR 1947.
Remarks: Currently in the genus Charodotes fide Roth & Sadeghian (2003).
miwoka, Epiphragmophora californiensis Bartsch, 1919: Proc. Biol. Soc. Washington 32:248.
PARATYPES: LACM 1334 (4 dry), Pt. Reyes, Marin Co., California, Leg.: A.G. Smith, 28 APR
1918.
Remarks: Currently in the genus Helminthoglypta and currently a subspecies of H. stiversiana
(Cooper, 1875) fide Roth & Sadeghian (2003).
neglecta, Sonorella Gregg, 1951: Bull. So. Calif. Acad. Sci. 50(3):156-158, pl. 52.
HOLOTYPE: LACM 1451 (dry); PARATYPES: LACM 1087 (195 dry), 1615 m, N of the Portal
to Paradise Rd., 3 mi. W of Portal, Chiricahua Mountains, Cochise Co., Arizona, Leg.: W.O.
Gregg & M.L. Walton, 13 OCT 1949.
Remarks: Holotype ex W.O. Gregg collection 5317.
opuntia, Micrarionta (Micrarionta) Roth, 1975a: Bull. So. Calif. Acad. Sci. 74(2):94-96.
PARATYPE: LACM 1773 (1 dry), NE San Nicolas Id., California Channel Ids., California,
Leg.: Larson, Regal, & Anderson, APR 1974.
Remarks: Currently in the subgenus Nicolenea fide Roth & Sadeghian (2003).
ora, Micrarionta Willett, 1929a: Bull. So. Calif. Acad. Sci. 28(2):17-19, pl. 6.
HOLOTYPE: LACM 1018 (dry), N end of Fish Mountains, Imperial Co., California, Leg.: G.
Willett, 31 MAR 1929.
Remarks: Currently in the genus Sonorelix and currently a subspecies of S. borregoensis
(Berry, 1929) fide Roth & Sadeghian (2003). Cited as missing by Roth & Sadeghian (2003), but
has since been located.
orocopia, Micrarionta Willett, 1939a: Bull. So. Calif. Acad. Sci. 38(1):15, pl. 2, figs. a-c.
HOLOTYPE: LACM 1060 (dry), 2.5 mi. N of Dos Palmas, Orocopia Mountains, Riverside Co.,
California, Leg.: G. Willett, 1 JAN 1939.
Remarks: Currently in the genus Eremarionta fide Roth & Sadeghian (2003).
pacoimensis, Helminthoglypta traski Gregg, 1931: The Nautilus 45(2):48-49, pl. 4, figs. 5-7.
HOLOTYPE: LACM 1033 (dry); PARATYPES: LACM 1653 (3 dry), Pacoima Canyon, San
Gabriel Mountains, Los Angeles Co., California, Leg.: W.O. Gregg, 21 FEB 1931.
Remarks: Currently in the subgenus Charodotes fide Roth & Sadeghian (2003).
painter, Sonorella Lang & Gilbertson, 2010: Proc. Biol. Soc. Washington 123(1):64-70, fig. 2.
PARATYPES: LACM 3073 (5 dry), PARATYPE: LACM 3086 (1 dry, coated), 1868.9 m,
northeastern reach of Lang Canyon, San Luis Mtns., Hidalgo Co., New Mexico (31°20.29'N,
108°45.33'W). Leg.: B.K. Lang & D.C. Rogers, 22 SEP 2000.
pedregosensis, Sonorella Gilbertson & Radke, 2006: Amer. Malac. Bull 21(1/2):18-22, figs. 1-4.
HOLOTYPE: LACM 3046; PARATYPE: LACM 3047, 1421 m elev., 26 km N of Douglas,
Leslie Canyon National Wildlife Refuge, Pedregosa Mtns., Cochise Co., Arizona (31°35.53'N,
109°30.40'W), Leg.: W.R. Radke, 10 AUG 202; PARATYPES: 1421 m elev., Leslie Canyon
National Wildlife Refuge, Pedregosa Mtns., Cochise Co., Arizona (31°35.53'N, 109°30.40'W),
Leg.: L. Gilbetson, 10 APR 2001.
Remarks: Anatomies mounted on glass slides.
piutensis, Helminthoglypta cuyamacensis Willett, 1938b: Bull. So. Calif. Acad. Sci. 37(2):
53-54, pl. 12.
HOLOTYPE: LACM 1056 (dry), 2133 m, Piute Mountains, Kern Co., California, Leg.: G.
Willett, 3 JUL 1938.
Remarks: Identified as Helminthoglypta piutensis by Roth & Sadeghian (2003).
pronotis, Monadenia fidelis Berry, 1931a: The Nautilus 44(4):122-123.
PARATYPES: LACM 1226 (6 dry), Pt. St. George, 3 mi. N of Crescent City, Del Norte Co.,
California, Leg.: E.P. Chace, 25 AUG 1929.
reediana, Helminthoglypta Willett, 1932b: The Nautilus 45(4):134-135, pl. 11, figs. 8-10.
HOLOTYPE: LACM 1030 (dry), Lowe Canyon, Monterey Co., California, Leg.: G. Willett, 31
MAR 1931.
Remarks: Currently in the genus Charodotes fide Roth & Sadeghian (2003).
oremota, Micrarionta indioensis Willett, 1937b: The Nautilus 50(4):124-125.
HOLOTYPE: LACM 1050 (dry), Borrego Mountains, San Diego Co., California, Leg.: G.
Willett, 14 FEB 1937.
Remarks: Currently in the genus Cahuillus fide Roth & Sadeghian (2003) as Cahuillus
indioensis remotus (Willett, 1937).
rex, Helminthoglypta tudiculata Church & Smith, 1938: The Nautilus 51(4):119-122.
PARATYPE: LACM 1227 (1 dry), middle fork Tule River, 2 mi. above Springville, Tulare Co.,
California, Leg.: C.C. Church, JUL 1933.
Remarks: Currently a subspecies of Helminthoglypta cypreophila (Binney & Bland, 1869) fide
Roth & Sadeghian (2003).
salviae salviae, Helminthoglypta (Helminthoglypta) Roth, 1987: The Veliger 30(2):184-186.
PARATYPES: LACM 2219 (3 dry), 1280 m, Apache Canyon, 4 km WSW of Nettle Spring
Campground and 10 km E of State Highway 33, Ventura Co., California, Leg.: W.B. Miller &
others, 22 APR 1986.
Remarks: Currently in the genus Charodotes fide Roth & Sadeghian (2003).
similans, Helminthoglypta Hanna & Smith, 1937: The Nautilus 51(1):13-15.
PARATYPES: LACM 1223 (2 dry), 3.75 mi. SE of Oil City, Fresno Co., California, Leg.: G D.
Hanna & C.C. Church.
Remarks: Currently in the subgenus Charodotes fide Roth & Sadeghian (2003).
simmonsi, Sonorella Miller, 1966: The Nautilus 80(2):48-50.
PARATYPES: LACM 1245 (3 dry), W side of canyon that trends SW from Newman Peak,
Picacho Mountains, Pinal Co., Arizona, Leg.: J.C. Bequaert, M.L. Walton, & W.B. Miller, 27
DEC 1965.
stageri, Helminthoglypta Willett, 1938b: Bull. So. Calif. Acad. Sci. 37(2):52-53, pl. 11.
HOLOTYPE: LACM 1055 (dry), 1676 m, Erskine Creek, Piute Mountains, Kern Co., California,
Leg.: G. Willett, 29 MAY 1938.
taylori, Helminthoglypta (Coyote) Reeder & Roth, 1988: The Veliger 31(3/4):254-256.
PARATYPES: LACM 2222 (5 dry), 960 m, at junction of Cedar Springs Rd. and Summit Valley
Rd., W fork of Mojave River, San Bernardino Co., California, Leg.: W.B. Miller, 9 MAY 1963.
taylori, Humboldtiana Drake, 1951b: Revista Sociedad Malac. 8(2):93-96.
PARATYPE: LACM 1675 (1 dry), surface of cultural deposits, Site C96, Coahuila, México,
Leg.: W.W. Taylor, OCT 1947.
thermimontis, Helminthoglypta Berry, 1953a: Trans. San Diego Soc. Nat. Hist. 11(12):333-336.
PARATYPES: LACM 1652 (3 dry), 1524-1676 m, SE side of Hot Springs Mountain, San Diego
Co., California, Leg.: S.S. Berry & W.G. Craig, 5 APR 1928.
tryoni, Helix Newcomb, 1864: Proc. Calif. Acad. Sci., 1st ser., 3:116.
PARATYPES: LACM 1962 (21 dry), San Nicolas Id., California Channel Ids., California
Remarks: Currently in the genus Xerarionta fide Roth & Sadeghian (2003).
unifasciata, Micrarionta hutsoni Willett, 1930a: The Nautilus 44(1):6.
HOLOTYPE: LACM 1023 (dry), Newberry Springs, San Bernardino Co., California, Leg.: G.
Willett, 11 FEB 1929.
Remarks: Currently in the genus Eremarionta and currently a subspecies of E. rowelli
(Newcomb, 1865) fide Roth & Sadeghian (2003).
walkeriana, Helminthoglypta Hemphill, 1911: Trans. San Diego Soc. Nat. Hist. 1:102.
PARATYPES: LACM 1684 (2 dry), Near Morro Bay, San Luis Obispo Co., California, Leg.: H.
Hemphill.
Remarks: Currently in the genus Charodontes fide Roth & Sadeghian (2003). Ex W.O. Gregg
collection 7539.
waltoni, Helminthoglypta Gregg & Miller, 1976: Bull. So. Calif. Acad. Sci. 75(1):10-13, figs. 1,
3 (lower fig.).
HOLOTYPE: LACM 1760 (dry), 1372 m, in small ravine on E side of Scove Canyon Rd.,
Laguna Mountains, San Diego Co., California (32 50.3 N, 116 29.3 W), Leg.: M.L. Walton &
W.O. Gregg, 28 MAR 1947; PARATYPE: LACM 1107 (1 dry), 1372 m, NE side Laguna
Mountain Rd., 2.8 mi. NE of Laguna Junction, Laguna Mountains., San Diego Co., California,
Leg.: M.L. Walton & W.O. Gregg, 28 MAR 1947.
waltoni, Sonorella Miller, 1968: The Nautilus 82(2):61-63.
PARATYPES: LACM 1247 (4 dry), 0.5 mi. S canyon mouth W side of wash, West Doubtful
Canyon, Peloncillo Mountains, Cochise Co., Arizona, Leg.: W.B. Miller, J.C. Bequaert, & M.L.
Walton, 23 OCT 1966.
williamsi, Helminthoglypta arrosa A.G. Smith, 1938: The Nautilus 51(3):79-81.
PARATYPES: LACM 1220 (4 dry), Hog Id., Tomales Bay, Marin Co., California, Leg.: W.
Williams, 1936.
