Zootaxa 3780 (1): 051–107
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Article
Copyright © 2014 Magnolia Press
ZOOTAXA
ISSN 1175-5334 (online edition)
http://dx.doi.org/10.11646/zootaxa.3780.1.3
http://zoobank.org/urn:lsid:zoobank.org:pub:74DE381B-0185-4735-8CCB-60AD25E0DA01
Catalogue of the type specimens deposited in the Mollusca Collection of the
Museu Nacional / UFRJ, Rio de Janeiro, Brazil
ALEXANDRE DIAS PIMENTA1,4, JÚLIO CÉSAR MONTEIRO2, ANDRÉ FAVARETTO BARBOSA3,
NORMA CAMPOS SALGADO1 & ARNALDO CAMPOS DOS SANTOS COELHO1
1
Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro (UFRJ)
Instituto de Estudos do Mar “Almirante Paulo Moreira” (IEAPM)
3
Instituto Brasileiro do Meio Ambiente e dos Recursos Naturais Renováveis (IBAMA)—Coordenação-Geral de Petróleo e Gás, Rio de
Janeiro
4
Correspondent author. E-mail: alexpim@mn.ufrj.br
2
Abstract
A curatorial revision of the type specimens deposited in the Mollusca Collection of the Museu Nacional / UFRJ, Rio de
Janeiro, Brazil (MNRJ) revealed the existence of 518 lots of type specimens (holotypes, neotypes, syntypes and paratypes)
for 285 names of molluscan taxa from 88 families, including 247 gastropods, 30 bivalves, three cephalopods and five scaphopods. A total of 106 holotypes and one neotype are deposited in the MNRJ. Type material for ten nominal taxa described as being deposited in the MNRJ was not located; the probable reasons are discussed. Some previously published
erroneous information about types in the MNRJ is rectified. A total of 37 type specimens are illustrated.
Key words: types, malacological collection, mollusc, malacology, nomenclature
Introduction
Founded in 1818 by D. João of Portugal as the “Museu Real”, the Museu Nacional is the oldest museum in Brazil
and also its oldest scientific institution (Museu Nacional, 2007). Since 1946, the museum has been part of the
Federal University of Rio de Janeiro (Universidade Federal do Rio de Janeiro—UFRJ) and it has academic,
scientific and educational missions, aiming to develop and disseminate knowledge of natural history and
anthropology. As part of its scientific activities, the Museu Nacional holds the most important zoological
collections in Brazil.
Following a recommendation of the International Code of Zoological Nomenclature (ICZN, 1999: 72F.4) to
encourage the publication of lists of types housed in institutions, this paper presents a catalogue of the molluscan
types held in the Mollusca Collection of the Museu Nacional.
The Mollusca Collection of the Museu Nacional
The Mollusca Collection of the Museu Nacional (acronym MNRJ), one of the most important of its kind in South
America, holds more than 32,000 registered lots and about 10,000 unregistered lots, which are currently being
processed, including dry shells and preserved specimens from Brazilian and foreign provenance. The type material
stems from studies carried out since the end of the 19th century by Brazilian and foreign researchers.
The early beginnings of this collection are difficult to establish. The earliest reference was by Netto (1870),
who mentioned that molluscs were exhibited in the first building of the Museu Nacional; followed by Lacerda
(1905), who stated that these specimens were exhibited at the present location of the museum. Miranda-Ribeiro
(1924) referred to the mollusc collection, which at that time contained 584 Brazilian species, as determined by E.
von Martens from “Zoologisches Museum”, presently “Museum für Naturkunde”, in Berlin.
Accepted by A. Nutzel: 7 Feb. 2014; published: 20 Mar. 2014
51
In the 1910s and 1920s, the naturalist Bourgny de Mendonça curated the collection. In the early 1930s, Paulo
de Miranda-Ribeiro established the first catalogue, with about 6,230 records of molluscan specimens included in
the Zoological Collection of the Museu Nacional. This catalogue used the old numbering system of the Mollusca
Collection, which ranged from 30,001 to 36,302. In 1942, the naturalist Emmanoel de Azevedo Martins, from
“Divisão de Geologia e Minearologia” was responsible for managing the Mollusca Collection, initiating a new
numbering system, which has consistently been followed until since. From 1950 on, the curators were,
successively, Fausto Luiz de Souza Cunha and Arnaldo Campos dos Santos Coelho, with the important
collaboration of Dr. Hugo de Souza Lopes. Arnaldo Campos dos Santos Coelho took part in the restructuring of the
physical space for the collection and continued to develop it until his retirement in 1996, when the collection was
transferred to the responsibility of Norma Campos Salgado. She retired in 2008, transferring the collection to its
present curator, Alexandre Dias Pimenta.
In 1970, Hugo de Souza Lopes, a dipterologist from the Instituto Oswaldo Cruz in Rio de Janeiro (IOC),
donated his entire private collection of molluscs, which was stored in IOC, to the MNRJ molluscan collection. This
collection, containing more than 7,500 lots, had been catalogued by Hugo de Souza Lopes using his own
numbering system, and since much of that material had been listed in various publications (sometimes with IOC
acronym), the lots were entered into the MNRJ molluscan collection catalogue with their original collection
numbers, with the added acronym HSL. In addition to being a renowned entomologist, Hugo de Souza Lopes was
an enthusiast of molluscan taxonomy, which led him to establish contact with many malacologists throughout the
world, resulting in an extensive exchange of material. For this reason, the MNRJ molluscan collection includes a
large number of specimens, including some types that were originally listed as belonging to other collections.
Exchange of paratypes was a common practice in the 1970s, mainly with other Brazilian collections, including
the present Museu Oceanográfico do Rio Grande, Rio Grande do Sul state (MORG) and the Museu de Zoologia,
São Paulo state (MZSP), and also with institutions in other countries, such as the Museo Argentino de Ciencias
Naturales "Bernardino Rivadavia" in Buenos Aires, Argentina (MACN) and the Museo Nacional de Historia
Natural y Antropología, in Montevideo, Uruguay (MNHNM).
A particular case regarding donation of type material to MNRJ molluscan collection involves some taxa
described by Wolfgang Weyrauch in a series of papers describing the terrestrial gastropods from Peru (Weyrauch,
1957, 1960a, b, 1967a, b). Weyrauch deposited several paratypes in his private collection and often sent duplicate
material to museums and private collectors throughout the world (Barbosa et al., 2008).
Material and methods
All material stored in the MNRJ Mollusca Collection and labeled as any type category was examined and
compared with the original descriptions of their names, as well as with information from the labels and catalogue
books of the collection. Topotypes are not included in this catalogue. Only taxa names published until 2013
(including) are considered.
The names of the taxa are presented in their original spelling, arranged by class and organized in alphabetical
order by the lowest taxonomic category, followed by the original generic affiliation. This entry is followed by
information, in brackets, regarding the currently adopted generic (when different from the original) and family
classification, as well as the valid name (if the original name was preoccupied and subsequently replaced). The
family classification used here follows Bouchet et al. (2005) for Gastropoda and Bieler et al. (2010) for Bivalvia,
both with the modifications for marine taxa suggested in WoRMS database (Appeltans et al. 2013); WoRMS
database (WoRMS Editorial Board 2013) for Scaphopoda, Cephalopoda and Polyplacophora. Appendix 1 lists all
names in systematic order, using the currently adopted generic combination, which is mostly based on WoRMS
database (Appeltans et al. 2013) for marine species; Simone (2006a) for non-marine Brazilian species; Ramirez et
al. (2003), for non-marine gastropods from Peru. It is beyond the scope of this paper to discuss classifications at
genus or family levels, or synonymies of the names; also, in the few cases where syntypes exist, lectotypes have
not been designated.
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Each presented entry encompasses three sections:
- data for the species description: Author and year of description, along with the published page number where
the species name was introduced (except for the title and abstract) and indication of illustrations in the original
publication. Next, the type locality is given, based on the original publication and/or the original label of the
holotype or lectotype (if it is deposited in the MNRJ collection); in some cases, a general complement (e.g.,
country, state, region, etc.) is added; data for geographical coordinates and depth (dep) or altitude (alt) are included
only when originally provided;
- the type status of the specimens currently stored in the MNRJ Mollusca Collection, followed by the collection
number, are given. For paratypes, the number of specimens in each lot is given, as well as their collection locality,
based on the original publication and/or data on the labels; in some cases, a general complement (e.g., country,
state, region, etc.) is added. The type localities for the nominal taxa described by Weyrauch (1957, 1960a, b, 1967a,
b) follows Breure (2012). For each lot, the state of preservation and related information are given in brackets, with
the following conventions: shell (sh)—when only the shell is preserved, without soft parts; specimen (spm) - for a
complete specimen; valve—when the valve(s) of Bivalvia shell(s) are separated. In some cases of dissected
fragments of soft parts, mounted radulae or laminae, if necessary, an appropriate remark is added. Whenever
necessary, a new collection number is provided, and the reasons are discussed in the remarks section;
- remarks: When necessary, this section presents a brief discussion about the types, indicating any discrepancy
between the material found in the collection and the information published in the original description of the taxon,
such as the number of paratypes, the use of a different acronym for the MNRJ collection, the lack of a collection
number etc. When necessary, a decision about the type status of the specimens is taken, following the rules and
recommendations of the ICZN (1999). In some cases, paratype lots in the MNRJ collection were not explicitly
cited in the original publication. In these cases, we followed the ICZN (1999: Article 72.4.1.1) which states that,
“for a nominal species or subspecies established before 2000, any evidence, published or unpublished, may be
taken into account to determine what specimens constitute the type series”; in most cases, information on the label
was considered, always in comparison with information in the original publication, such as dimensions of the
specimes, localities, date of collecting etc. For cases of names published after 2000, in which a specimen deposited
in the MNRJ collection is labeled as paratype but was not mentioned in the original description, it is not considered
as a type. In some cases, a lot cited in the original publication as paratype lot, contains a greater number of
specimens than originally expressed; strictly, according to ICZN (1999), only those specimens cited as paratypes
should be considered types, but in such cases, it is impossible to determine which were the actual paratypes;
although these cases could be considered lapsus calami by the authors when referring to the lot in the original
description, we preferred to keep them as non types.
For some species, photographs of some type specimens (Figures 1–37) are presented, in the following cases:
holotype not illustrated, for which the illustration errs in some way (e.g. with wrong or incomplete legend),
illustrated by a drawing or in a poor figure, or illustrated in rare publications. In any case in which a type from the
MNRJ was illustrated in a subsequent publication, this publication is cited.