Remarks: Currently a subspecies of Helminthoglypta stiversiana (Cooper, 1875) fide Roth &
Sadeghian (2003).
wintu, Monadenia (Shastelix) troglodytes Roth, 1981: Proc. Calif. Acad. Sci., 4th ser.,
42(15):396-397.
PARATYPE: LACM 2026 (1 dry), 62 mi. E of Brock Mountain Lookout, above Squaw Creek
arm Shasta Lake, Shasta Co., California, Leg.: R.E. Seib, 25 MAR 1978.
CLASS BIVALVIA Linnaeus, 1758
Subclass Protobranchia Pelseneer, 1889
Order Solemyoida Dall, 1889
Superfamily SOLEMYOIDEA Gray, 1840
Family Solemyidae Gray, 1840
reidi, Solemya Bernard, 1980: Venus 39(1):19-22.
PARATYPE: LACM 1722 (1 wet), 326 m, Texada Id., Strait of Georgia, Nanaimo Dist., British
Columbia, Canada (49 04 N, 124 33.2 W), Leg.: D.B. Quayle, R/V G.B. Reed, 24 JUN 1974.
Order Nuculoida Dall, 1889
Superfamily NUCULOIDEA Gray, 1824
Family Nuculidae Gray, 1824
beringiana, Acila (Trunacila) Slodkevich, 1967: Tertiary Acila of Sakhalin Id., p. 56.
PARATYPES: LACM 2635 (2 dry), 135 m, Sea of Okhotsk, W of Oktyabr'skiy, Kamchatka
Oblast, Russia (52 34 N, 154 59.08 E), Leg.: P. Ushakov, R/V Gagara, 4 AUG 1932
Remarks: Junior synonym of Acila (Truncacila) castrensis (Hinds, 1844) fide Coan & others
(2000).
interflucta, Nucula (Nucula) Marincovich, 1973: Nat. Hist. Mus. L.A. Co. Sci. Bull. 16:7-8, fig.
1.
HOLOTYPE: LACM 1581 (dry); PARATYPES: LACM 1582 (1968 dry), 0 m, intertidal, Punta
Morro, Iquique, Tarapaca Prov., Chile (20 13 S, 70 10.42 W), Leg.: L. Marincovich, AUG 1964.
keenae, Nucula (Lamellinucula) Bernard, 1983: Venus 41(4):253-256, figs. 2, 6.
HOLOTYPE: LACM 2027 (dry), 2000 m, Cascadia Abyssal Plain off Oregon, East Pacific
(44 39.1 N, 125 19.5 W), Leg.: Oregon State Univ., OCT 1962.
Remarks: Junior synonym of Nucula (Lamellinucula) carlottensis Dall, 1897 fide Coan & others
(2000).
leptodonta, Nucula (Leionucula) Bernard, 1983: Venus 41(4):255-257, figs. 4, 8.
HOLOTYPE: LACM 2029 (dry), 800 m, Cascadia Abyssal Plain off Oregon, East Pacific
(44 34.4 N, 125 59.3 W), Leg.: Oregon State Univ., JAN 1965.
Remarks: Junior synonym of Ennucula tenuis (Montagu, 1808) fide Coan & others (2000).
ovatotruncata, Leionucula Scarlato, 1981, Guide to the fauna of the USSR, p. 174.
PARATYPES: LACM 2638 (7 dry), 35 m, N of Ostrov Kozakevich, Zaliv Amur (= Amur Bay),
Sea of Japan, Primorye Kray, Russia (43 04 N, 131 50 E), Leg.: P. Schmidt, 1 MAR 1900.
Remarks: Junior synonym of Ennucula tenuis (Montagu, 1808) fide Coan & others (2000).
pigafettae, Nucula Dall, 1908: Bull. Mus. Comp. Zool. 43(6):369-370.
PARATYPES: LACM 1167 (3 dry), 1238 m, off Valdivia, Valdivia Prov., Chile (38 08 S,
81 00 W), Leg.: R/V Albatross.
Remarks: Currently in the subgenus Leionucula.
romboides, Leionucula inflata Scarlato, 1981: Guide to the fauna of the USSR, p. 173-174.
PARATYPES: LACM 2644 (40 dry), 25 m, N of Mys Shelagskiy, East Siberian Sea, Chukchi
Auton. Okrug, Russia (72 15 N, 167 38 E), Leg.: R/V Sibiryakov, 1932.
Remarks: Considered a valid species by Coan & others (2000).
takashii, Nucula (Lamellinucula) Bernard, 1983: Venus 41(4):255, figs. 3, 7.
HOLOTYPE: LACM 2028 (dry), 2000 m, Cascadia Abyssal Plain off Oregon, East Pacific
(44 39.1 N, 125 19.5 W), Leg.: Oregon State Univ., OCT 1962.
Remarks: Junior synonym of Nucula (Lamellinucula) charlottensis Dall, 1897 fide Coan &
others (2000).
yaquinae, Nucula (Leionucula) Bernard, 1983: Venus 41(4):257, figs. 5, 9.
HOLOTYPE: LACM 2030 (dry), 1540 m, Cascadia Abyssal Plain off Oregon, East Pacific
(45 50.1 N, 125 14 W), Leg.: Oregon State Univ., JUL 1967.
Remarks: Junior synonym of Ennucula cardara (Dall, 1916) fide Coan & others (2000).
Superfamily PRISTIGLOMOIDEA Sanders & Allen, 1973
Family Pristiglomidae Sanders & Allen, 1973
fragilis, Pseudoglomus Bernard, 1989c: Venus 48(2):67-68, fig. 1.
HOLOTYPE: LACM 2345 (dry), 400 m, off Lincoln Co., Oregon (44 31.5 N, 124 54 W)
Leg.: Oregon State Univ., 16 JUN 1964.
Superfamily NUCULANOIDEA H. Adams & A. Adams, 1853
Family Nuculanidae H. Adams & A. Adams, 1853
angusticauda, Nuculana (Nuculana) minuta Scarlato, 1981: Guide to the fauna of the USSR, p.
186.
PARATYPES: LACM 2634 (2 dry), 75 m, Tatarskiy Provliv, Sea of Japan, W of Sakhalin Id.,
Russia, Leg.: Z. Kobyakova; R/V Blukher, 13 JUL 1933.
Remarks: Junior synonym of Nuculana minuta (Müller, 1776) fide Coan & others (2000).
burchi, Nuculana Willett, 1944: Bull. So. Calif. Acad. Sci. 43(2):71, pl. 14, fig. 3.
HOLOTYPE: LACM 1066 (dry), PARATYPE: LACM 2139 (1 dry), 91 m, off Redondo Beach,
Los Angeles Co., California, Leg.: J.Q. & T.A. Burch.
Remarks: Junior synonym of Nuculana (Juperteria) cellulita (Dall, 1895) fide Coan & others
(2000). Paratype ex UCLA 20018.
exoptata, Leda Pilsbry & Lowe, 1932a: Proc. Acad. Nat. Sci. Philadelphia 84:107-108.
PARATYPES: LACM 2169 (4 dry), 37 m, Guaymas, Sonora, México, Leg.: H.N. Lowe
Remarks: Currently in the genus Adrana fide Keen (1971). Ex UCLA 3148.
habei, Robaia Scarlato, 1981: Guide to the fauna of the USSR, p. 188-189.
PARATYPE: LACM 2645 (1 dry), Zaliv Petra Velikogo (= Peter the Great Bay), Sea of Japan,
Primorye Kray, Russia, Leg.: K. Derjugin, 10 JUL 1932.
Remarks: Currently in the genus Nuculana.
impar, Leda Pilsbry & Lowe, 1932a: Proc. Acad. Nat. Sci. Philadelphia 84:106-107.
PARATYPES: LACM 2168 (4 dry), Bahía Concepción, Golfo de California, Baja California
Sur, México, Leg.: H.N. Lowe.
Remarks: Currently in the genus/subgenus Nuculana (Saccella) fide Keen (1971). Ex UCLA
3147.
laeviradius, Leda Pilsbry & Lowe, 1932a: Proc. Acad. Nat. Sci. Philadelphia 84:106.
PARATYPES: LACM 2167 (2 dry), 37 m, Guaymas, Sonora, México, Leg.: H.N. Lowe,
1929-1931.
Remarks: Junior synonym of Nuculana acrita (Dall, 1908) fide Keen (1971). Ex UCLA 3146.
pontonia, Leda Dall, 1890, Proc. USNM 12(773):257.
PARATYPES: LACM 2395 (6 dry),1503 m, off San Diego, San Diego Co., California, Leg.:
R/V Albatross.
Remarks: Currently in the genus/subgenus Nuculana (Jupeteria) fide Coan & others (2000).
redondoensis, Nuculana J.Q. Burch, 1944a: Min. Conch. Club So. Calif. 33:9-10.
PARATYPES: LACM 1316 (7 dry), Redondo Beach, Los Angeles Co., California, Leg.: J.Q.
Burch, 1944; PARATYPES: LACM 1696 (2 dry), 46 m, off Redondo Beach, Los Angeles Co.,
California, Leg.: J.Q. Burch, AUG 1941.
Remarks: Junior synonym of Nuculana (Jupetria) penderi (Dall & Bartsch, 1910) fide Coan &
others (2000).
robusta, Ledella Bernard, 1989c: Venus 48(2):68, fig. 2.
HOLOTYPE: LACM 2346 (dry), 4883 m, Tufts Abyssal Plain off Oregon, East Pacific
(44 53.6 N, 147 36.5 W), Leg.: Oregon State Univ., 14 AUG 1976.
spinuliformis, Ledella Bernard, 1989c: Venus 48(2):70, fig. 4.
HOLOTYPE: LACM 2337 (dry), 2800 m, off Lincoln Co., Oregon (44 40.9 N, 125 35.3 W),
Leg.: Oregon State Univ., 14 JUN 1964.
Remarks: Junior synonym of Ledellina convexirostrata (Filatova & Shileyko, 1984), fide Coan
& others (2000).
turnerae, Ledella Bernard, 1989c: Venus 48(2):68-70, fig. 3.
HOLOTYPE: LACM 2347 (dry), 3650 m, W of Patton Escarpment, California (31 12 N,
119 37.4 W), Leg.: Scripps Inst. Oceanog., 18 OCT 1965.
Remarks: Junior synonym of Ledella ultima (E.A. Smith, 1885) fide Coan & others (2000).
Family Malletiidae H. Adams & A. Adams, 1858
alata, Malletia (Malletiella) Bernard, 1989c: Venus 48(2):71-72, fig. 7.
HOLOTYPE: LACM 2343 (dry), 1200 m, San Diego Trough, California (32 25.8 N,
117 27.5 W), Leg.: Scripps Inst. Oceanog., 22 OCT 1971.
Family Yoldiidae Habe, 1977
beringiana, Yoldia Dall, 1916: Proc. USNM 52(2183):399.
PARATYPE: LACM 2394 (1 broken valve), 1805 m, Pribilof Ids., Bering Sea, Alaska, Leg.:
US Fish Commission.
Remarks: Junior synonym of Megayoldia montereyensis (Dall, 1893) fide Coan & others
(2000). Specimen broken into numerous pieces.
derjugini, Yoldiella Scarlato, 1981: Akad. Nauk SSSR Zool. Inst., Issledovaniia Fauny Morei
14(22):206-207.
PARATYPES: LACM 2639 (32 dry), 68 m, S of Mya Gamov, Zaliv Petra Velikogo (= Peter the
Great Bay), Sea of Japan, Primorye Kray, Russia (42 33 N, 131 13 E). Leg. Somov, SEP 1932.
Remarks: Incorrectly thought to be a junior homynym of Tindaria derjugini Gorbunov, 1946.
ensifera, Yoldia Dall, 1897: Bull. Nat. Hist. Soc. of Br. Columbia 2(1):9.
PARATYPES: LACM 1327 (7 dry), 68 m, Monterey Bay, Monterey Co., California, Leg.: R/V
Albatross.
Remarks: Junior synonym of Yoldia (Cnesterium) seminuda Dall, 1871 fide Coan & others
(2000).
montereyensis, Yoldia Dall, 1893: Nautilus 7(3):29-39.
PARATYPES: LACM 1125 (10 dry), 699 m, Monterey Bay, Monterey Co., California, Leg.:
R/V Albatross.
Remarks: Currently in the genus Megayoldia fide Coan & others (2000).
pseudonotabile, Yoldia (Cnesterium) keppeliana Scarlato, 1981: Guide to the fauna of the
USSR, p. 203.
PARATYPE: LACM 2636 (1 dry), 14 m, off Mya Sheveleva, Zaliv Ussuriyskiy (= Usuri Bay),
Sea of Japan, Primorye Kray, Russia (43 10 N, 132 21 E), Leg.: A. Razin, 14 AUG 1932;
PARATYPE: LACM 2652 (1 dry), 20 m, off Mya Sheveleva, Zaliv Ussuriyskiy (= Usuri Bay),
Sea of Japan, Primorye Kray, Russia (43 15 N, 132 19 E), Leg.: A. Razin, 8 JUN 1932.
Remarks: Junior synonym of Yoldia (Cnesterium) seminuda Dall, 1871.
toporoki, Yoldia (Cnesterium) Scarlato, 1981: Guide to the fauna of the USSR, p. 203-204.
PARATYPES: LACM 2632 (3 dry), 18-22 m, NE of Mys Terpeniya, Sakhalin Id., Sea of
Okhotsk, Russia (48 39 N, 144 45 E), Leg.: Brashnikov, R/V Storozh, 26 JUN 1899.
Family Siliculidae Allen & Sanders, 1973
alleni, Silicula Bernard, 1989c: Venus 48(2):70-71, fig. 5.
HOLOTYPE: LACM 2344 (dry), 3650 m, W of Patton Escarpment, California (31 12 N,
119 37 W), Leg.: Scripps Inst. Oceanog.
sandersi, Lamellileda Bernard, 1989c: Venus 48(2):71, fig. 6.
HOLOTYPE: LACM 2348 (dry), 5180 m, Tufts Abyssal Plain off Oregon, East Pacific
(45 02.2 N, 153 51.5 W), Leg.: Oregon State Univ., 19 AUG 1976.
Remarks: Currently in the genus Silicula fide Coan & others (2000).
Family Tindariidae Verrill & Bush, 1897
brunnea, Tindaria Dall, 1916: Proc. USNM 52(2183):401.
PARATYPES: LACM 1161 (131 dry), 2562 m, midway between St. Paul & Umnak Ids.,
Aleutian Ids. Co., Alaska, Leg.: R/V Albatross.
Remarks: Currently in the genus Neilonella (Family Neilonellidae) fide Coan & others (2000).
Subclass Pteriomorphia Beurlen, 1944
Order Arcoida Stoliczka, 1871
Superfamily ARCOIDEA Lamarck, 1809
Family Arcidae Lamarck, 1809
aleutica, Samacar (Pseudoporterius) Kamenev, 2007: Jour. Conch. 39(3):317-319, figs. 12-13.
HOLOTYPE: LACM 3067 (dry); PARATYPE: LACM 3068 (1 dry, gold coated), Western
Aleutian Ids, Aleutians West Co., Alaska, Leg.: R. Baxter, SEP 1986; PARATYPE: LACM 3069
(1 dry), 88 m, SSW of Cape Lutke, off SW end of Unimak Id., Aleutians East Co., Alaska
(54º24.13'N, 165º37.78'W), Leg.: R. Baxter, 24 JUL 1986
baughmani, Anadara Hertlein, 1951: Texas Jour. Sci. 3(3):487-489.
PARATYPE: LACM 2058 (1 dry), 91 m, off Aransas Pass, Aransas Co., Texas
Superfamily GLYCYMERIDOIDEA Newton, 1916
Family Glycymerididae Newton, 1916
keenae, Glycymeris Willett, 1943: Bull. So. Calif. Acad. Sci. 42(3):114, pl. 12, figs. 4-7.
HOLOTYPE: LACM 1065 (dry); PARATYPES: LACM 2137 (5 dry), 37-55 m, Forrester Id.,
Prince of Wales Co., Alaska, Leg.: G. Willett.
Remarks: Currently in the subgenus Axinola fide Coan & others (2000). Paratypes ex UCLA
20055.
Superfamily LIMOPSIDAE Dall, 1895
Family Limopsidae Dall, 1895
diegensis, Limopsis Dall, 1908: Bull. Mus. Comp. Zool. 43(6):395.
PARATYPES: LACM 1166 (61 dry), 146 m, off San Diego, San Diego Co., California, Leg.:
R/V Albatross.
Remarks: Junior synonym of Limopsis panamensis Dall, 1902 fide Coan & others (2000).
Order Mytiloida Férussac, 1822
Superfamily MYTILOIDEA Rafinesque, 1815
Family Mytilidae Rafinesque, 1815
americanum, Amygdalum Soot-Ryen, 1955: Allan Hancock Pac. Exped. 20(1):70-71, pl. 8, fig.
37.
HOLOTYPE: LACM 2111 (dry), 37 m, off North Pt., Isla de Gorgona, Cauca Dept., Colombia
(3 02 N, 78 01.5 W), Leg.: R/V Velero III, 22 JAN 1935.
azoricus, Bathymodiolus Cosel & Comtet in Cosel, Comtet & Krylova, 1999: The Veliger 42(3):
220-231. PARATYPE: LACM 3111 (wet), 866 m, Menez Gwen hydrothermal field, MidAtlantic Ridge (37°50.5 N, 31°31.3 W), Leg.: A.-M. Alayse, DIVA 2 Exped., R/V Nadir, 15 JUN
1995.
brevior, Bathymodiolus Cosel, Metivier, & Hashimoto, 1994: The Veliger 37(4):375-380.
PARATYPES: LACM 2759 (2 wet), 1842 m, Hine Hina hydrothermal field, Lau Basin, SE of
Fiji (22 32 S, 176 43 W), Leg.: BIOLAU Exped., 13 MAY 1989.
elegantulum, Dacrydium (Quendreda) Soot-Ryen, 1955: Allan Hancock Pac. Exped. 20(1):87,
pl. 8, fig. 41.
HOLOTYPE: LACM 2112 (dry); PARATYPES: LACM 2113 (32 dry), 64 m, Bahía Gardner,
Isla Espanola (Hood), Islas Galápagos (1 21 S, 89 38 W), Leg.: R/V Velero III, 31 JAN 1934.
elongatus, Bathymodiolus Cosel, Metivier, & Hashimoto, 1994: The Veliger 37(4):380-386.
PARATYPES: LACM 2760 (2 wet), 2765 m, Mussel Valley, North Fiji Basin, NW of Fiji
(18 50 S, 173 29 E), Leg.: R/V Starmer II, 13 JUL 1989.
hancocki, Lithophaga (Leiosolenus) Soot-Ryen, 1955: Allan Hancock Pac. Exped. 20(1):
102-103, pl. 10, fig. 60.
HOLOTYPE: LACM 2175 (wet); PARATYPES: LACM 1973 (3 dry), crater, Isla Onslow, N of
Isla Santa Maria (= Charles Id.), Islas Galápagos (1 14.3 S, 90 26.12 W), Leg.: R/V Velero III,
23 JAN 1938.
neglectus, Modiolus Soot-Ryen, 1955: Allan Hancock Pac. Exped. 20(1):64-65, pl. 7, figs.
31-32.
HOLOTYPE: LACM 2109 (dry); PARATYPE: LACM 2110 (1 dry), San Diego, San Diego Co.,
California.
Remarks: Currently in the subgenus Modiolusia.
phenax, Musculus Dall, 1915: The Nautilus 28(12):138.
PARATYPES: LACM 1077 (23 dry), St. George Id., Pribilof Ids., Bering Sea, Alaska, Leg.: G
D. Hanna, MAY 1939.
Remarks: Provisionally in the genus “Musculus” fide Coan & others (2000).
puteoserpentis, Bathymodiolus Cosel, Metivier, & Hashimoto, 1994: The Veliger 37(4):
387-391.
PARATYPE: LACM 2761 (1 dry valve; 1 wet valve), 3515 m, "L'Elan" (The Moose) site, Snake
Pit hydrothermal field, Mid-Atlantic Ridge (23 22 N, 47 57 W), Leg.: Hydrosnake Exped., 21
JUN 1988.
rogersi, Lithophaga (Labis) attenuate Berry, 1957: Leaflets in Malacology 1(14):76.
PARATYPES: LACM 1117 (6 dry), NW side of Caelta Cholla, near Puerto Peñasco, Sonora,
México, Leg.: M. Rogers & F. Wright, 30 DEC 1956.
rostriferum, Dacrydium (Dacrydium) Bernard, 1978: Venus 37(2):62-63, figs. 1, 12.
HOLOTYPE: LACM 1880 (dry), 2532 m, W of Cape Flattery, Clallam Co., Washington
(48 26.6 N, 126 54.5 W), Leg.: Oregon State Univ., SEP 1971.
washingtonia, Habepegris Bernard, 1978: Venus 37(2):63-64, figs. 3, 13.
HOLOTYPE: LACM 1881 (dry), 2189 m, W of Cape Flattery, Clallam Co., Washington
(48 38 N, 126 58.1 W), Leg.: Oregon State Univ., SEP 1971.
Remarks: Currently in the genus Idas fide Coan & others (2000).
Order Pterioida Newell, 1965
Suborder Pteriina Newell, 1965
Superfamily PTERIOIDEA Gray, 1847
Family Pteriidae Gray
galtsoffi, Pinctada Bartsch, 1931: Proc. USNM 79(2878):1-2.
PARATYPES: LACM 2393 (4 dry articulated specimens; 8 dry disarticulated valves), Pearl &
Hermes Atoll, NW Hawaiian Ids., Leg.: P.S. Galtsoff, Aug 1930.