Abbreviations used: Collections: ANSP—Academy of Natural Sciences of Philadelphia, USA; CPMSC—
Coleção “Paulo Marcio Santos Costa”, private;—IOC Instituto Oswaldo Cruz, Rio de Janeiro; MACN—Museo
Argenino de Ciencias Naturales “Bernardino Rivadavia”, Buenos Aires, Argentina; MNHNM—Museo Nacional
de Historia Natural y Antropologia, Montevideo, Uruguay; MNRJ—Museu Nacional/Universidade Federal do Rio
de Janeiro, Brazil; MORG—Museu Oceanográfico do Rio Grande, Rio Grande, Brazil; MZSP—Museu de
Zoologia da Universidade de São Paulo, São Paulo, Brazil; collecting expeditions and others: AMASSEDS—A
Multidisciplinary Amazon Shelf Sedimentary Study; BC—Bacia de Campos; CF VII—Comissão Oceanográfica
Cabo Frio VII, 1983, Oceanographic Ship Almirante Saldanha coll.; HAB—Projeto Habitats—“Heterogeneidade
Ambiental da Bacia de Campos” (Habitats Project—Campos Basin Environmental Heterogeneity), carried out by
CENPES/PETROBRAS; JOPS—Joint Oceanographic Projects, Research Vessel "Victor Hansen" coll; MD55—
Oceanographic cruise Marion Dufresne, southeast Brazil, 1987; Lima—Programa de Geologia Marinha do Brazil,
Oceanographic Ship Almirante Câmara coll.; PADCT—Importance and Characterization of edge of Continental
Shelf to Living and Nonliving Resources (“Programa de Apoio ao Desenvolvimento Científico e Tecnológico”),
carried out by Instituto Oceanográfico da Universidade de São Paulo, 1997−1998; PCABS—Projeto de
Caracterização Ambiental da Bacia de Sergipe (Sergipe Basin Environmental Caracterization Project);
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REVIZEE—Program of Evaluation of the Sustainable Potential of Living Resources in the Economic Exclusive
Zone (“Recursos Vivos da Zona Econômica Exclusiva”), carried out by Ministério do Meio Ambiente, 1996−2002;
WW—“Wolfgang Karl Weyrauch” private collection.
Results
The 518 type lots from MNRJ Mollusca Collection belong to 247 names of taxa in the class Gastropoda, 30 taxa in
the class Bivalvia, three names in the class Cephalopoda, and five names in the class Scaphopoda (not including the
non located type material). Of these, 13 names refer to freshwater taxa (ten gastropods and three bivalves), 32
terrestrial gastropods, and 240 marine molluscan taxa.
Type specimens for 10 species mentioned in taxonomic publications to be deposited in the MNRJ Mollusca
Collection were not found. The reasons for this are not always easy to uncover. In most cases, the lots were never
received by the MNRJ collection, and their whereabouts remains unknown.
A total of 106 holotypes and one neotype are present in the collection, in addition to several paratypes and
some syntypes. Most of the names (269) concern species described from type localities in Brazil, with a few types
from other countries, especially terrestrial pulmonates from Peru (13 names).
Type catalogue
Class Gastropoda
abrolhosensis, Acmaea (Collisella) [Lottia, Lottiidae]. Petuch, 1979: 513, figs. 1A−B. Type locality: Santa
Barbara Island, Abrolhos Archipelago (17°58’S, 38°42’W), Bahia state, Brazil.
Paratypes. MNRJ 3733 [5 sh], type locality (Fig. 2).
Remarks. The original publication mentioned two different latitudinal coordinates for the type locality
(17°58’S and 19°57’S); the second was a clear lapsus calami, since the correct latitude for the Abrolhos
Archipelago is 17°58’S. The original publication listed three paratypes in MNRJ 3733, but the lot contains five
shells.
aembe, Neopleurotomoides [Raphitomidae]. Figueira & Absalão, 2012: 8, figs. 5−8. Type locality: Campos Basin,
Southeast Brazil, Southwestern Atlantic (22°11’16"S, 39°43’44"W, 1968 m dep).
Holotype. MNRJ 30285 [sh]. Paratype: MNRJ 30286 [1 sh], type locality.
albertoi, Caelatura [Barleeidae]. Santos & Absalão, 2006a: 121, figs 1−7. Type locality: Off Amapá state, Brazil,
AMASSEDS station 4134 (02°21’00"N, 48°29’00"W, 72 m dep).
Holotype. MNRJ 10305 [sh]; paratypes: MNRJ 10306 [5 sh], type locality.
albovittata, Mitrella [Mokumea, Columbellidae]. Lopes, Coelho & Cardoso, 1965: 21−23, figs. 1−2. Type locality:
Itapuã, (farol), Salvador, Bahia state, Brazil.
Holotype. MNRJ HSL6502 [sh] (Fig. 20); paratypes: MNRJ 3030 [1 sh], type locality; MNRJ 3031 [2 sh],
Praia do Pontal, Cabo Frio, Rio de Janeiro, Brazil; MNRJ HSL6503 [1 sh], type locality; MNRJ HSL6504 [3 sh],
type locality; MNRJ HSL6505 [1 sh], Ponta Verde, Maceió, Alagoas, Brazil.
Remarks. Holotype and paratypes originally stated to be deposited in the IOC malacological collection were
later donated to the MNRJ.
alvesi, Gerdiella [Cancellariidae]. Lima, Barros & Petit, 2007: 100, figs. 1−5. Type locality: Northeastern Brazil,
off the state of Pernambuco (08°46.5′S, 34°44.5′W, 690 m dep).
Paratypes. MNRJ 10718 [2 sh], type locality.
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amandus, Megalobulimus [Strophocheilidae]. Simone, 2012a: 436, figs. 22–26. Type locality: Santa Maria da
Vitória, Bahia state, Brazil (~13°24’S, 44°12’W, ~460 m alt).
Paratype. MNRJ 30380 [1 sh], type locality.
Remarks. Original description did not provide catalogue number for the paratypes.
amphorale, Prunum [Marginellidae]. De Souza, 1992: 239, figs. 2−3. Type locality: Off Pará state, Brazil
(01°52′27"S, 48°16′12"W, 47 m dep).
Paratype. MNRJ 6205 [1 sh], off Pará state, Brazil (00°12′S, 47°03′W, 33 m dep).
analuciae, Bullata [Marginellidae]. de Souza & Coovert, 2001: 5, figs. 2, 8, 11. Type locality: Off Vitória, Espírito
Santo state, Brazil (20°20’S, 40°00’W, 60−70 m dep).
Holotype. MNRJ 7186 [sh]; paratype: MNRJ 7187 [1 sh], type locality.
anfractura, Metula [Buccinidae]. Matthews & Rios, 1968: 151, figs. 1−4. Type locality: Off Tramandaí, Rio
Grande do Sul state, Brazil (30°05’S, 49°31’W, 85 m dep).
Paratype. MNRJ 3489 [1 sh], off Rio Grande, Rio Grande do Sul state (32°45’S, 50°43’W, 87 m dep), Brazil.
angularis, Corinnaeturris [Turridae]. Figueira & Absalão, 2010a: 739, figs. 2G, H, I, J. Type locality: Campos
Basin, southeast Brazil (21°57’17"S, 39°56’01"W, 750 m dep).
Paratype. MNRJ 17882 [1 sh], (21°57’11"S, 39° 56’04"W, 698 m dep), Campos Basin, southeast Brazil.
antares, Mitrella [Astyris, Columbellidae]. Costa & de Souza, 2001: 18, figs. 5−8, 12−14. Type locality: Off
Guarapari, Espírito Santo state, Brazil (20°41’S, 40°22’W, 25 m dep).
Paratypes. MNRJ 7185 [1 sh], Marataízes, Espírito Santo state (20°10’S, 40°37’W, 30 m dep), Brazil; MNRJ
7257 [1 sh], Itapuã, Salvador, Bahia state, Brazil.
aquilonaria, Turbonilla [Pyramidellidae]. Absalão, Santos & Tenório, 2003: 8, figs. 15−17. Type locality: Near
Cabo de Santo Agostinho, Pernambuco state, northeast coast of Brazil (08°10’30"S, 34°42’01"W, intertidal).
Paratype. MNRJ 9826 [1 sh], north coast of Brazil.
aquilonia, Oscilla [Pyramidellidae]. Pimenta, Santos & Absalão, 2008: 176, fig. 2G−L. Type locality: Off Pará
state, north Brazilian coast − AMASSEDS station 3228 (03°25.1’N, 49°56.4’W, 64 m dep).
Holotype. MNRJ 10825 [sh]; paratypes: MNRJ 10819 [2 sh], type locality.
Remarks. The original publication listed three shells in the lot MNRJ 10819, but only two shells were found.
aracruzensis, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2004a: 124, 126, figs. 61−65. Type locality: Off
Aracruz, Espírito Santo state, southeast coast of Brazil.
Holotype. MNRJ 8928 [sh]; paratypes: MNRJ 8921 [1 sh], Camburi, Espírito Santo state, Brazil; MNRJ 8929
[1 sh], type locality.
arionata, Olivella [Olivellidae]. Absalão, 2000: 11, figs. 1, 3−6. Type locality: Off state of Rio de Janeiro, southern
Brazil (21°40’117 S, 40°48’004 W, 18 m dep), Brazil.
Paratype. MNRJ 8289 [1 spm mummified], type locality.
arnaldoi, Epitonium [Epitoniidae]. Tursch & Pierret, 1964: 36, fig. 4. Type locality: Off Punta de Juatinga, Brazil
(23°22’S, 48°28’W, 50 m dep).
Paratype. MNRJ 3118 [1 sh], type locality (Fig. 15).
arnaldoi, Protoglyptus [Naesiotus, Bulimulidae]. Lanzieri & Rezende, 1971: 255, figs. 1−45. Type locality: Pico
do Desejado, Ilha da Trindade, Espírito Santo state, Brazil.
Holotype. MNRJ 3105 [spm] (Fig. 35); paratypes: MNRJ 3430 [109 sh + 3 fragments], Praia dos Cabritos,
Ilha da Trindade, Espírito Santo state, Brazil; MNRJ 3429 [8 sh], Praia do Andrada, Ilha da Trindade, Espírito
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Santo state, Brazil; MNRJ 3426 [79 shells + 3 fragments], type locality; MNRJ 3428 [29 sh], Praia dos Cabritos,
Ilha da Trindade, Espírito Santo state, Brazil; MNRJ 3427 [26 sh], type locality; MNRJ 7874 [51 spms + 4 sh +
fragments of soft parts of dissected spm], type locality.
Remarks. The holotype shell is somewhat corroded. Simone (2006a: 125, fig. 395) figured a paratype, but
without providing the collection number (probably MNRJ 3426). Salvador et al. (2013) figured the holotype and a
paratype reported by those authors as being deposited with the same collection number as the holotype; however,
MNRJ 3426 contains only the holotype.
atlanticus, Caducifer [Monostiolum, Buccinidae]. Coelho, Matthews & Cardoso, 1970: 185, figs. 1−3. Type
locality: Praia do Andrada, Trindade Island, Espírito Santo state, Brazil.
Holotype. MNRJ 3550 [sh]; paratypes: MNRJ 3548 [1 sh], Mucuripe beach, Fortaleza, Ceará state, Brazil;
MNRJ 3549 [1 sh], Ilha da Trindade, Brazil; MNRJ 3551 [1 sh], Praia da Enseada da Cachoeira, Fortaleza, Ceará
state, Brazil.
Remarks. Photographs of the holotype and of the paratype in the lot MNRJ 3548 were published by Watters
(2009). Gomes et al. (2006: 195, fig. B5) also figured the holotype without mentioning type status or collection
number of the illustrated shell.
atlanticus, Leptotrophon [Muricidae]. Pimenta, Couto & Costa, 2008: 248, figs. 13−26. Type locality: Canopus
Bank, 96 miles off Ceará state (240–260 m dep), Brazil.
Holotype. MNRJ 11004 [spm and mounted radula]; paratypes: MNRJ 11009 [3 spm], type locality.
aulakion, Caelatura [Barleeidae]. Santos & Absalão, 2006a: 122, figs. 8−13. Type locality: Amapá state, Brazil
(04°27’00"N, 49°58’00"W, 160 m dep).
Holotype. MNRJ 10308 [sh].
Remarks. MNRJ 10312 [3 sh], from the type locality, contains a label written by the authors indicating
“paratype”, but this was not cited in the original publication and therefore, according to ICZN (1999: Article
72.4.1.1), it is here considered as a topotype.
aurisflamma, Annulobalcis [Eulimidae]. Simone & Martins, 1995: 224, figs. 1−25. Type Locality: Enseada
Beach, Ubatuba, São Paulo state, Brazil (23°29’S, 45°05’W).
Paratypes. MNRJ 6374 [1 spm], type locality; MNRJ 6375 [1 spm], type locality.
baiano, Conus [Conidae]. Coltro, 2004: 4−5, pl. 4, fig. A, pl 12, figs. H1, P1−11, pl 16, fig. F. Type locality: 25 km
SW Abrolhos Archipelago, off Alcobaça, Bahia state, Brazil (17°24’S, 38°20’W).
Paratypes. MNRJ 10194 [1 sh], type locality; MNRJ 10195 [1 sh], type locality.
baixoguanduensis, Thaumastus (Thaumastus) [Bulimulidae]. Pena, Coelho & Salgado, 1996: 1, figs. 1, 13, 3−11.
Type locality: Baixo Guandu, Espírito Santo state, Brazil (19°32’09"S, 41°01’06"W).
Holotype. MNRJ 6924 [sh + dissected soft parts]; paratypes: MNRJ 6925 [23 sh], type locality; MNRJ 6926
[2 sh + 23 spms + dissected soft parts], type locality; MNRJ 6927 [47 spms + 5 soft parts], type locality; MNRJ
6928 [24 spms + dissected soft parts], type locality; MNRJ 6929 [3 sh + 47 spms + dissected soft parts], type
locality; MNRJ 6930 [20 spmns + dissected soft parts], type locality; MNRJ 6931 [2 sh + 7 spms + dissected soft
parts], type locality; MNRJ 6932 [1 sh + dissected soft parts], Fazenda da Serra, Espírito Santo state, Brazil.
Remarks. The original publication listed 26 shells in lot MNRJ 6925 and, as result of a lapsus calami,
reversed the photographs of the dorsal views of the holotype of this species and the other species described in that
paper; Pena et al. (1996: page 7, fig. 2) is actually a photograph of the holotype of Thaumastus tiradentensis in
dorsal view, while Pena et al. (1996: page 10, fig. 13) is the photograph of the holotype of T. baixoguanduensis is
dorsal view.
Simone (2006a: 152, fig. 516) figured a shell referred to as holotype MNRJ 6931; however, the shell illustrated
is actually a paratype from MNRJ 6931.
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balaustium, Cerithiopsis [Cerithiopsidae]. Figueira & Pimenta, 2008: 75, figs. 19−23. Type locality: Off São Paulo
state, southeast coast of Brazil (24°17.129’S, 44°12.149’W, 163 m dep).
Paratype. MNRJ 10981 [1 sh], PADCT station 6577 (25°15.76’S, 45°04.62’W, 124 m dep), off São Paulo
state, Brazil.
barcellosi, Bursa [Ranella, Ranellidae]. Matthews, Rios & Coelho, 1973: 52, figs. 1−5. Type locality: Off Torres,
Rio Grande do Sul state, Brazil, 280 m.
Paratype. MNRJ 3647 [1 sh], off Conceição, Rio Grande do Sul state (31°50’S, 49°58’W, 260 m dep), Brazil
(Fig. 13).
barcellosi, Caelatura [Barleeidae]. Absalão & Rios, 1995: 89, figs. 4−7. Type locality: Off Bahia state,
northeastern Brazil (50 m dep).
Paratype. MNRJ 7159 [1 sh], Atol das Rocas, Bahia state, Brazil (3m dep).
barrosi, Brasityphis [Muricidae]. Absalão & Santos, 2003: 2, figs. 1−7. Type locality: Off the coast of Sergipe
state, northeastern Brazil (11°23’48"S, 37°04’48"W, 90 m dep).
Paratypes. MNRJ 9854 [6 sh, but see remarks], station 2,3 PCABS 37°05’08"S, 11°24’14"W, 99 m dep.
Remarks. The original publication stated that MNRJ 9854 corresponds to “paratye 5”, but the lot contains five
additional shells, all labelled by the authors as “paratypes”. According to the ICZN (1999: Article 72.4.1.1) only
one, originally expressly referred to as paratype is considered as a paratype, but it is impossible to determine which
specimen that is. All other specimens are topotypes.
beckeri, Oxyloma [Succineidae]. Lanzieri, 1966: 182, figs. 5−7, 67−79. Type locality: Pico S. Bonifácio, Ilha da
Trindade, Espírito Santo state, Brazil.
Paratypes. MNRJ 3177 [5 spms + 1 spm dissected + 5 sh], type locality (Fig. 31).
Remarks. The holotype (published as MNRJ 3177) was not located; it has probably been mixed-up with the
paratypes. It is not possible to identify the holotype from the information given in the original publication or the
catalogue of the collection. The shell shown in the photograph provided by Simone (2006a: 237, fig. 911) and
referred to as “holotype MNRJ 3177”, is actually one of the paratypes. The original publication stated that the lot
MNRJ 3177a bears 12 specimens (four preserved in ethanol), eight dissected for anatomical and microanatomical
studies and 37 shells. Gomes et al. (2006: 195, fig. C6) figured another paratype, but did not mention type status or
collection number of the illustrated shell.
beckyae, Latirus [Hemipolygona, Fasciolariidae]. Snyder, 2000: 161, figs. 1. Type locality: Off Vitória, Espírito
Santo state, Brazil (30−50 m dep).
Paratype. MNRJ 7696 [1 sh], type locality.
belmonti, Synaptocochlea [Trochidae]. Simone, 2009a: 14, figs. 13−17. Type locality: Belmonte Island, São Pedro
e São Paulo Archipelago (00°51’01"N, 29°20’44"W), Brazil.
Paratypes. MNRJ 12869 [3 sh, but see remarks], type locality.
Remarks. The original publication mentioned only two shells in the lot MNRJ 12869, but the lot includes
three shells. According to ICZN (1999: Article 72.4.1.1), only two specimens are paratypes as originally expressed.
It is not possible to identify which specimens these are.
bernardoi, Pisania [Buccinidae]. Costa & Gomes, 1998: 15, figs. 1−4. Type locality: Continental slope off the
coast of Salvador, Bahia state, Brazil.
Paratype. MNRJ 7163 [1 sh], off Guarapari, Espírito Santo state, Brazil.
bertarollae, Conus [Conidae]. Costa & Simone, 1997: 4, figs. 1−20. Type locality: Off Alcobaça, Bahia state,
Brazil.
Paratype. MNRJ 7666 [1 sh], type locality.
Remarks. The collection number was not mentioned in the original publication.
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bicincta, Pisinna [Anabathridae]. Santos & Absalão, 2004: 4, figs. 1−5. Type locality: Off the coast of Espírito
Santo, southern Brazil (19°16'05"S, 38°00'32"W, 67 m dep).
Holotype. MNRJ 10302 [sh]; paratype: MNRJ 10303 [sh], type locality.
biminiense, Thericium [Cerithium, Cerithiidae]. Pilsbry & McGinty, 1949: 13, pl. 1, figs. 6, 6a−b. Type locality:
Turtle Rocks, Binimi Island, Bahamas.
Paratypes. MNRJ HSL6151 [3 sh], type locality (Fig. 8).
Remarks. The paratypes in MNRJ were originally published as belonging to “McGinty collection”; part of
this type collection was later donated to HSL collection.
birmani, Crenilabium [Acteonidae]. Simone, 2006b: 68, figs. 1−12. Type locality: Off Ubatuba, São Paulo state,
Brazil (23°46'S, 44°51'W, 55 m dep).
Paratypes. MNRJ 10703 [9 sh], 22°18’52.95"S 40°11’58.95"W, 190−200 m.
biumbilicata, Parthenina [Pyramidellidae]. Pimenta, 2012: 445, figs. 31–36. Type locality: Continental shelf of
north of Rio de Janeiro state, southeast of Brazil, REVIZEE Central sta C1-D3 (22°52’S, 41°09’W, 80 m dep).
Paratypes. MNRJ 16306 [1 sh], continental shelf of Campos Basin, Rio de Janeiro state, southeast of Brazil,
HAB 16 sta H4 (21°42’49"S, 40°10’21"W, 97 m); MNRJ 16307 [1 sh], continental shelf of Campos Basin, Rio de
Janeiro state, southeast of Brazil, HAB 16 sta F3 (22°7’39”S, 40°18’53”W, 72 m).
bodarti, Conus [Conidae]. Coltro, 2004: 2, pl 1, fig. A, pl. 8 fig. H1, P1−P11. Type locality: 125 km NE Abrolhos
Archipelago, off Alcobaça, Bahia state, Brazil (15°50’S, 37°57’W).