Order Limoida Waller, 1978
Superfamily LIMOIDEA Rafinesque, 1815
Family Limidae Rafinesque, 1815
californica, Limatula Bernard, 1988: Venus 47(4):227-228, fig. 3.
HOLOTYPE: LACM 2335 (dry), 144 m, off Santa Catalina Id., California Channel Ids.,
California (33 17.3 N, 118 15.5 W), Leg.: R/V Velero IV, 25 OCT 1941.
Remarks: Junior synonym of Limatula saturna Bernard, 1978 fide Coan & others (2000).
coani, Limatuletta Bernard, 1988: Venus 47(4):230-231, fig. 5.
HOLOTYPE: LACM 2333 (dry), 3706-3806 m, off Patton Ridge, California (31 20 N,
129 09.9 W), Leg.: Scripps Inst. Oceanog., 23 MAR 1970.
Remarks: Currently in the genus/subgenus Limatula (Limatuletta) fide Coan & others (2000).
disalvoi, Lima Raines, 2002a: La Conchiglia 34(304):31-32, fig. 35.
HOLOTYPE: LACM 2946 (dry); PARATYPE: LACM 2947 (1 dry); PARATYPES: LACM
2948 (10 wet), 10-30 m, off Anakena, Isla de Pascua (= Easter Id.), Leg.: B.K. Raines.
macleani, Limatula Bernard, 1988: Venus 47(4):227, fig. 2.
HOLOTYPE: LACM 2336 (dry), 560 m, off San Pedro, Los Angeles Co., California (33 42 N,
118 15 W), Leg.: R/V Velero IV, 9 OCT 1965.
Remarks: Junior synonym of Limatula saturna Bernard, 1978 fide Coan & others (2000).
maui, Acesta Marshall, 2001: Mem. Museum National d'Histoire Naturelle 185:99-103.
PARATYPE: LACM 3036 (1 dry), 487-480 m, SE of Campbell Id., New Zealand (53°14.30’S,
170°12.30’E), Leg.: M. Yates, R/V Chiyo Maru, 31 AUG 1987.
nasca, Plicacesta Bernard, 1988: Venus 47(4):228-230, fig. 4.
HOLOTYPE: LACM 2332 (dry), 228 m, off Shoal Guyot, Nasca Ridge, ca. 1500 km W of
Antofagasta Prov., Chile (25 44 S, 82 25 W), Leg.: Scripps Inst. Oceanog.; Downwind Exped.,
26 JUN 1958.
Remarks: Currently in the genus Acesta.
saturna, Limatula Bernard, 1978: Venus 37(2):71-72.
PARATYPE: LACM 1879 (1 dry), 13 m, near Saturna Id., Strait of Georgia, Capital Dist.,
British Columbia, Canada (48 48.2 N, 123 13.2 W), Leg.: Fisheries Res. Board of Canada,
Scallop Survey, MAY 1960.
valdiviesae, Limaria Bernard, 1988: Venus 47(4):226, fig. 1.
HOLOTYPE: LACM 2334 (dry), 100 m, northern Chile (6 21 S, 80 56.1 W), Leg.: Instituto del
Mar, R/V Humboldt, 4 OCT 1980.
Remarks: The coordinates cited by Bernard are actually off northern Peru rather than Chile.
Suborder Pectinina Waller, 1978
Superfamily PECTINOIDEA Rafinesque, 1815
Family Pectinidae Rafinesque, 1815
cranmerorum, Somalipecten Waller, 1986: The Nautilus 100(2):42-45.
PARATYPE: LACM 2198 (1 dry), 150-300 m, off Somalia.
galapagensis, Pecten (Oppenheimopecten) Grau, 1959: Allan Hancock Pac. Exped. 23:152-154,
pl. 56.
HOLOTYPE: LACM 1749 (dry), 18-33 m, Caleta Tagus, Isla Isabela (Albemarle), Islas
Galápagos (0 16.13 N, 91 22.63 W), Leg.: R/V Velero III, 15 JAN 1934.
hancocki, Pecten (Oppenheimopecten) Grau, 1959: Allan Hancock Pac. Exped. 23:154-155, pl.
57.
HOLOTYPE: LACM 1750 (dry), 86 m, Bahía Chatham, Isla del Coco, East Pacific (5 33.8 N,
86 58.7 W), Leg.: R/V Velero III, 14 JAN 1938; PARATYPES: LACM 2114 (2 dry), 57-91 m,
off Isla Nuez, Isla del Coco, East Pacific (5 34 N, 86 59 W), Leg.: R/V Velero III, 13 JAN 1938.
phalara, Chlamys Roth, 1975b: Jour. Malacological Soc. Aust. 3(2):81-84.
PARATYPES: LACM 2653 (4 dry), 80-200 m, off Isla Más a Tierra, Islas Juan Fernández, East
Pacific (33 36 S, 75 55 W), Leg.: R/V Anton Bruun, 15 DEC 1975.
Family Propeamussiidae Abbott, 1954
acutus, Cyclopecten Grau, 1959: Allan Hancock Pac. Exped. 23:31-32, pl. 10, fig. 2.
HOLOTYPE: LACM 1746 (dry), 59 m, N side of Isla de Gorgona, Cauca Dept., Colombia
(3 01 N, 78 11 W), Leg.: R/V Velero III, 24 FEB 1938
argenteus, Cyclopecten Bernard, 1978: Venus 37(2):66-68, figs. 4, 14.
HOLOTYPE: LACM 1883 (dry), 1530 m, NW of Cape Blanco, Curry Co., Oregon (44 25.6 N,
125 13.4 W), Leg.: Oregon State Univ., AUG 1964.
barbarensis, Cyclopecten Grau, 1959: Allan Hancock Pac. Exped. 23:37-38, pl. 14, fig. 1.
HOLOTYPE: LACM 1748 (dry), 49-55 m, San Nicolas Id., California Channel Ids., California
(33 18.3 N, 119 28.7 W), Leg.: R/V Velero III, 11 APR 1940.
Remarks: Grau cites that the type locality is “off Santa Barbara Id., southern Califonria, in 27
fm., Hancock expeditions bottom sample 1064, 33º30'01"N, 119º02'20"W, May 29, 1939” but
his original label notes that the locality is actually AHF locality 1086 (see data above).
benthalis, Cyclopecten Grau, 1959: Allan Hancock Pac. Exped. 23:24, pl. 5.
HOLOTYPE: LACM 1744 (dry), 896 m, 14.5 mi. NNW of, San Nicolas Id., California Channel
Ids., California (33 28.07 N, 119 34.68 W), Leg.: R/V Velero IV, 07 MAY 1955.
catalinensis, Pecten (Cyclopecten) Willett, 1931b: The Nautilus 45(2):65, pl. 4, figs. 1-2.
HOLOTYPE: LACM 1020 (dry); PARATYPE: LACM 2138 (1 dry), 183 m, off Whites Landing,
N side of Santa Catalina Id., California Channel Ids., California, Leg.: George Willett, JUL 1928.
Remarks: Currently in the genus Cyclopecten fide Coan & others (2000). Paratype ex UCLA
20111.
exquisitus, Cyclopecten Grau, 1959: Allan Hancock Pac. Exped. 23:34-36, pl. 12.
HOLOTYPE: LACM 1747 (dry), 183-274 m, off Isla Wenman (= Wolf Id.), Islas Galápagos
(1 23.1 N, 91 48.7 W), Leg.: R/V Velero III, 11 JAN 1934.
knudseni, Cyclopecten Bernard, 1978: Venus 37(2):68-69, figs. 5, 15.
HOLOTYPE: LACM 1882 (dry), 2605 m, NW of Yaquina Head, Lincoln Co., Oregon
(45 17.4 N, 125 49 W), Leg.: Oregon State Univ., MAR 1970.
squamiformis, Cyclopecten Bernard, 1978: Venus 37(2):69-70, figs. 6, 16.
HOLOTYPE: LACM 1884 (dry), 2650 m, SW of Astoria, Clatsop Co., Oregon (45 52.7 N,
126 29.4 W), Leg.: Oregon State Univ., FEB 1971.
zephyrus, Cyclopecten Grau, 1959: Allan Hancock Pac. Exped. 23:25-27, pl. 7, fig. 1.
HOLOTYPE: LACM 1745 (dry), 732-786 m, 5.5 mi. off Long Pt., Santa Catalina Id., California
Channel Ids., California (33 27.92 N, 118 20.28 W), Leg.: R/V Velero IV, 2 OCT 1948.
Family Plicatulidae Yonge, 1977
anomioides, Plicatula Keen, 1958: Bulls. Amer. Paleo. 38(172):241-242.
PARATYPE: LACM 2267 (1 dry), on breakwater in front of Miramar Hotel, Guaymas, Sonora,
México.
Remarks: Ex UCLA 46026.
Subclass Palaeoheterodonta Newell, 1965
Order Unionoida Stoliczka, 1871
Superfamily UNIONOIDEA Fleming, 1828
Family Unionidae Fleming, 1828
brooksiana, Anodonta van der Schalie, 1938: Ann. Carnegie Mus. 27(12):167-170.
PARATYPES: LACM 2715 (6 dry), Spout Pond, Ferryland Dist., Newfoundland, Canada, Leg.:
Dr. & Mrs. S.T. Brooks, 21 SEP 1937.
conchos, Disconaias Taylor, 1997: Occas. Pap. Moll. 5(75):420-423, pl. 21, fig. 1.
HOLOTYPE: LACM 2257 (dry); PARATYPES: LACM 2849 (2 dry), Rio Conchos, about 0.5
km west of Julimes, Chihuahua State, México, Leg.: D.W. Taylor & A.L. Metcalf, 29 AUG
1969.
lananensis, Quadrula Frierson, 1901: The Nautilus 15(7):75-76.
SYNTYPE: LACM 2018 (1 dry), Lanana Creek, near Nacogdoches, Nacogdoches Co., Texas,
Leg.: Askew, Strode, & Frierson, 10 JUL 1901.
Remarks: Currently in the genus Fusconaia.
moreleti, Unio Deshayes, 1848: Explor. sci. Algerie, Zool. 1: Mollusca, pl. 109,
figs. 1-4, pl. 112, fig. 5.
SYNTYPE: LACM 2232 (1 dry), Ech Cheliff (= Orleansville), El Asnan Dist., Algeria.
Remarks: Junior synonym of Unio elongatulus durieui Deshayes, 1847. Ex UCLA 32172.
ortmanni, Unio Frierson, 1913: The Nautilus 27(2):14-15.
PARALECTOTYPES: LACM 2017 (2 dry), designated by Johnson (1972:146), Occas.
Papers on Moll. 3(41), Rio Cochins, near Quirigua, Izabal Dept., Guatemala.
Remarks: Junior synonym of Elliptio sphenorhynchus (Crosse & Fischer, 1893).
patsaligensis, Pleurobema Simpson, 1900: Proc. Acad. Nat. Sci. Philadelphia 52:82.
PARATYPES: LACM 2074 (2 dry), Little Patsaliga Creek, Crenshaw Co., Alabama.