Paratypes. MNRJ 10186 [1 sh], type locality; MNRJ 10187 [1 sh], type locality.
bodarti, Poirieria (Pazinotus) [Muricidae]. Costa, 1993: 50, figs. 1−3. Type locality: Between Escalvada Island
and Rasa Island, off Guarapari, Espírito Santo state, Brazil (20°41’S, 40°22’, 25 m dep).
Paratype. MNRJ 21650 (ex-CPMSC 1650) [1 sh], type locality.
Remarks. Costa & Pimenta (2012) showed that the figured shells of P. bodarti in Costa (1993: 49, fi gs. 1, 2)
cited as holotype and paratype 2, respectively, actually correspond to the same shell, the paratype 2. This paratype
was originally published in CPMSC collection and was subsequently donated to the MNRJ collection.
brasiliana, Volvarina [Marginellidae]. Boyer, 2000: 49, figs. 28−31. Type locality: Barra, Salvador, Bahia state,
Brazil.
Paratype. MNRJ 8410 [1 sh], type locality.
Remarks. The original publication did not provide the collection number.
brasilianum, Abyssochrysos [Abyssochrysidae]. Bouchet, 1991: 306, figs. 1−4, 9−11, 23. Type locality: Northeast
of Cabo de São Tomé, Brazil MD55 station CB 98 (21°35’S, 40°31’W, 900 m dep), Brazil.
Paratypes. MNRJ 6200 [2 sh], off Foz do Rio Doce, MD55 station CB93 (19°36’S, 28°53’W, 640 m dep),
Brazil.
Remarks. The original publication stated that paratypes are deposited in the “Museu Nacional de Ciencias
Naturais (Rio de Janeiro)”, which corresponds to the MNRJ, with no collection number provided. The publication
provided a list of localities from which the paratypes originated (19°36’S, 28°53’W, 640 m dep), off Foz do Rio
Doce.
brasiliensis, Onchidoris [Onchidorididae]. Alvim, Padula & Pimenta, 2011: 506, figs. 1−3. Type locality: Praia do
Forno, Arraial do Cabo, state of Rio de Janeiro, south-eastern coast of Brazil (22°58′7.69′′S, 42°0′52.41′′W, 2 m
dep).
Holotype. MNRJ 12057 [spm]; paratypes: MNRJ 13388 [1 spm, dissected], type locality; MNRJ 11759 [1
spm, dissected], type locality; MNRJ 12806 [7 spm], type locality; MNRJ 13454 [2 spm, dissected], Ilha do
Papagaio, Cabo Frio, Rio de Janeiro state (3 m dep); MNRJ 13455 [1 spm, dissected], Ilha do Papagaio, Cabo Frio,
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Rio de Janeiro state (3 m dep); MNRJ 13453 [2 spm, one dissected], Parcel da Pedra Lisa, Ilha de Búzios, São
Sebastião, São Paulo state (6 m dep).
brunoi, Loboa [Bulimulus, Bulimulidae]. Ihering, 1917: 121, Pl. 4, fig. 7. Type locality: Ilha da Trindade, Espírito
Santo state, Brazil.
Holotype. MNRJ 1128 [sh] (Fig. 34).
Remarks. MNRJ 1128 contains a label written by A. Breure in vi/1975, indicating “lectotype”; a second shell
(MNRJ 1129), collected in the same expedition to Trindade Island, also labeled by A. Breure, indicates
“paralectotype”. However, Breure (1979: 62) stated that the holotype of Bulimulus brunoi is at the MNRJ.
In fact, the original publication mentioned only one shell, clearly stated when Ihering (1917: 122) wrote that “the
unique specimen, found at the Brazilian Island of Trindade, is kept in the National Museum of Rio de Janeiro”; this
shell is therefore the holotype. Although, according to Lobo (1919), all molluscs collected in Trindade island were
sent to Hermann von Ihering, there is no evidence that the second shell (MNRJ 1129) was examined by Ihering
when describing the species. Therefore, it is a topotype only; this topotype was illustrated by Lobo (1919: 145) and
Salvador et al. (2013).
Dall (1918) stated that "the specimen, which may be the type, belongs to the National Museum of Brazil".
Simone (2006a: 117, fig. 356) figured the holotype, erroneously referred to as the lectotype. Gomes et al. (2006:
195, fig. C3) also figured the holotype, but did not mention the type status or collection number. Holotype figured
by Salvador et al. (2013).
buckuporum, Olivancillaria [Olividae]. Thomé, 1966: 163, figs. 1−8. Type locality: Bom Jesus Beach, Torres, Rio
Grande do Sul state, Brazil.
Paratype. MNRJ 6933 [1 sh], Praia do Cassino, Rio Grande, Rio Grande do Sul state, Brazil.
Remarks. The paratype in the lot MNRJ 6933 was originally published with the collection number MORG
7949 and was later donated to the MNRJ collection.
bullisi, Calliostoma (Emerlinia) [Calliostomatidae]. Clench & Turner, 1960: 36, pl. 23, figs. 1−2. Type locality:
Oregon, station 2049, about 65 miles ESE of Cabo Orange, Amapá, Brazil (04°02’N, 50°33’W, 38 fathoms (~ 70 m
dep).
Paratypes. MNRJ 12725 [2 sh], type locality general area (Fig. 3).
Remarks. The original publication stated that “there are two paratypes from the same general area in the
collection of H. Bullis”. This collection was later purchased by J. Coltro and M. Coltro, who donated it to the
MNRJ Mollusca Collection.
cabofrioensis, Mitrella [Columbellidae]. Costa & de Souza, 2001: 16, figs. 2−4, 9−11. Type locality: Off Cabo
Frio, Rio de Janeiro state, Brazil (23°18’00"S, 42°00’00"W, 140 m dep).
Paratype. MNRJ 7162 [1 sh], type locality.
cabrali, Solatisonax [Architectonicidae]. Tenório, Barros, Francisco & Silva, 2011: 178, figs. 2B−C, E−F). Type
locality: Off Alagoas state, Brazil, (10°06'S, 35°46'W, 720 m dep).
Holotype. MNRJ 13027 [sh]; paratypes: MNRJ 13028 [1 sh], off Pernambuco state, Brazil (08°46.5'S,
34°44.5'W, 690 m dep); MNRJ 13029 [1 sh] off Pernambuco state, Brazil (09°04.7'S, 34°51.2'W, 520 m dep);
MNRJ 13030 [1 sh], off Pernambuco state, Brazil (08°46.5'S, 34°44.5'W, 690 m dep).
Remarks. Cavallari et al. (2013) redescribed S. cabrali and showed that the figures 2A and 2G in the original
description were mistakenly indicated as illustrations of the holotype and the paratype MNRJ 13028, respectively.
According to Cavallari et al. (2013) figure 2A does neither match the holotype nor any of the paratypes; figure 2A
shows probably a representative of another architectonicid species. Figure 2G does not match any of the types.
New images of holotype and paratype MNRJ 13028 were provided by Cavallari et al. (2013).
caetensis, Thaumastus (Thaumastus) [Bulimulidae]. Pena, Salgado & Coelho, 2011: 533, figs. 11−21. Type
locality: Caeté (19°49’19"S; 43°40’46"W), Serra da Piedade, Minas Gerais state, Brazil.
Holotype. MNRJ 8097 [spm]; paratypes: MNRJ 8098 [3 sh + fragments of dissecetd soft parts], type locality;
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MNRJ 8099 [2 sh + fragments of dissected soft parts + 4 histological laminae], type locality; MNRJ 8100 [1 sh],
type locality; MNRJ 8101 [1 spm dissected + MEV stub], type locality; MNRJ HSL7810 [1 sh], type locality;
MNRJ 3115 [3 sh], type locality; MNRJ 9806 [12 spm + fragments of dissected soft parts + MEV stub with
protoconch + 24 histological laminae], type locality.
capixaba, Cerithiopsis [Cerithiopsidae]. Figueira & Pimenta, 2008: 77, figs. 24−28. Type locality: Off Espírito
Santo state, Brazil (20°38’S, 40°00’W, 33 m dep).
Paratypes. MNRJ 10982 [3 sh], REVIZEE station y7 (77m dep)
Remarks. The original publication did not give details about the locality of REVIZEE station y7, wich was
dredged off Espirito Santo state (20.84°S, 40.16°W, 75 m dep).
capixaba, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2004a: 131, figs. 101−106. Type locality: Off Espírito
Santo state, REVIZEE station VV38 (19°28’S, 38°22’W, 71.4 m dep), southeast coast of Brazil.
Holotype. MNRJ 8926 [sh]; paratypes: MNRJ 8937 [2 sh], type locality; MNRJ 8938 [2 sh], type locality.
carcellesi, Conus [Conidae]. Martins, 1945: 260, figs. a−g. Type locality: Mar del Plata, Buenos aires, Argentina.
Holotype. MNRJ 8462 (ex-MNRJ 36300) [sh]; paratypes: MNRJ 8463 (ex-MNRJ 36301−36302) [2 sh], type
locality (Fig. 26).
Remarks. The holotype was originally published with the collection number MNRJ 36300 and the paratypes
with collection number MNRJ 36301 and MNRJ 36302, which correspond to the old numbering system of the
collection.
carceralis, Odostomella [Pyramidellidae]. Pimenta, Absalão, & Alencar, 2000: 67, figs. 2−7. Type locality: Dois
Rios inlet, Ilha Grande, Rio de Janeiro state, Brazil (23°11’2"S, 44°11’0"W, 2−15 m dep).
Paratypes. MNRJ 8080 [2 sh], type locality.
careorugula, Olivella (Anasser) [Olivellidae]. Absalão & Pimenta, 2003: 182, figs. 1G−I. Type locality: South of
Cabo Frio, Rio de Janeiro state, Brazil, MD 55 station CB105 (23°47’S, 42°10’W, 610 m dep).
Paratypes. MNRJ 8913 [2 sh], type locality; MNRJ 8915 [2 sh], type locality.
cargilei, Conus [Conidae]. Coltro, 2004: 5−6, pl. 5, fig. A, pl 13, figs. H1, P1−11, pl. 16, fig. H. Type locality: 75
km NNE sic Abrolhos Archipelago, off Alcobaça, Bahia state, Brazil (15°57’S, 38°01’W).
Paratypes. MNRJ 10196 [1 sh], type locality; MNRJ 10197 [1 sh], type locality.
carinata, Caelatura [Barleeidae]. Santos & Absalão, 2006a: 122, figs. 14−19. Type locality: REVIZEE Nordeste
station 130A, northeast Brazil (03°20’00"S, 38°11’00"W, 71 m dep).