Remarks: Junior synonym of Pleurobema strodeanum (Wright, 1898).
sajensis, Elliptio Frierson, 1927: Check list of No. Amer. Naiades, Baylor Univ. Press, p. 37.
PARATYPES: LACM 2019 (3 dry), Rio Saja, 4 mi. W of Esmeralda, Jutiapa Dept., Guatemala.
Subclass Heterodonta Neumayr, 1884
Order VENEROIDA H. Adams & A. Adams, 1856
Family Thyasiridae Dall, 1900
careyi, Axinulus Bernard, 1979b: Nat. Hist. Mus. L.A. Co. Cont. Sci. 313:33-34, fig. 50.
HOLOTYPE: LACM 1900 (dry); PARATYPE: LACM 1901 (1 dry), 695 m, Beaufort Sea,
Barrow Co., Alaska (71 14.3 N, 149 22.9 W), Leg.: Oregon State Univ., 5 SEP 1971.
orecta, Thyasira Bernard, 1982: Venus 41(3):177-180, figs. 1-3c.
HOLOTYPE: LACM 1804 (dry), 2520 m, W of Cape Flattery, Clallam Co., Washington
(48 18.4 N, 127 04.2 W), Leg.: Oregon State Univ.
phrygiana, Thyasira Miloslavskaya, 1977: Studies of sea fauna 14(22):402.
PARATYPES: LACM 2647 (2 dry), 53 m, Ostrov Vrangelya (= Wrangel Id.), East Siberian Sea,
Chukchi Auton. Okrug, Russia (73 02 N, 178 43 E), Leg.: R/V Krasin, 7 SEP 1935.
redondoensis, Aligena T. Burch, 1941: The Nautilus 55(2):50-51, pl. 4, figs. 5-6.
HOLOTYPE: LACM 1844 (dry); PARATYPES: LACM 1010 (4 dry); PARATYPES: LACM
1845 (4 dry), 137 m, off Redondo Beach, Los Angeles Co., California, Leg.: T.A. & J.Q. Burch,
AUG 1938.
Remarks: Holotype ex AHF 382. Currently in the genus Axinodon fide Coan & others (2000).
sphaericosa, Adontorhina Scott, 1986: The Veliger 29(2):150-153.
PARATYPES: LACM 2087 (3 wet), 280 m, Boca de Quadra Fjord, Ketchikan Co., Alaska
(55 10.2 N, 130 37.5 W), Leg.: R. Cimberg, SEP 1981.
Superfamily ASTARTOIDEA d’Orbigny, 1844
Family Astartidae d’Orbigny, 1844
derjugini, Astarte (Astarte) Filatova, 1957: Trudy Inst. Okean. Akad. Nauk SSSR, p. 298.
PARATYPES: LACM 2643 (2 dry), 253 m, Sea of Okhotsk, SW of Kamchatka Region, Russia,
Leg.: R/V Vityaz, JUN 1951.
Remarks: Junior synonym of Astarte crenata (Gray, 1824) fide Coan & others (2000).
derbecki, Elliptica alaskensis Scarlato, 1981: Guide to the fauna of the USSR, p. 294.
PARATYPE: LACM 2649 (1 dry), 84 m, SE of Mys Levenshterna, Sea of Okhotsk, Sakhalin Id.,
Russia (53 47 N, 143 22 E), Leg.: P. Ushakov; R/V Gagara, 11 AUG 1932; PARATYPES:
LACM 2650 (3 dry), 20 m, Yakshina Bight, Ostrova Bol'shoy Shantar; Shantarskiye Ostrova,
Sea of Okhotsk, Khabarovsk Kray, Russia (54 44 N, 137 36 E), Leg.: I. Zaks, 10 SEP 1927.
Remarks: Junior synonym of Astarte elliptica (Brown, 1827) fide Coan & others (2000).
inaequilatera, Nicania Filatova, in Scarlato, 1981: Guide to the fauna of the USSR, p. 302.
PARATYPES: LACM 2631 (11 dry), 60-70 m, Shikotan-To, Kurile Ids., Russia. Leg.: E.
Gujanova, 19 SEP 1949.
Remarks: Currently in the genus Astarte fide Coan & others (2000).
mirabilis, Rictocyma Dall, 1871: Amer. Jour. Conchology 7:151.
PARATYPES: LACM 2396 (5 dry valves), 15 m, Unga Id., Shumagin Ids., Aleutian Ids. Co.,
Alaska, Leg.: W.H. Dall.
Remarks: Junior synonym of Astarte esquimalti (Baird, 1863) fide Coan & others (2000).
vernicosa, Astarte (Nicania) Dall, 1903a: Proc. USNM 26(1342):945, 948-949.
PARATYPES: LACM 1204 (6 dry), 18-29 m, Nazan Bay, Atka Id., Aleutian Ids. Co., Alaska.
willetti, Astarte Dall, 1917b: The Nautilus 31(1):11-12.
PARATYPES: LACM 2170 (7 dry), 91 m, Forrester Id., Prince of Wales Co., Alaska, Leg.: G.
Willett.
Remarks: Junior synonym of Astarte compacta Carpenter, 1864 fide Coan & others (2000). Ex
UCLA 6735.
Superfamily CRASSATELLOIDEA Férussac, 1822
Family Crassatellidae Férussac, 1822
skoglundae, Crassinella Coan, 1979: The Veliger 22(1):6-7.
PARATYPE: LACM 1904 (1 dry), 7.5-15 m, La Cruz, Bahía Banderas, Nayarit, México
(20 45 N, 105 24 W), Leg.: C. & P. Skoglund, 31 JUL 1973.
Superfamily CARDITOIDEA Fleming, 1828
Family Carditidae Fleming, 1828
montereyensis, Cardita ventricosa Smith & Gordon, 1948: Proc. Calif. Acad. Sci., 4th ser.,
26(8):212-216.
PARATYPES: LACM 1002 (4 dry), 115 m, 4.6 mi. NW of Pt. Pinos, Monterey Bay, Monterey
Co., California, Leg.: G.C. McGinitie, 9 MAR 1931.
Remarks: Junior synonym of Cyclocardia ventricosa (Gould, 1850) fide Coan & others (2000).
redondoensis, Cardita ventricosa T. Burch, 1944b: Min. Conch. Club So. Calif. 39:14.
PARATYPES: LACM 1299 (3 dry); PARATYPE: LACM 1769 (1 dry), 137-183 m, off
Redondo Beach, Los Angeles Co., California, Leg.: T.A. & J.Q. Burch, AUG 1941.
Remarks: Junior synonym of Cyclocardia ventricosa (Gould, 1850) fide Coan & others (2000).
umnaka, Cardita Willett, 1932c: Trans. San Diego Soc. Nat. Hist. 7(9):87-88, pl. 5, figs. 1a-2b.
HOLOTYPE: LACM 1034 (dry); PARATYPES: LACM 2140 (120 dry valves); PARATYPES:
LACM 2278 (2 dry), Umnak Id., Aleutian Ids. Co., Alaska, Leg.: G. Willett, JUL 1926.
Remarks: Currently in the genus Cyclocardia fide Coan & others (2000). Paratype lot LACM
2140 ex UCLA 20132; paratype lot LACM 2278 ex UCLA 1406.
Family Condylocardiidae Bernard, 1897
elongata, Condylocardia Coan, 2003: The Nautilus 117(2):54, figs. 21-23.
HOLOTYPE: LACM 2926 (dry); PARATYPES: LACM 2927 (5 dry), 15-30 m, Punta Espinosa,
Isla Fernandina (= Narborough Id.) (0°16.5'S, 91°26.20'W), Leg.: R/V Searcher, 25 JAN 1972.
fernandina, Condylocardia Coan, 2003: The Nautilus 117(2):53-54, figs. 15-17.
HOLOTYPE: LACM 2920 (dry); PARATYPES: LACM 2921 (7 dry), 12 m, Punta Espejo, E
side of Isla Marchena (= Bindloe Id.), Islas Galápagos (0°19.30'N, 90°24'W), Leg.: J.H. McLean,
11 MAY 1984.
galapagana, Carditella Coan, 2003: The Nautilus 117(2):57-, figs. 29-32.
HOLOTYPE: LACM 2930 (dry); PARATYPES: LACM 2931 (8 dry), 183-274 m, Isla Wenman
(= Wolf Id.), Islas Galápagos (1°23'N, 91°49'W), Leg.: R/V Velero III, 11 JAN 1934.
geigeri, Condylocardia Coan, 2003: The Nautilus 117(2):56-57, fig. 28.
HOLOTYPE: LACM 2962 (dry); PARATYPES: LACM 2963 (135 dry), 37 m, Bahía Sullivan,
Isla San Salvador (= James Id.), Islas Galápagos (0°16'S, 90°34'W), Leg.: Velero III, 24 JAN
1934; PARATYPE: LACM 2964 (1 dry), 16 m, Bahía Academy, Isla Snta Cruz (= Indefatigable
Id.), Islas Galápagos (0°45'S, 90°18'W), Leg.: R/V Velero III, 7-8 DEC 1934.
kaiseri, Condylocardia Coan, 2003: The Nautilus 117(2):54-56, figs, 24-27.
HOLOTYPE: LACM 2928 (dry); PARATYPES: LACM 2929 (21 dry), 12 m, Punta Espejo, E
side of Isla Marchena (= Bindloe Id.), Islas Galápagos (0°19.30'N, 90°24'W), Leg.: J.H. McLean,
11 MAY 1984.
koolsae, Condylocardia Coan, 2003: The Nautilus 117(2):53-54, figs. 18-20.
HOLOTYPE: LACM 2924 (dry); PARATYPES: LACM 2925 (5 dry), 274 m, Isla Wenman (=
Wolf Id.), Islas Galápagos (1°23'N, 91°49'W), Leg.: R/V Velero III, 11 JAN 1934.
pusilla, Benthocardiella Powell, 1930: Trans. Proc. New Zealand Inst. 60:533-534.
PARATYPE: LACM 1361 (1 dry), 18-22 m, off Wanganui, Wellington, North Id., New Zealand.
sparsa, Condylocardia Coan, 2003: The Nautilus 117(2):50-53, figs. 12-14.
HOLOTYPE: LACM 2918 (dry); PARATYPES: LACM 2919 (10 dry), 8-12 m, Punta Penca, N
of Bahía Potrero, Guanacaste Prov., Costa Rica (10º29.15'N, 85º48.85'W), Leg.: P.I. LaFollette &
D. Cadien, 20 FEB 1972.
Superfamily CHAMOIDEA Lamarck, 1809
Family Chamidae Lamarck, 1809
arcana, Chama Bernard, 1976: Nat. Hist. Mus. L.A. Co. Cont. Sci. 278:14-15, fig. 4a.
HOLOTYPE: LACM 1723 (dry), 0 m, intertidal, Newport Bay, Orange Co., California (33 37 N,
117 55 W), Leg.: J.H. McLean, MAR 1965.
cerion, Chama Matsukuma, Paulay, & Hamada, 2003: Venus 62(1-2):21-25.