Holotype. MNRJ 10307 [sh].
carioca, Conus [Conidae]. Petuch, 1986: 9, figs. 1−2. Type locality: Off Cabo Frio, Rio de Janeiro state, Brazil
(100m dep).
Paratype. MNRJ 8459 [1 sh], type locality (Fig. 27).
Remarks. The paratype in the lot MNRJ 8459 was originally published with the collection number MORG
14−242 and later donated to the MNRJ collection.
carioca, Crepidula [Calyptraeidae]. Simone, 2006c: 26, figs. 22−25, 33, 107−110. Type locality: Praia de Boa
Viagem, Niterói, Rio de Janeiro state, Brazil (22°56’S, 43°12’W).
Holotype. MNRJ HSL7464 [spm]; paratypes MNRJ 9518 [84], off Santana Island, Rio de Janeiro state, Brazil
(22°30’S, 41°23’W, 48 m dep); MNRJ 2336 [4 spm], Itaipu Beach, Rio de Janeiro state, Brazil; MNRJ 13657 [18
spm], Boa Viagem Beach, Niterói, Rio de Janeiro state, Brazil; MNRJ HSL5480 [1 sh], Boa Viagem Beach,
Niterói, Rio de Janeiro state, Brazil; MNRJ 2337 [2 spm], Forno Beach, Arraial do Cabo, Rio de Janeiro state,
Brazil; MNRJ HSL3366 [3 sh], Arraial do Cabo, Rio de Janeiro state, Brazil; MNRJ 9516 [1 sh], Praia Rasa,
Buzios, Rio de Janeiro state, Brazil; MNRJ 9517 [3 spm], Joana Beach, Cassimiro de Abreu, Rio de Janeiro state,
Brazil.
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Remarks. The holotype was erroneously published as MNRJ 7464, without an HSL acronym. The catalogue
numbers for paratypes MNRJ HSL3366 and MNRJ HSL5480 were originally provided without the HSL acronym.
The original publication lists five specimens in MNRJ 2336, five specimens in MNRJ 9517 and four shells in
MNRJ 3366, but does not provide numbers for the specimens in MNRJ 9518. Paratype MNRJ 13657 was
originally published as MNRJ 7464 (the same collection number as the holotype).
Dornellas & Simone (2011) stated that paratypes of this species deposited in the MZSP malacological
collection were removed from MNRJ 7464, but they were actually removed from MNRJ HSL7464.
carioca, Oenopota [Turridae]. Figueira & Absalão, 2010b: 476, fig. 1G, H, I. Type locality: Campos Basin,
southeast Brazil, southwestern Atlantic (22°27’31"S, 40°09’23"W, 750 m dep).
Paratypes. MNRJ 13556 [2 sh], Campos Basin, southeast Brazil, (22°04’34"S, 39°54’05"W, 750 m dep),
Brazil.
carloslirai, Anachis (Costoanachis) [Columbellidae]. Costa, 1996: 45, figs. 1, 3−5. Type locality: Off Guarapari,
Espírito Santo state, Brazil (20°41’S, 40°00’W, 20−25 m dep).
Paratype. MNRJ 7152 [1 sh], type locality.
carlucioi, Protoglyptus [Naesiotus, Bulimulidae]. Rezende & Lanzieri, 1963: 112, figs. 1−38. Type locality: Gruta
calcárea near Planaltina Nova, Goiás state, Brazil.
Paratypes. MNRJ HSL7780 [36 sh]; MNRJ HSL7915 [38 sh], type locality.
Remarks. The type material was originally deposited in the Instituto Oswaldo Cruz (IOC) in Rio de Janeiro
and later transferred to MNRJ; the original description mentioned that the holotype, 10 dissected paratypes and 39
shells were deposited in lot MNRJ HSL7780. However, the presumed holotype is not in a separate vial, but mixedup with the remaining paratypes. It is impossible to recognize the holotype based on the information provided by
the original desciption. The 10 dissected paratypes and three shells from MNRJ HSL7780, the lot MNRJ HSL7916
and 20 paratypes from MNRJ HSL7915 mentioned in the original publication were not located.
Simone (2006a: 125, fig. 396) figured a paratype from MNRJ HSL7915, but did not use the acronym HSL.
cearense, Opaliopsis [Nystiellidae]. Andrade, Costa & Pimenta, 2011: 1564, figs. 10−16. Type locality: Canopus
Bank, 96 miles off Ceará state, Brazil (02°14′25′′ S, 38022′50′′W, 240–260 m dep).
Holotype. MNRJ 15500 [sh]; paratypes: MNRJ 13347 [3 sh], type locality.
cepene, Cerithiella [Newtoniellidae]. Lima & Barros, 2007: 66, figs. 10−14. Type locality: Continental slope of
Pernambuco, Brazil (08°46.5’00"S, 34°44.5’00"W, 690 m dep).
Holotype. MNRJ 10837 [sh].
chiletensis, Scutalus (Scutalus) [Orthalicidae]. Weyrauch, 1967a: 373, figs. 24−30. Type locality: Left bank of Rio
Jequetepeque, mountain opposite Chilete (07°13’18"S, 078°50’40"W, 850–950 m alt, Dept. Cajamarca, Peru.
Paratypes. MNRJ 4416 [10 sh], type locality.
Remarks. The paratype MNRJ 4416 bears a label indicating “WW 1354/10 paratypes”; it was part of the
paratype lot from Weyrauch private collection originally published as “WW 1354/836”, and was later donated to
MNRJ.
clenchi, Conus [Conidae]. Martins, 1943: 2, figs (not numbered). Type locality: Barra do Furado, Campos, Rio de
Janeiro state, Brazil.
Holotype. MNRJ 11720 [sh] (Fig. 28).
clertoni, Psilaxis [Architectonicidae]. Tenório, Barros, Francisco & Silva, 2011: 181, fig. 3). Type locality: Off
Pernambuco state, Brazil (08°46.5'S, 34°44.5'W, 690 m dep).
Holotype. MNRJ 13031 [sh].
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cloveri, Marginella [Marginellidae]. Rios & Matthews, 1972: 31, fig. 1. Type locality: Off Cabo Cassiporé, 100 m,
Amapá state, Brazil.
Paratypes. MNRJ 3642 [1 sh], off Maranhão state collection station 1875 (00°56’S, 43°41’W, 45 m dep),
Brazil.
coelhoi, Bostryx [Bulimulidae]. Breure, 1978: 61, figs. 80−85. Type locality: Dept. Ancash, 7 km NE Sam Marcos
(2,720 m alt), Peru.
Paratypes. MNRJ 8481 [3 sh], type locality.
Remarks. The paratype MNRJ 8481 was originally published as UF 22799, from which it was later separated
and donated to MNRJ.
coltrorum, Spixia [Orthalicidae]. Simone, 2012a: 433, figs. 9–14. Type locality: Santa Maria da Vitória, Bahia
state, Brazil (~13°24’S, 44°12’W, ~460 m alt).
Paratype. MNRJ 30378 [1 sh], type locality.
Remarks. Original description did not mention the catalogue number for MNRJ paratypes.
conifera, Chrysallida [Pyramidellidae]. Pimenta, 2012: 441, figs. 13–18. Type locality: Continental shelf of
Campos Basin, Rio de Janeiro state, southeast of Brazil, HAB 16 sta C3 (22°46’49"S, 41°3’16"W, 78 m dep).
Holotype. MNRJ 16300 [sh]. Paratypes: MNRJ 16301 [1 sh], continental shelf of Bacia de Campos, Rio de
Janeiro state, southeast of Brazil, HAB 16 sta G3 (22°3’41.0"S, 40°10’5.38"W, 75 m); MNRJ 16302 [1 sh], type
locality; MNRJ 16303 [1 sh], continental shelf of Bacia de Campos, Rio de Janeiro state, southeast of Brazil, HAB
16 sta B3 (22°59’43"S, 41°21’13"W, 77 m).
compta, Gaza [Margaritidae]. Simone & Cunha, 2006: 4, figs. 1−10, 49−52, 67−77, 92. Type locality: Brazil, off
Cabo Frio, Rio de Janeiro state, 22°53'S, 42°01'W, 700−800 m dep.
Paratype. MNRJ 10531 [1 sh], type locality.
Remarks. MNRJ 10566 [1 sh], from the type locality, contains a label written by the authors indicating
“paratype”, but this lot was not mentioned in the original publication and therefore, according to ICZN (1999:
Article 72.4.1.1), it is a topotype.
corallina, Kora [Orthalicidae]. Simone, 2012a: 432, figs. 1–8. Type locality: Santa Maria da Vitória, Bahia state,
Brazil (~13°24’S, 44°12’W, ~460 m alt).
Paratype. MNRJ 30377 [2 sh], type locality.
Remarks. Original description did not mention the catalogue number for MNRJ paratypes.
coronata, Benthonellania [Rissoidae]. Absalão & Santos, 2004: 336, figs. 7−8. Type locality: REVIZEE station
A3, south of Bahia state, Brazil (18°30’00"S, 37°42’20"W, 260 m dep).
Paratype. MNRJ 9960 [1 sh], REVIZEE station 52C, Rio de Janeiro state (21°45’52"S, 40°04’52"W, 450 m
dep).
costifer, Bostryx (Elatibostryx) imeldae [Orthalicidae]. Weyrauch, 1960b: 129, pl. 12, figs. 35−38. Type locality: 5
Km from Laraos on footpath to Yauyos, left margin of Río Mayo [Río Laraos], [Cerro] Quichao] (12°20’45"S,
75°48’53"W, 3500m alt), Peru, Departamento de Lima.
Paratypes. MNRJ 4446 [3 sh], type locality.
Remarks. The paratype MNRJ 4446 bears a label indicating “WW 3319/3 paratypes”; it was part of the
paratype lot from Weyrauch private collection originally published as “WW 3319/53 & 32 juv.”, and was later
donated to the MNRJ collection. Although MNRJ 4446 was not cited by Barbosa et al. (2008), probably because
was not found by them in the collection by that time, it fills the the requirements here taken into account to be
considered a paratype lot.
crassa, Thala [Nodicostellaria, Costellariidae]. Simone, 1995a: 805−810, figs. 1−11. Type locality: Continental
slope off Ubatuba, São Paulo state, Brazil (24°25’0"S, 44°16’5"W, 320 m dep).
Paratypes. MNRJ 6934 [2 spm], type locality.