PARATYPES: LACM 3065 (3 dry), Moorea, Society Arch., French Polynesia, Leg.: E.
Lovatelli.
clarionensis, Pseudochama Willett, 1938a: The Nautilus 52(2):48-49, pl. 4, figs. 1-2.
HOLOTYPE & PARATYPE (attached): LACM 1058 (3 dry), 55 m, off SW side of Isla Clarión,
Islas Revillagigedo, México, Leg.: G. Willett, 24 MAR 1938; PARATYPES: LACM 1627 (26
dry), 37-64 m, Isla Clarión, Islas Revillagigedo, México (18 22 N, 114 44 W), Leg.: G. Willett,
24 MAR 1938.
garthi, Chama Bernard, 1976: Nat. Hist. Mus. L.A. Co. Cont. Sci. 278:18, fig. 9.
HOLOTYPE: LACM 2006 (dry); PARATYPE: LACM 2007 (1 dry), Bahía Octavia, Choco
Dept., Colombia (6 50 N, 77 41.1 W), Leg.: R/V Velero III, 28 JAN 1935.
Remarks: Holotype ex AHF 167, paratype ex AHF 168.
granti, Pseudochama Strong, 1934: The Nautilus 47(4):137.
PARATYPES: LACM 2162 (4 dry), 27-55 m, off the S side of Santa Catalina Id., California
Channel Ids., California.
Remarks: Ex UCLA 3398.
tinctoria, Chama Bernard, 1976: Nat. Hist. Mus. L.A. Co. Cont. Sci. 278:22, fig. 5.
HOLOTYPE: LACM 2008 (dry); PARATYPE: LACM 2009 (1 dry), 27-37 m, Bahía Honda, Isla
Coiba, Veraguas Prov., Panamá (7 43.27 N, 81 32.92 W), Leg.: R/V Velero III, 22 FEB 1934.
Remarks: Holotype ex AHF 169, paratype ex AHF 170.
Superfamily GALEOMMATOIDEA Gray, 1840
Family Galeommatidae Gray, 1840
parmaleeana, Bellascintilla Coney, 1990: The Nautilus 104(4):138-141, figs. 21-30.
HOLOTYPE: LACM 2446 (dry), 37 m, off Bahía Herradura, Puntarenas Prov., Costa Rica
(9 38.8 N, 84 40.8 W), Leg.: R/V Searcher, 10 MAR 1972; PARATYPES: LACM 2447 (5 dry),
12 m, Isla del Cano, Puntarenas Prov., Costa Rica (8 44.0 N, 83 52.5 W), Leg.: J.H. Mclean, W.
Wheeler; R/V Searcher, 14-19 MAR 1972; PARATYPE: LACM 2448 (1 dry), 18-37 m,
between Rancho Palmilla & Rancho El Tule, Golfo de California, Baja California Sur, México
(22 58 N, 109 48 W), Leg.: J.H. McLean et al., 5 APR 1966; PARATYPE: LACM 2449 (1 dry),
Playa Nancite, N side Golfo de Papaguayo, Parque Nacional Santa Rosa, Guanacaste Prov.,
Costa Rica (10 48 N, 85 42 W), Leg.: E.V. Coan, R. Hollywood, 15 JAN 1986; PARATYPES:
LACM 2450 (7 dry), Punta Chame, Golfo de Panamá, Panamá Prov., Panamá (8 41 N,
79 39 W), Leg.: D.R. Shasky, JUN 1977; PARATYPE: LACM 2451 (1 dry), 6-12 m, N side Isla
Salango, Manabi Prov., Ecuador (1 35.5 S, 80 53.4 W), Leg.: D.R. Shasky, 26 JUN-4 JUL 1980.
Family Lasaeidae Gray, 1842
cistula, Lasaea Keen, 1938: Proc. Malac. Soc. London 23(1):25-26.
PARATYPES: LACM 2172 (4 dry), Half Moon Bay, San Mateo Co., California.
Remarks: Junior synonym of Lasaea. adansoni (Gmelin, 1791) fide Coan & others (2000). Ex
UCLA 20344.
nipponica, Lasaea Keen, 1938: Proc. Malac. Soc. London 23(1):26-28.
PARATYPE: LACM 1079 (1 dry), Watanoha, Matsusima, Miyagi Pref., Honshu, Japan.
vadosa, Boreacola Bernard, 1979b: Nat. Hist. Mus. L.A. Co. Cont. Sci. 313:37-38, fig. 56.
HOLOTYPE: LACM 1902 (dry); PARATYPE: LACM 1903 (1 dry, missing), 0.5 m, intertidal,
Pitt Pt., Beaufort Sea, Barrow Co., Alaska (70 55.73 N, 153 12.73 W), Leg.: Western
Washington St. Coll.
Remarks: Junior synonym of Boreacola maltzani (Verkrüzen, 1875) fide Coan & others (2000).
Superfamily GLOSSOIDEA Gray, 1847
Family Vesicomyidae Dall & Simpson, 1901
magnifica, Calyptogena (Ectenagena) Boss & Turner, 1980: Malacologia 20(1):165-183.
PARATYPES: LACM 1951 (2 dry), 2451 m, Galapagos Rift, East Pacific (0 48.0 N,
86 13.00 W), Leg.: R/V Alvin, 1 DEC 1979.
Superfamily CYAMIOIDEA Sars, 1878
Family Cymiidae Sars, 1878
aucklandica, Legrandina Powell, 1933: Proc. Malac. Soc. London 20(5):233.
PARATYPE: LACM 1356 (1 dry), Faith Harbour, Auckland Ids., New Zealand.
Family Sportellidae Dall, 1899
obliqua, Basterotia Coan, 1999: Veliger 42(2):137-138, figs. 6-7, 32-33.
HOLOTYPE: LACM 2846 (1 dry valve); PARATYPES: LACM 2847-2848 (single dry valves),
100 m, 3 mi. south of Peak, Tetas de Cabra, Bahía San Carlos, near Guyamas, Golfo de
California, Sonora, Mexico (27 53 N, 111 06 W), Leg.: Roy & Forrest Poorman, APR 1978.
Superfamily CORBICULOIDEA Gray, 1847
Family Pisidiidae Gray, 1857
sanguinichristi, Pisidium (Cyclocalyx) Taylor, 1987: New Mexico Bur. Mines Min. Res. Bull.
116:47-48, fig. 24.
HOLOTYPE: LACM 2258 (dry), 3305 m, Middle Fork Lake (a cirque lake), Sangre de Cristo
Mountains, Taos Co., New Mexico, 21 MAY 1981
Superfamily VENEROIDEA Rafinesque, 1815
Family Veneridae Rafinesque, 1815
catharia, Callocardia Dall, 1902: Proc. USNM 26(1312):402.
PARATYPES: LACM 1205 (48 dry), 37 m, off Guaymas, Sonora, México, Leg.: R/V Albatros.
confusa, Transennella S. Gray, 1982: Malac. Review 15:109-112.
PARATYPES: LACM 1950 (21 dry), Lawson's flat, Tomales Bay, Marin Co., California
(38 13 N, 122 57 W), Leg.: S. Gray, 10 AUG 1979.
Remarks: Currently in the genus Nutricola fide Coan & others (2000)
Superfamily TELLINOIDEA Blainville, 1814
Family Tellinidae Blainville, 1814
dexioptera, Macoma Baxter, 1977: The Veliger 19(3):272-276.
PARATYPES: LACM 1786 (2 dry), MacLeod Harbor, Montague Id., Prince William Sound,
Valdez-Chitina-Whittier Co., Alaska (59 53.4 N, 147 49.8 W), Leg.: R. Baxter, JUL 1965
Remarks: Currently in the subgenus Rexithaerus fide Coan & others (2000).
inflatula, Macoma Dall, 1897: Nat. Hist. Soc. Br. Columbia, Bull. 2(1):11.
PARATYPES: LACM 1328 (12 dry), 22 m, Captain's Bay, Unalaska Id., Aleutian Ids. Co.,
Alaska.
Remarks: Junior synonym of Macoma (Psammacoma) carlottensis Whiteaves, 1880 fide Coan
& others (2000).
morroensis, Macoma (Psammacoma) T. Burch, 1945: Min. Conch. Club So. Calif. 47:33.
PARATYPE: LACM 2040 (1 dry); PARATYPES: LACM 2153 (2 dry), near entrance of Morro
Bay, San Luis Obispo Co., California, Leg.: M.E. Caruthers.
Remarks: Junior synonym of Macoma (Psammacoma) acolasta Dall, 1921 fide Coan & others
(2000). Paratype lot LACM 2153 ex UCLA 20516.
pacis, Macoma Pilsbry & Lowe, 1932a: Proc. Acad. Nat. Sci. Philadelphia 84:95.
PARATYPES: LACM 2182 (1 dry), La Paz, Golfo de California, Baja California Sur, México,
Leg.: H.N. Lowe.
Remarks: Junior synonym of Macoma viridotincta (Carpenter, 1856) fide Keen (1971). Ex
UCLA 3896.
Family Donacidae Fleming, 1828
dorotheae, Donax Morrison, 1971: Proc. Biol. Soc. Washington 83(48):554-555.
PARATYPES: LACM 1604 (approx. 2800), 0 m, intertidal, Grand Isle, Jefferson Parish,
Louisiana, Leg.: J.P.E. Morrison, 3 JAN 1970.
Family Psammobiidae Fleming, 1828
kussakini, Asaphis Ivanova, 1985: Vladivostok Far East Acad. Akad. Sci., p. 83.
PARATYPES: LACM 2646 (2 dry), 5 m, Ostrov Moneron, Sea of Japan, W of Sakhalin Id.,
Russia (46 15 N, 141 16 E), Leg.: V. Fadeev, 15 JUL 1977.
petri, Soletellina (Nuttallia) Bartsch, 1929: Inst. Hydro. Explor. mers d'URSS 10:133.
PARATYPE: LACM 2648 (1 dry valve), Zaliv Petra Velikogo (= Peter the Great Bay), Sea of
Japan, Primorye Kray, Russia., Leg.: K. Derjugin.
Remarks: Currently in the genus Nuttallia.
purpureus, Heterodonax bimaculatus Williamson, 1893: Proc. USNM 15(898):187.
LECTOTYPE: LACM 1168 (dry), designated by Coan (1973:46), The Veliger 16(1);
PARALECTOTYPES: LACM 1176 (9 dry) (see above), San Pedro, Los Angeles Co., California.
Remarks: Junior synonym of Heterodonax pacificus (Conrad, 1837) fide Coan & others (2000).
salmoneus, Heterodonax bimaculatus Williamson, 1893: Proc. USNM 15(898):187.
LECTOTYPE: LACM 1169 (dry), designated by Coan (1973:47), The Veliger 16(1);
PARALECTOTYPES: LACM 1177 (12 dry) (see above), San Pedro, Los Angeles Co.,
California.