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cylindrata, Eulimella [Pyramidellidae]. Pimenta, Santos & Absalão, 2011: 30, figs. 26−31. Type locality: MD 55
station CB96: 21°31.8’S, 40°08.7’W (295–300 m dep), east of Cabo de São Tomé, off southeastern Brazil.
Paratype. MNRJ 19486 [1 sh], MD55 station CB77 (19°40’6"S, 37°48’1"W, 790–940 m dep), Abrolhos
continental slope, northeastern Brazil.
debilisculptus, Scutalus (Scutalus) coraeformis [Orthalicidae]: Weyrauch, 1967a: 376, pl. 2, figs. 20−23; pl. 9,
figs. 137, 138. Type locality: Rio Jequetepeque, 16 km N Tembladera on the road Pacasmayo-cajamarca, near
Quinden, 600–700 m alt, Dept. Cajamarca, Peru.
Paratype. MNRJ 4420 [1 sh], type locality.
Remarks. The original publication (Weyrauch, 1967a: 377) mentioned the holotype as WW 1964a and 116
paratypes as WW 1964; however, in the legends of the figures 20−23, it is indicated holotype WW 1984a and
paratypes WW 1984. Breure (2012: figs 49–50) figured the holotype (currently stored at FML 1984, Fundación
Miguel Lillo, Tucumán, Argentina) which has an ink handwritten mark in the shell indicating “WW 1984
holotype”. Thus we believe that the indication of the number 1964 in Weyrauch (1967a: 377) was a lapsus calami
and the correct collection number of this material in WW collection was 1984.
The paratype MNRJ 4420 bears a label indicating “WW 1984/2 paratypes”, but only one shell was found, the
second one is missing. This shell has an ink handwritten mark, very similar to the figure in Breure (2012)
indicating “1984”. For these reasons, we believe that MNRJ 4420, although not cited in Barbosa et al. (2008), was
part of the paratype lot from Weyrauch private collection originally published simultaneously as “WW1864” and
“WW1984”, and was latter donated to the MNRJ collection.
delucai, Conus [Conidae]. Coltro, 2004: 3, pl. 3, fig. a, pl. 10, figs. h1, p1–11. Type locality: 75 km E Abrolhos
Archipelago, off Alcobaça, Bahia state, Brazil (15°57’S, 38°01’W).
Paratypes. MNRJ 10190 [1 sh], type locality; MNRJ 10191 [1 sh], type locality.
devyanae, Latirus [Fasciolariidae] Rios, Costa & Calvo, 1994: 33, figs. 1−5. Type locality: Off Cabo Frio, Rio de
Janeiro state, Brazil (140 m dep).
Paratype. MNRJ 7060 [1 sh], type locality.
diabula, Oenopota [Turridae]. Figueira & Absalão, 2010b: 475, figs. 1D−F. Type locality: Campos Basin,
southeast Brazil, southwestern Atlantic, (22°10’43"S, 39°54’46"W, 750 m dep).
Paratypes. MNRJ 13554 [1 sh], Campos Basin, southeast Brazil (22°31’28"S, 40°03’50"W, 1,050 m dep);
MNRJ 13555 [2 sh], Campos Basin, southeast Brazil (22°26’28"S, 39°58’53"W, 1,046 m dep),
ejuncida, Eulimella [Pyramidellidae]. Pimenta, Santos & Absalão, 2011: 33, figs. 40−47. Type locality: Off São
Paulo state, PADCT station 6631 (25°46' S, 45°28.8' W, 164 m dep), southeast coast of Brazil.
Paratypes. MNRJ 15399, [2 sh], PADCT station 6553, (27°10.38' S, 47°27.54' W, 129 m dep), off Santa
Catarina state, south Brazil; MNRJ 15398, type locality, [1sh].
ektopa, Egila [Pyramidellidae]. Pimenta & Absalão, 2004b: 170, figs. 6E−H. Type locality: Off Espírito Santo
state, Brazil, REVIZEE, station VV38 (19°44’S, 38°22’W, 71.4 m dep).
Holotype. MNRJ 10255 [sh].
eliezeri, Caecum [Caecidae]. Absalão, 1997: 271, figs. 1−4. Type locality: Off Rio de Janeiro state, Brazil,
GEOMAR XII station 58 (21°31.7’S, 40°19’W, 41 m dep).
Paratype. MNRJ 7158 [1 sh], off Rio de Janeiro state, Brazil, GEOMAR XII station 111 (22°17.2’S,
40°49.6’00"W, 51.8 m dep).
elisae, Lucapina [Fissurellidae]. Costa & Simone, 2006: 1, figs. 1−18. Type locality: Canopus Bank, off Fortaleza,
Ceará state, Brazil (02°14’25"S, 38°22’50"W, 240−260 m dep).
Holotype. MNRJ 10661 [spm]; paratypes: MNRJ 10680 [2 spm], type locality; MNRJ 10681 [1 sh], type
locality; MNRJ 10683 [2 sh], type locality.
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elongata, Cochlespira [Turridae]. Simone, 1999a: 105, figs. 1−2, 5, 7, 9, 11−22. Type locality: Off Ubatuba, São
Paulo state, Brazil (24°31’0"S, 44°28’0"W, 250 m dep).
Paratypes. MNRJ 7161 [2 spm], type locality.
enceladus, Benthomangelia [Turridae]. Figueira & Absalão, 2010a: 735, figs. 1D, E, F. Type locality: Campos
Basin, southeast Brazil (22°02’51"S, 39°52’22"W, 1,048 m dep).
Paratype. MNRJ 13551 [1 sh], type locality.
equatorialis, Thylaeodus [Vermetidae]. Spotorno & Simone, 2013: 89, figs. 1−18. Type locality: Belmonte Inlet,
Enseada, São Pedro e São Paulo Archipelago, Pernambuco, Brazil (00°55’01"N, 29°20’44"W, 3−10 m dep).
Paratytpes: MNRJ 30504 [2 sh], type locality.
esamianus, Tomigerus [Bulimulidae]. Salgado & Coelho, 1990: 2, figs. 1−7. Type locality: Mossoró, Passagem do
Rio (rodovia RN76), Rio Grande do Norte state, Brazil.
Holotype. MNRJ 5980 [sh]; paratypes MNRJ 5981 [2 spms + fragments of a dissected spm], type locality;
MNRJ 5982 [53 sh], type locality; MNRJ 5983 [6 + 15 sh], type locality; MNRJ 5984 [63 sh], Governador DixSept Rosado, Rio Grande do Norte state.
Remarks. The original publication listed 26 specimens in MNRJ 5981; 8 specimens + 16 shells in MNRJ
5983; 40 shells in MNRJ 5982; and 65 shells in MNRJ 5984.
Holotype figured by Simone (2006a: 176−177, fig. 627).
exiguum, Eulimastoma [Pyramidellidae]. Pimenta, 2012: 444, figs. 19–24. Type locality: Continental shelf of
Campos Basin, Rio de Janeiro state, southeast of Brazil, HAB 17 sta I3 (21°23’33.709"S, 40°15’43.349"W, 88 m
dep).
Holotype. MNRJ 16308 [sh]; paratypes: MNRJ 16309, type locality [2 shells].
farinatiae, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2004a: 114, figs. 1−5. Type locality: Off Cabo Frio,
north coast of Rio de Janeiro state, CF VII station 6147 (22°53.7’S, 41°50.5’W, 50 m dep), southeast coast of
Brazil.
Holotype. MNRJ 8931 [sh]; paratype: MNRJ 8930 [1 sh], off Rio de Janeiro state, CF VII station 6165
(23°02.8’S, 042°46’W, 56 m dep), Brazil.
farroupilha, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2004a: 116, figs. 13−18. Type locality: Off Rio
Grande (42 m dep), Rio Grande do Sul state, south coast of Brazil.
Paratype. MNRJ 8919 [1 sh], type locality.
faustoi, Ancilla [Olividae]. Matthews, Matthews & Dijck, 1977: 115, fig. 1a−b. Type locality: Canopus station 99,
off Rio Grande do Norte state, Brazil (05°05’S, 34°57’W, 69 m dep).
Holotype. MNRJ 3790 [sh] (Fig. 23).
flemingi, Kerguelenella [Siphonariidae]. Powell, 1955: 121, pl. 5, fig. 44. Type locality: Bay south of Crozier
Point, Auckland Islands, New Zealand.
Paratypes. MNRJ HSL4069 [2 sh], type locality (Fig. 29).
Remarks. The original publication did not mention the number of paratypes, but provided some clues that
several specimens were used to describe the species. The catalogue by H. S. Lopes mentioned: “Powell leg XI.55 2
ex paratypes”. Therefore, we consider the shells at MNRJ HSL4060 as part of the type series, later exchanged
between Powell and H. S. Lopes.
flexispina, Arene [Areneidae]. Leal & Coelho, 1985: 45, figs. 1−3. Type locality: Brazil, Bahia state, 40 km off
Ponta do Corumbaú, (16°56’30"S, 38°39’30"W, 80 m dep).
Holotype. MNRJ 5352 [sh] (Fig. 4); paratypes: MNRJ 5353 [1 sh], type locality; MNRJ 5354 [1 sh],
GEOMAR X station 047, 120 km off Abrolhos Reef, Bahia state, Brazil (17°44’00"S, 37°21’00"W, 54 m dep).
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fluminensis, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2004a: 120, figs. 31−34. Type locality: Off Macaé,
Rio de Janeiro state, southeast coast of Brazil, (22°48’S, 41°09’W, 69 m dep).
Holotype. MNRJ 8943 [sh]; paratypes: MNRJ 8923 [1 sh], off north coast of Rio de Janeiro state, Brazil,
REVIZEE station D3 (22°52’S, 041°09’W, 80 m dep); MNRJ 8924 [1 sh], Bacia de Campos, Rio de Janeiro state,
Brazil, (22°15'20"S, 040°19'54"W, 100 m dep).
formicacorsii, Olivella (Orbignytesta) [Olivellidae]. Klappenbach, 1962: 95, figs. 1−6. Type locality: Playa La
Coronilla, Departamento de Rocha, Uruguay.
Paratypes. MNRJ 2909 [3 sh], type locality (Fig. 24).
Remarks. The paratype MNRJ 2909 was originally published under the collection number MNHN 0722 and
was later donated to the MNRJ collection.
franklini, Eulimastoma [Pyramidellidae]. Pimenta, 2012: 444, figs. 25–30. Type locality: Ponta Grande, Ilha
Grande Bay, South coast of Rio de Janeiro state, southeast of Brazil; Ilha Grande sta 25 (23°5.098’S, 44°18.603’W,
17 m).
Holotype. MNRJ 17168 [sh]; paratypes: MNRJ 17167 [2 sh], type locality; MNRJ 17169 [4 sh], Enseada de
Dois Rios, Ilha Grande, Rio de Janeiro state, Brazil.
frenguellii, Lathyrus [Fusinus, Fasciolariidae]. Carcelles, 1953: 8, figs. 13−16, 19. Type locality: a 30/35 millas as
S.S.E. de Puerto Quequén, Argentina.
Paratype. MNRJ 13204 [1 sh], type locality (Fig. 18).
Remarks. The paratype MNRJ 13204 was originally published under MACN 24442; in 1971, it was
exchanged for a paratype of Bursa pacamoni (see also Tablado & Mantinian 2004). Misspelled Lathyrus, error for
Latirus.
gigliottii, Metula [Buccinidae]. Coltro, 2005: 2, Pl. C, figs. 1−11. Type locality: Off Cabo de Santa Marta, Santa
Catarina state, Brazil.
Paratypes. MNRJ 10976 [1 sh], type locality; MNRJ 10977 [1 sh], type locality.
Remarks. Original publication did not provide MNRJ collection number; MNRJ 10977 published as paratype
3, P. M. Santos Costa collection, was later donated to the MNRJ collection.
gili, Pazinotus [Muricidae]. Costa & Pimenta, 2012: 121, figs. 5–6. Type locality: south slope of the Canopus
Bank, off Ceará state (02°14′25″S, 038°22′ 50″W, 240–260 m dep).
Holotype. MNRJ 11037 [spm]; paratypes: MNRJ 11063, type locality [5 spm]; MNRJ 11008, type locality [1
spm dissected].
glomeratus, Bostryx (Bostryx) zilchi [Orthalicidae]. Weyrauch, 1960b: 124, taf. 12, figs. 29−34. Type locality: 1
km from Laraos on footpath to Yauyos, (Cerro) Quichao (12°20’45"S, 75°48’53"W, 3500 m alt), Peru,
Departamento Lima.
Paratypes. MNRJ 4445 [5 sh], type locality.
Remarks. The paratype MNRJ 4445 bears a label indicating “WW 3320/5 paratypes”; it was part of the
paratype lot from Weyrauch private collection originally published as “WW 3320/zahlreich”, and was later donated
to the MNRJ collection.
goncalvesi, Engina [Anna, Buccinidae]. Coltro, 2005: 1, Pl. B, figs. 1−11. Type locality: Off Cabo Frio, Rio de
Janeiro state, Brazil.
Paratypes. MNRJ 10978 [1 sh] type locality; MNRJ 10979 [1sh], type locality; MNRJ 15187 [1 sh], type
locality.
Remarks. The original publication did not list the MNRJ collection numbers. MNRJ 10979, published as
paratype 4, P. M. Santos Costa collection, was later donated to the MNRJ collection.
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gongyloskymnus, Metaxia [Triphoridae]. Fernandes & Pimenta, 2011: 826, figs. 25–31. Type locality: Bacia de
Campos, off Rio de Janeiro state, southeast Brazil (22°43´S, 40°40´W, 120 m).
Paratype. MNRJ 30293 [1 sh], type locality.
goytacazi, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2004a: 116, figs. 6−12. Type locality: Bacia de
Campos, Rio de Janeiro state, (22°15’25"S, 40°19’39"W, 100 m dep), southeast coast of Brazil.
Holotype. MNRJ 8935 [sh]; paratype: MNRJ 8936 [1 sh], Bacia de Campos, Rio de Janeiro state (22°15'07"S,
040°19'56"W, 95 m dep), Brazil.
grandiportus, Zilchiella [Clausiliidae]. Weyrauch, 1957: 10, Taf. 1, figs. 5−10. Type locality: Río Llaucan, Peña
Rota (? 06°37’22"S, 78°28’38"W, 2550−2750m alt), Peru, Departamento Cajamarca.
Paratypes. MNRJ 4461 [10 sh], type locality.
Remarks. The paratype MNRJ 4461 bears a label indicating “WW 2005/10 paratypes”; it was part of the
paratype lot from Weyrauch private collection originally published as “WW 2005”, and was later donated to the
MNRJ collection. Barbosa et al. (2008) erroniously mentioned this lot as MNRJ 4460 instead of 4461.
grandiventris, Scutalus [Orthalicidae]. Weyrauch, 1960a: 42, taf. 5, figs. 27−33. Type locality: between Trujillo
and Cajamarca, ca. 100 km NE Trujillo, above Cascas (07°29’S, 78°49’W, 1400, Peru, Departamento Cajamarca.
Paratypes. MNRJ 4417 [2 sh], type locality.
Remarks. The paratype MNRJ 4417 bears a label indicating “WW 1364/2 paratypes”; it was part of the
paratype lot from Weyrauch private collection originally published as “WW 1364/zahlreich”, and was later donated
to the MNRJ collection.
guerrinii, Bullata [Marginellidae]. de Souza & Coovert, 2001: 2, figs. 1, 7, 10. Type locality: Pedra da Lixa, Parcel
das Paredes, 27 km E off Caravelas, Bahia state, Brazil (17°41,5’S, 38°59’W, 5−15 m dep).
Holotype. MNRJ 7818 [sh].
harasewychi, Brocchinia [Cancellariidae]. Barros & Lima, 2007: 61, figs. 6−11. Type locality: along the state of
Pernambuco, Brazil, station D−11, 08°46,5’00"S, 34°44,5’00"W, 690m.
Paratype. MNRJ 10574 [1 sh], type locality.
helenae, Anachis [Cosmioconcha, Columbellidae]. Costa, 1983: 95, figs. 1−3. Type locality: Off Ilha da Boa
Viagem, Niterói, Rio de Janeiro state (22°54’43"S, 43°07’32"W, 8 m dep), Brazil.
Paratype. MNRJ 4864 [1 sh], off Amapá, 56m (Fig. 21).
Remarks. Published under the collection number MORG 14810H; later donated to the MNRJ Mollusca
Collection. The locality of each paratype was not given in the original description, but the label indicates “off
Amapá, 56 m”.
henckesi, Conus [Conidae]. Coltro, 2004: 3, pl. 2, fig. A, pl. 9, figs. H1, P1−P11. Type locality: Mar Grande,
Itaparica Island, Bahia state, Brazil.
Paratypes. MNRJ 10188 [1 sh], type locality; MNRJ 10189 [1 sh], type locality.
hubrechti, Diptychophlia [Turridae]. Cunha, 2005: 12, figs. 1−6. Type locality: Canopus Bank, off Fortaleza,
Ceará state, Brazil (02°14'25"S, 38°22'50"W, 260 m dep).
Paratypes. MNRJ 10530 [4 sh], type locality.
hyphala, Olivella (Olivina) [Olivellidae]. Absalão & Pimenta, 2003: 183, figs. 2A−C. Type locality: Abrolhos
continental slope, Brazil, MD 55 stn DC75, (18°59’S, 37°50’W, 295 m dep).
Paratypes. MNRJ 8914 [2 sh], type locality; MNRJ 8917 [2 sh], type locality.
iansa, Conus [Conidae]. Petuch, 1979: 524, figs. 4G−H. Type locality: 2 km east of Santa Barbara Island,
Abrolhos Archipelago, Bahia state, Brazil (17°57’S, 38°41’W, 25 m dep).
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Paratype. MNRJ 4155 [1 sh], type locality.
Remarks. The paratype MNRJ 4155 was not listed in the original publication. According to the information
on the label of this lot and the length of the shell, this specimen corresponds to the paratype listed in the original
publication as deposited in the author’s collection, which was later transferred to MNRJ.
iemanja, Taringa [Discodorididae]. Alvim & Pimenta, 2013: 192, figs. 2A−B, 19−20. Type locality: Continental
slope of Campos Basin, Rio de Janeiro state, Brazil (22°06’9875"S, 40°10’8204"W, 150 m dep).
Holotype. MNRJ 13180 [spm, dissected].
iheringi, Peltella [Orthalicidae]. Leme, 1968: 197, figs. 1−13. Type locality: Floresta da Tijuca, Rio de Janeiro
state, Brazil.
Paratype. MNRJ 3513 [1 spm], type locality.
Remarks. The paratype MNRJ 3513 was originally publicated as MZSP 17987, and was subsequently donated
to MNRJ.
indigestus, Adelomelon [Volutidae]. Ihering, 1908: 433, fig. 2. Type locality: São Sebastião Island, São Paulo state
and Guaratiba, Rio de Janeiro state, Brazil.
Syntype. MNRJ 950 [sh], Guaratiba, Rio de Janeiro state, Brazil.
Remarks. According to Wiggers & Veitenheimer-Mendes (2005), who studied the syntypes of this species,
there is no indication for a lectotype designation. The syntype at MNRJ 950 was illustrated in the original
publication, and also by Wiggers & Veitenheimer-Mendes (2005: 230, fig. 1e−g).
indiscreta, Rissoina [Rissoidae]. Leal & Moore, 1989: 140, figs. 2−7. Type locality: Ponta Verde, Maceió, Alagoas
state, Brazil (09°38’S, 35°44’W).
Holotype. MNRJ 5760 [sh]; paratypes: MNRJ HSL6025 [4 sh], type locality.
Remarks. The original publication listed MNRJ 5761, with three shells, from type locality, but this lot was not
located.
inflexus, Australorbis [Biomphalaria, Planorbidae]. Paraense & Deslandes, 1956: 149, figs. 1−7. Type locality:
Pouso Alegre, Minas Gerais state, Brazil.
Syntypes. MNRJ HSL7555 [27 sh], Pouso Alegre; MNRJ HSL7556 [79 sh], Barbacena; MNRJ HSL7557 [5
sh], Cachoeira; MNRJ HSL7558 [5 sh], Ouro Preto; MNRJ HSL7559 [14 sh], Retiro; MNRJ HSL7560 [40 sh],
Santa Rita do Sapucaí; all types from Minas Gerais state, Brazil.
Remarks. The original publication stated that the types were deposited in the “collection of the Instituto
Oswaldo Cruz (Rio de Janeiro)”; the lots were subsequently donated to the MNRJ collection. The original
publication listed five shells under number 7559, but 14 shells were found; the five dissected genitalia mentioned
as no. 7561 were not located. The original publication presented a list of both syntypes and paratypes. The
summary of the article stated that the type locality was “Pouso Alegre, state of Minas Gerais, Brazil”. This is the
locality of the syntypes in MNRJ HSL7555. On the original label of the type lots, the term “cotipos” (cotypes) is
used to refer to the lots listed as paratypes. All lots of “paratypes” or “cotypes” are herein interpreted as syntypes.