Remarks: Junior synonym of Heterodonax pacificus (Conrad, 1837) fide Coan & others (2000).
Family Semelidae Stoliczka, 1870
guaymasensis, Semele (Leptomyaria) Pilsbry & Lowe, 1932a: Proc. Acad. Nat. Sci.
Philadelphia 84:92.
PARATYPES: LACM 2185 (4 dry), 37 m, Guaymas, Sonora, México, Leg.: H.N. Lowe, JAN
1930.
Remarks: ExUCLA 3914
Superfamily SOLENOIDEA Lamarck, 1809
Family Solenidae Lamarck, 1809
tairona, Solen (Solen) Cosel, 1985: Bull. Mus. Nat. Hist. Nat., Paris, 4th ser., sect. A,
7(2):328-333, fig. 2a; pl. 2, fig. 2.
PARATYPE: LACM 2210 (1 dry), Cartagena-Crespo, Bolivar Dept., Colombia.
Family Pharidae H. Adams & A. Adams, 1858
myrae, Ensis Berry, 1953b: Trans. San Diego Soc. Nat. Hist. 11(15):398-401.
PARATYPE: LACM 1104 (1 dry), Terminal Id., San Pedro, Los Angeles Co., California, Leg.:
W.H. Eschaur.
Order Myoida Goldfuss, 1820
Suborder Myina Goldfuss, 1820
Superfamily MYOIDEA Lamarck, 1809
Family Corbulidae Lamarck, 1818
grovesi, Corbula (Varicorbula) Coan, 2002: Malacologia 44(1):74-75, figs. 23, 49.
HOLOTYPE: LACM 2891 (dry); PARATYPE: LACM 2892 (2 dry valves), 731.7 m, 8 mi. 283
deg. T from S end of Isla San Lorenzo, Golfo de California, Baja California Sur, México
(28°38.75'N, 113°0'W), Leg.: R/V Velero IV, 2 DEC 1967.
rosea, Corbula luteola Williamson, 1905b: Bull. So. Calif. Acad. Sci. 4(8):120.
SYNTYPE: LACM 1421 (1 dry), Terminal Id., Los Angeles Co., California.
Remarks: Junior synonym of Juliacorbula luteola (Carpenter, 1864) fide Coan & others (2000).
Family Spheniopsidae Gardner, 1928
frankbernardi, Spheniopsis Coan, 1990: Veliger 33(4):396-397, figs. 4-6.
HOLOTYPE: LACM 2427 (dry); PARATYPE: LACM 2428 (1 dry); PARATYPE: LACM 2429
(1 dry), 18-37 m, off Cabo San Lucas, Golfo de California, Baja California Sur, México
(22 08 N, 110 W), Leg.: P.M. Oringer & L. Marincovich, 4 APR 1966.
Order Pholadina H. Adams & A. Adams, 1858
Superfamily PHOLADOIDEA Lamarck, 1809
Family Pholadidae Lamarck, 1809
fitchi, Penitella Turner, 1955: Johnsonia 3(34):71-75.
PARATYPES: LACM 1444 (2 wet), N side of Bahia San Bartolome, Pacific Coast Baja
California Sur, México, Leg.: J. Fitch, E. Swyer & D. Joseph.
hopkinsi, Penitella Kennedy & Armentrout, 1989: The Veliger 32(3):320-324, figs. 5, 7, 9, 11.
HOLOTYPE: LACM 2387 (dry); PARATYPES: LACM 2388 (12 dry), SW side of Wingham
Id., NW of Kayak Id., Cordova-Mc Carthy Co., Alaska (59 59.47 N, 144 24.6 W), Leg.: John M.
Armentrout, 16 JUL 1975; PARATYPES: LACM 2389 (2 dry), 0 m, intertidal, Nuchek, N side
Pt. Etches, Hinchinbrook Id., Prince William Sound, Cordova-McCarthy Co., Alaska (60 20 N,
146 40 W), Leg.: R. Baxter, JUN 1965.
richardsoni, Penitella Kennedy, 1989: The Veliger 32(3):316-318, figs. 7, 13.
HOLOTYPE: LACM 2378 (wet), 0 m, intertidal, cove at mouth of Coon Creek, 0.6 km NE Pt.
Buchon, S of Morro Bay, San Luis Obispo Co., California (35 15.60 N, 120 53.80 W), Leg.: C.
C. Coney, 19 MAY 1984; PARATYPE: LACM 2379 (1 wet), 0 m, intertidal, N side of Sunset
Bay, Coos Co., Oregon (43 20.15 N, 124 22.03 W), 29 JUN 1942; PARATYPES: LACM 2380
(5 dry), 18 m, off Del Monte, Monterey Bay, Monterey Co., California (36 37 N, 121 52.5 W),
Leg.: J.H. McLean; R/V Tage, 1960-1962; PARATYPES: LACM 2381 (2 dry), 0 m, intertidal,
Pt. Fermin, Palos Verdes Peninsula, Los Angeles Co., California (33 43 N, 118 17 W), Leg.: M.
Rogers, 12 JUN 1960; PARATYPE: LACM 2382 (1 dry), 0 m, intertidal, Carpinteria Reef, Santa
Barbara Co., California (34 23 N, 119 31 W), Leg.: M. Rogers, 12 FEB 1961; PARATYPES:
LACM 2383 (2 dry), 0 m, intertidal, Camalu-by-the-Sea, Pacific Coast Baja California, México
(30 50 N, 116 05 W), Leg.: J.H. McLean & P.M. Oringer, 5-6 JAN 1966; PARATYPES: LACM
2384 (2 wet), 0 m, intertidal, Bolinas Bay, beach near Duxbury Reef, Marin Co., California
(37 55 N, 122 40 W), Leg.: E.C. Wilson & W. Carney, 20 SEP 1968; PARATYPE: LACM
2385 (1 dry), 0 m, intertidal, Middle Cove, Cape Arago State Park, Coos Co., Oregon (43 18 N,
124 24 W), Leg.: G. Sphon & G.L. Kennedy, 10 AUG 1972; PARATYPES: LACM 2386 (3 dry,
2 wet), 0 m, intertidal, 0.4 km NW of South Pt., Shell Beach, San Luis Obispo Co., California
(35 09.3 N, 120 40.5 W), Leg.: P.I. LaFollette, 10 DEC 1977; PARATYPES: LACM 2390 (5
dry), Tacoma, Pierce Co., Washington
Remarks: Paratype lot LACM 2390 ex UCLA 20643.
Subclass Anomalodesmata Dall, 1889
Order Pholadomyoida Newell, 1965
Superfamily PANDOROIDEA Rafinesque, 1815
Family Pandoridae Rafinesque, 1815
forresterensis, Pandora (Kennerlyia) Willett, 1918: The Nautilus 31(4):134-135.
PARATYPES: LACM 2327 (12 dry), 91 m, Forrester Id., Prince of Wales Co., Alaska, Leg.:
G. Willett, 1916-1917.
Remarks: Junior synonym of Pandora (Pandorella) wardiana A. Adams, 1859 fide Coan &
others (2000). Ex UCLA 20232.
Family Lyonsiidae Fischer, 1887
cucumerina, Lyonsia Ivanova, in Scarlato, 1981: Guide to the fauna of the USSR, p. 280-281.
PARATYPE: LACM 2642 (1 dry), Intertidal, Yakshina Bight, Ostrov Bol'shoy Shantar,
Shantarskiye Ostrova, Sea of Okhotsk, Khabarovsk Kray, Russia (54 44 N, 137 36 E), Leg.: M.
Ivanova, AUG 1966.
delicata, Lyonsia Marincovich, 1973: Nat. Hist. Mus. L.A. Co. Sci. Bull. 16:15, figs. 23-24.
HOLOTYPE: LACM 1583 (dry); PARATYPES: LACM 1584 (29 dry), 0 m, intertidal, rocky
beach at N end of Playa Cavancha, Iquique, Tarapaca Prov., Chile, (20 13 S, 70 10 W), Leg.: L.
Marincovich, AUG 1964.
nuculaniformis, Lyonsia Scarlato, 1981: Guide to the fauna of the USSR, p. 279-280.
PARATYPE: LACM 2640 (1 dry), SE of Mys Kastri, Zaliv Chikhacheva, Tatarskiy Provliv, Sea
of Japan, Sakhalin Id., Russia (51 29 N, 140 50 E), Leg.: Tarasov, 1929.
vniroi, Lyonsia Scarlato, 1981: Guide to the fauna of the USSR, p. 280.
PARATYPE: LACM 2641 (1 dry), Yakshina Bight, Ostrov Bol'shoy Shantar, Shantarskiye
Ostrova, Sea of Okhotsk, Khabarovsk Kray, Russia (54 44 N, 137 36 E), Leg.: I. Zaks, 5 AUG
1927.
Family Periplomatidae Dall, 1895
rosewateri, Periploma (Septentrioploma) Bernard, 1989a: Venus 48(1):6, fig. 11.
HOLOTYPE: LACM 2331 (dry), 600 m, off Lincoln Co., Oregon (44 38.8 N, 124 54.9 W),
Leg.: Oregon State Univ.; R/V Yaquina, 19 FEB 1970.
Superfamily PHOLADOMYOIDEA Gray, 1847
Family Parilimyidae Morton, 1847
chilensis, Panacca Coan, 200: Malacologia 42(1-2):165-168, figs. 1-2.
HOLOTYPE: LACM 2876 (1 dry valve): 130-180 m, off NW side of Isla Mas a Tierra, Islas
Juan Fernandez (33°34'S, 78°45'W), R/V Anton Bruun, 13-15 DEC 1965.
Order Septibranchida Pelseneer, 1888
Superfamily CUSPIDARIOIDEA Dall, 1886
Family Cuspidariidae Dall, 1886
ascoldica, Cuspidaria Scarlato, 1972: Trudy Zool. Inst. Akad. Nauk SSSR 52:121.
PARATYPE: LACM 2637 (1 dry), 200 m, off Mya Povorotny, Sea of Japan, Primorye Kray,
Russia (42 40 N, 133 02 E), Leg.: A. Tarasov, 21 AUG 1930.
buccina, Cuspidaria (Cuspidaria) Bernard, 1989b: Venus 48(2):62, fig. 1.
HOLOTYPE: LACM 2341 (dry), 3585 m, off Lincoln Co., Oregon (44 58.1 N, 132 14.7 W),
Leg.: Oregon State Univ., 6 OCT 1972.
Remarks: Upper specimen of fig.1 (anterior portion of right valve) missing.
disa, Cardiomya (Labromysa) Bernard, 1989b: Venus 48(2):64-65, fig. 4.
HOLOTYPE: LACM 2338 (dry), 3700 m, W of Patton Ridge, southern California (32 22 N,
120 42 W), Leg.: Scripps Inst. Oceanog., 6 FEB 1981.
Remarks: Currently in the genus Bathyneaera fide Coan & others (2000).
filatovae, Cuspidaria Bernard, 1979a: Venus 38(1):14-16, fig. 1.
HOLOTYPE: LACM 1906 (dry), 3932 m, NW of Newport, Lincoln Co., Oregon (45 01.1 N,
135 13.6 W), Leg.: Oregon State Univ., OCT 1972.
Remarks: Currently in the genus Rhinoclama fide Coan & others (2000).
isolirata, Cardiomya Bernard, 1969: Jour. Fish. Res. Board Canada 26(8):2231-2232, fig. 1.
HOLOTYPE: LACM 2075 (dry), 60 m, off Pt. Loma, San Diego Co., California (32 32.27 N,
117 15.47 W), Leg.: R/V Velero III, 23 FEB 1941.
Remarks: Ex AHF 163. Junior synonym of Cardiomya pectinata (Carpenter, 1865) fide Coan &
others (2000).
marmorea, Cuspidaria (Rhinoclama) Bernard, 1989b: Venus 48(2):62, fig. 2.
HOLOTYPE: LACM 2340 (dry), 3950 m, W of Patton Escarpment, southern California
(31 37 N, 120 18.3 W), Leg.: Scripps Inst. Oceanog., 23 MAR 1970.
Remarks: Junior synonym of Rhinoclama filatovae Bernard, 1979 fide Coan & others (2000).
okutani, Cardiomya behringensis Scarlato, 1972: Trudy Zool. Inst. Akad. Nauk (Leningrad)
52:122
PARATYPE: LACM 2651 (1 dry), 120-135 m, Ostrova Askolda, Zaliv Petra Valikogo (= Peter
the Great Bay), Sea of Japan, Primorye Kray, Russia (42 46 N, 132 20 E), JUN-JUL 1932.
Remarks: Junior synonym of Cardiomya behringensis (Leche, 1883) fide Coan & others (2000).
parapodema, Cuspidaria Bernard, 1969: Jour. Fish. Res. Board Canada 26(8):2232, fig. 2.
HOLOTYPE: LACM 2076 (dry); PARATYPE: LACM 2339 (1 dry), 79-155 m, off Redondo
Beach, Los Angeles Co., California (33 49.0 N, 118 26.0 W), Leg.: R/V Velero III, 5 MAY
1940.
Remarks: Holotype ex AHF 162.
variola, Cuspidaria Bernard, 1979a: Venus 38(1):16-17, fig. 2.
HOLOTYPE: LACM 1907 (dry), 2850 m, NW of Newport, Lincoln Co., Oregon (45 01.6 N,
127 31 W), Leg.: Oregon State Univ., DEC 1973.
Superfamily VERTICORDIOIDEA Stolczka, 1871
Family Verticordiidae Stoliczka, 1871
aequacostata, Verticordia Howard, 1950: The Nautilus 63(4):109-110, pl. 7.
HOLOTYPE: LACM 1752 (dry); PARATYPE: LACM 1753 (1 dry), 165 m, N of Isla Angel de
la Guarda, Golfo de California, México, Leg.: R/V Velero III ; PARATYPE: LACM 1754 (1
dry), 192 m, off Whites Cove, Santa Catalina Id., California Channel Ids., California;
PARATYPE: LACM 2080 (1 dry), 165 m, N of Isla Angel de la Guarda, Golfo de California,
México, Leg.: R/V VELERO III, 21 MAR 1937.
Remarks: Holotype ex AHF 4371. Currently in the genus Haliris fide Coan & others (2000).
hancocki, Verticordia Bernard, 1969: Jour. Fish. Res. Board Canada 26(8):2233-2234, fig. 6.
HOLOTYPE: LACM 2079 (dry); PARATYPE: LACM 2268 (1 dry), 73-110 m, off Isla de
Gorgona, Cauca Dept., Colombia (3 00.37 N, 78 19.78 W), Leg.: R/V Velero III, 24 FEB 1938.
Remarks: Holotype ex AHF 165. Currently in the genus/subgenus Trigonulina
(Verticordia).
pacifica, Trigonulina Jung, 1996: Bulls. Amer. Paleont. 110(351):48-51, pl. 2, figs. 11-12.
HOLOTYPE: LACM 2718 (dry); PARATYPES: LACM 2762 (9 dry), 82 m, 0.4-0.7 mi. 110 to
132 degrees T from Ship Rock, Santa Catalina Id., California Channel Ids., California (33 27 N,
118 30 W), Leg.: R. Reimer et al., R/V Velero IV, 13 FEB 1965; PARATYPES: LACM 2850 (3
dry), 60 m, Humpback Rock, off Hopkins Marine Station, Pacific Grove, Monterey Co,
California (36 38 N, 121 54 W), Leg.: J.H. McLean, R/V Tage, 26 NOV 1963; PARATYPES:
LACM 2851 (3 dry), 99-102 m, NW of Cavern Pt., Santa Cruz Id., California Channel Ids.,
California (34 04 N, 119 34.4 W), Leg.: R/V Velero III, 12 APR 1941; PARATYPES: LACM
2852 (2 dry), 27-91 m, Anacapa Passage, W of Anacapa Id., California Channel Ids., California
(33 59.0 N, 119 32.1 W), Leg.: R/V Velero III, 30 OCT 1940; PARATYPE: LACM 2853 (1
dry), 62-75 m, S of Gull Id., Santa Cruz Id., California Channel Ids., California (33 56.5 N,
119 49.6 W), Leg.: R/V Velero III, 11 APR 1941; PARATYPES: LACM 2854 (4 dry), 75 m, 4
mi. N of Islas Todos Santos, Pacific Coast Baja California, México (31 53.3 N, 116 48.3 W),
Leg.: R/V Velero III, 24 FEB 1941; PARATYPES: LACM 2855 (12 dry), 27 m, W of Isla Smith,
Bahía de los Angeles, Baja California, Golfo de California, México (29 04 N, 113 33 W), Leg.:
G. Sphon & D.K. Mulliner, 10 OCT 1975; PARATYPES: LACM 2856 (21 dry), 18-22 m, W of
Isla Smith, Bahía de los Angeles, Baja California, Golfo de California, México (29 03.7 N,
113 31.0 W), Leg.: G. Sphon & D.K. Mulliner, 10-16 MAY 1976; PARATYPES: LACM 2857
(25 dry), 31-37 m, Kellett Channel, S of Isla Cedros, Pacific Coast Baja California Sur, México
(27 57.0 N, 115 08.5 W), Leg.: J.H. McLean & P.I. LaFollette, R/V Searcher, 20 OCT 1971;
PARATYPES: LACM 2858 (24 dry), 43-55 m, of Isla Danzante, Bahía Escondido, Baja
California Sur, Golfo de California, México (25 46 N, 111 15 W), Leg.: DF.K. Mulliner & G.
Sphon, 6 NOV 1978; PARATYPES: LACM 2859 (13 dry), 27-37 m, off Punta Arena de la
Ventana, Baja California Sur, Golfo de California, México (24 04 N, 109 49 W), Leg.: J.H.
McLean, P.M. Oringer, & L. Marincovich, 8 APR 1966; PARATYPES: LACM 2860 (6 dry),
18-55 m, off anchorage at Bahía de los Muertos, Baja California Sur, Golfo de California,
México (23 58 N, 109 46 W), Leg.: J.H. McLean et al., 8 APR 1966; PARATYPES: LACM
2861 (6 dry), 18-37 m, between Rancho El Tule and Rancho Palmilla, Baja California Sur, Golfo
de California, México (22 58 N, 109 45 W), Leg.: J.H. McLean & P.M. Oringer, 5 APR 1966;
PARATYPES: LACM 2862 (10 dry), 37-73 m, Bahía Banderas, Jalisco, México (20 40 N,
105 25 W), Leg.: G. Willett, 14 FEB 1938; PARATYPES: LACM 2863 (31 dry), 26-33 m,
Bahía Braithwaite, Isla Socorro, Islas Revillagigedo, México (18 42.5 N, 110 56.22 W), Leg.:
R/V Velero III, 3 JAN 1934; PARATYPES: LACM 2864 (11 dry), 37 m, 0.5-1.5 mi. W of Roca
Vagares, Bahía Juanillo, Guanacaste Prov., Costa Rica (10 57.47 N, 85 45.3 W), Leg.: D.
Cadien & P.I. LaFollette, R/V Searcher, 14 FEB 1972; PARATYPES: LACM 2865 (27 dry),
37 m, off Bahía Herradura, Puntarenas Prov., Costa Rica (9 38.8 N, 84 40.8 W), Leg.: J.H.
McLean & W. Bussing, R/V Searcher, 10 MAR 1972; PARATTPES: LACM 2866 (15 dry), 21
m, anchorage at Bahía Herradura, Puntarenas Prov., Costa Rica (9 37.97 N, 84 40.5 W), Leg.:
J.H. McLean, R/V Searcher, 9 MAR 1972; PARATYPES: LACM 2867 (12 dry), 21 m,
anchorage inside small inlet 1.5 km S of Punta Quepos, Puntarenas Prov., Costa Rica (9 22.72 N,
84 09.68 W), Leg.: J.H. McLean, R/V Searcher, 11 MAR 1972; PARATYPES: LACM 2868 (8
dry), 73-128 m, Bahía Guatulco, Oaxaca, México, Leg.: G. Willett, 7 MAR 1938.
quaylei, Lyonsiella Bernard, 1969: Jour. Fish. Res. Board Canada 26(8):2232-2233, fig. 3.
HOLOTYPE: LACM 2077 (dry), 664 m, off Santa Catalina Id., California Channel Ids.,
California (33 20.02 N, 118 13.97 W), Leg.: R/V Velero IV, 25 APR 1954.
Remarks: Ex AHF 164.
spinosa, Verticordia Bernard, 1969: Jour. Fish. Res. Board Canada 26(8):2233, fig. 5.
HOLOTYPE: LACM 2078 (dry), 274 m, Bahía de Palmas, Golfo de California, Baja California
Sur, México (23 38.20 N, 109 30 W), Leg.: R/V Velero III, 21 JAN 1940
Remarks: Ex AHF 166. Currently in the genus Haliris.
Superfamily POROMYOIDEA Dall, 1886
Family Poromyidae Dall, 1886
hyalina, Poromya (Dermatomya) Bernard, 1989b: Venus 48(2):66, fig. 6.
HOLOTYPE: LACM 2342 (dry), 4882 m, Tufts Abyssal Plain off Oregon, East Pacific
(44 59.2 N, 147 40.8 W), Leg.: Oregon State Univ., 14 AUG 1976.
Remarks: Currently in the genus Dermatomya fide Coan & others (2000).
ochotensis, Poromya granuloderma Scarlato, 1981: Guide to the fauna of the USSR, p. 429.
PARATYPES: LACM 2633 (3 dry), 418 m, W of Ozernovskiy, Sea of Okhotsk, Kamchatka
Oblast, Russia (51 35.10 N, 154 46 E), Leg.: P. Ushakov; R/V Gagara, 10 JUL 1932.
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