Simone (2006a: 108, fig. 318B) figured a syntype referred to as MNRJ 7560 (without using the HSL acronym).
insularis, Gonyostomus [Strophocheilidae]. Leme, 1974: 2, figs. 1, 7−16, Pl. 1, figs. 2, 4. Type locality: Búzios
Island, São Paulo state, Brazil (23°48’S, 45°08’W).
Paratype. MNRJ 3643 [1 sh], type locality.
Remarks. The original publication used the acronym MNCM to refer to the MNRJ Mollusca Collection.
insularis, Sinezona [Scissurellidae]. Simone, 2009a: 12, figs. 2−9. Type locality: São Pedro e São Paulo
Archipelago (00°55’01"N, 29°20’44"W), Brazil.
Paratypes. MNRJ 12868 [2 sh], type locality.
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intermedius, Australorbis [Biomphalaria, Planorbidae]. Paraense & Deslandes, 1962: 343, figs. 1−6. Type
locality: Valparaiso, São Paulo state, Brazil (21°13’S, 50°51’W).
Syntypes: MNRJ HSL7918 [1 dissected spm and 10 sh], type locality.
Remarks. The original publication did not mention types, but stated that 10 shells and five dissected
specimens were deposited at the IOC (No. 7918); these were later transferred to the MNRJ, and herein are
considered as syntypes. On page 349 of the original publication, it is noted that the type locality is Valparaíso, but
the latitudinal coordinate is erroneously given as “N” (North), instead of South.
Simone (2006a: 107, fig. 314A) figured nine syntypes, referred to as MNRJ 7918 (without using the acronym
HSL).
intratesta, Crepidula [Calyptraeidae]. Simone, 2006c: 21, figs. 13−15, 112. Type locality: Off Ubatuba, São Paulo
state, Brazil (23°29’S, 44°52’W, 38 m dep).
Paratypes. MNRJ 2338 [1 spm, mummified], Ilha do Governador, Rio de Janeiro, Rio de Janeiro state, Brazil;
MNRJ 8989 [5 sh], off Vitória, Espírito Santo state, Brazil.
itanhura, Episcynia [Tornidae]. Simone, 2012b: 81, figs. 16–27. Type locality: São Mateus, Guriri, Espírito Santo
state, Brazil (18°47’S, 32°39’W, 3 m dep).
Paratypes. MNRJ 31326 [3 sh], type locality.
Remarks. Original description did not mention the catalogue number for MNRJ paratypes.
janowskyi, Engina [Buccinidae]. Coltro, 2005: 1, Pl. A, figs. 1−5. Type locality: Off Guarapari, Espirito Santo
state, Brazil.
Paratypes. MNRJ 10980 [1 sh], type locality; MNRJ 15186 [1 sh], type locality.
Remarks. The original publication did not provide the MNRJ collection number. MNRJ 15186, published as
“paratype 3, P. M. Santos Costa collection”, was later donated to the MNRJ collection.
jonasi, Microcancilla [Cancellariidae]. Barros & Petit, 2007: 96, figs. 1−8. Type locality: Continental Slope from
the state of Pernambuco, northeastern Brazil (08°42.1’S, 34°44.1’W, 425 m dep).
Paratypes. MNRJ 10839 [1 sh], type locality; MNRJ 10838 [2 sh], Pernambuco, Brazil, (08°46.5’S,
34°44.5’W, 690 m dep).
jonasi, Pseudotorinia [Architectonicidae]. Tenório, Barros, Francisco & Silva, 2011: 183, fig. 4). Type locality:
Off Pernambuco state, Brazil (08°45.1'S, 35°44.9'W, 500 m dep).
Holotype. MNRJ 13032 [sh].
juani, Natica [Naticidae]. Costa & Pastorino, 2012: 30, figs. 21–28, 30, 37–40. Type locality: Southern littoral of
São Paulo state, Brazil (~17–35 m dep).
Holotype. MNRJ 17876 [sh]; paratypes: MNRJ 15029, off Búzios, Rio de Janeiro state, Brazil (22°42’39.4"S,
40°40’50.2" W) [3 sh]; MNRJ 15030, off Búzios, Rio de Janeiro state, Brazil (22°41’49" S, 40°40’29.8" W) [3 sh];
MNRJ 15028, off Ilha de Cabo Frio, Arraial do Cabo, Rio de Janeiro state, Brazil (23°18’00" S, 40°00’00" W) [2
spm]; MNRJ 17875, between Rasa Is. and Ilha Grande, Rio de Janeiro state, Brazil [2 sh]; MNRJ 17877, type
locality [1 sh].
juliae, Hypselodoris [Felimare, Chromodorididae]. Dacosta, Padula & Schrödl, 2010: 16, figs. 1A−C, 2−4. Type
locality: Praia das Conchas, Cabo Frio, Rio de Janeiro state, Brazil (22°52’15"S, 41°58’52"W, intertidal).
Holotype. MNRJ 10940 [spm, dissected].
kaapor, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2004a: 129, figs. 85−89. Type locality: Off São Paulo
state, PADCT station 6631 (25°46’S, 45°28.8’W, 164 m dep), southeast coast of Brazil.
Paratype. MNRJ 8932 [1 sh], type locality.
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kaicherae, Vexillum (Costellaria) [Nodicostellaria, Costellariidae]. Petuch, 1979: 523, figs. 4C−D. Type locality:
South end of Guabatibas Reefs, Abrolhos Reef Complex, Bahia state, Brazil (17°25’S, 39°08’W, 1 m dep).
Paratypes. MNRJ 3735 [2 sh], type locality.
Remarks. The original publication referred to MNRJ as “Museu Nacional do Brasil”.
kamayura, Turbonilla [Pyramidellidae]. Absalão, Santos & Tenório, 2003: 5, figs. 8−10. Type locality: Off
Paraiba state, northeast coast of Brazil, REVIZEE station 7 (06°57’32"S, 34°29’22"W, 750 m dep).
Paratype. MNRJ 9824 [1 sh], Ilha de Itaparica, Praia do Despacho, Bahia state, Brazil (13°01’S, 38°27’W).
Remarks. The original publication indicated that the lot MNRJ 9824 contained two shells, but only one shell
was located.
kanela, Aclis [Aclididae]. Absalão, 2009: 138, figs. 2I−K. Type locality: Campos Basin, BC Sul I #75
(22°31´28"S, 40°03´50"W, 1050 m dep).
Types not located.
Remarks. The original publication listed MNRJ 12852 as a paratype lot from Campos Basin, Brazil, BC Sul I
#80 (22°24´31"S, 39°57´28"W), 1050 m dep, without providing the number of specimens. The specimens were not
sent to the MNRJ Mollusca Collection.
karsticus, Pothamolithus [Hydrobiidae]. Simone & Moracchioli, 1994: 449, figs. 2a−c, 4b, 5b, 6b, 7b, 8b, 9b, 10b,
11b. Type locality: Stream inside Calcário Branco Cave, Apiaí, São Paulo state, Brazil (24°30’20"S, 48°44’25"W).
Paratypes. MNRJ 6936 [2 spm], type locality.
klappenbachi, Olivella (Olivina) [Olivellidae]. Absalão & Pimenta, 2003: 184, figs. 2D−E. Type locality: South of
Cabo Frio, Rio de Janeiro state, Brazil, MD 55 station CB105, (23°47’S, 42°10’W, 610 m dep).
Paratypes. MNRJ 8916 [2 sh], type locality.
klappenbachi, Strophocheilus (Megalobulimus) [Megalobulimus, Strophocheilidae]. Leme, 1964: 98, figs. 5−8.
Type locality: Iguape, São Paulo state, Brazil.
Paratype. MNRJ 3419 [1 sh], Gruta do Morro Preto, Iporanga, São Paulo state.
Remarks. Published under collection number MZSP 14.608 and subsequently donated to the MNRJ
collection.
lacteum, Latirus [Polygona, Fasciolariidae]. Matthews-Cascon, Matthews & Rocha, 1991: 2, figs. 1−4. Type
locality: Off Pará state, Brazil, (1°32’5"N, 47°24’5"W, 63 m dep).
Holotype. MNRJ 6201 [sh] (Fig. 19).
laurae, Notocochlis [Naticidae]. Costa & Pastorino, 2012: 25, figs. 1–8, 15, 17–20. Type locality: Canopus Bank,
off Fortaleza, Ceará state, Brazil (02°14’25"S, 38°22’50" W, 240–260 m dep).
Holotype. MNRJ 11866 [spm]; paratypes: MNRJ 11867 [12 sh + 3 opercula], type locality.
Remarks. One paratype is missing in MNRJ 11867.
ledaluciae, Pyrene (Conella) [Conella, Columbellidae]. Rios & Tostes, 1981: 178, figs. 1−3. Type locality: Off
Maricá, Rio de Janeiro state, 80m, Brazil.
Paratypes. MNRJ 3900 [1 sh] (Fig. 22), type locality; MNRJ 3901 [1 sh], type locality.
Remarks. The paratype MNRJ 3900 has a different shell length than the published value. The paratype MNRJ
3901 was not listed in the original publication.
lepta, Peristichia [Pyramidellidae]. Pimenta, Santos & Absalão, 2008: 182, fig. 5D−H. Type locality: Off Paraná
state, Brazil; REVIZEE station 6656 (25°22.1’S, 46°47.5’W, 70 m dep).
Paratypes. MNRJ 10693 [2 sh], Ilha Grande station 36 (Ponta Alta de Parnaioca, 23°12.25’S, 44°30.35’W, 35
m dep), Rio de Janeiro state, Brazil.
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lepta, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2004a: 118, figs. 23−26. Type locality: Off Cabo Frio,
north coast of Rio de Janeiro state (22°53.7’S, 41°50.5’W, 50 m dep), southeast coast of Brazil.
Holotype. MNRJ 8939 [sh]; paratype: MNRJ 8940 [1 sh], Bacia de Campos, Rio de Janeiro state (22°15'36"S,
040°20'16"S, 100 m dep), Brazil.
litosbathron, Triptychus [Pyramidellidae]. Pimenta, Santos & Absalão, 2008: 180, fig. 4E−I. Type locality:
25°15.76’S, 45°04.62’W, 124 m; off Paraná state, Brazil.
Paratypes. MNRJ 10941 [5 sh], type locality; MNRJ 10694 [1 sh], off Espírito Santo state, Brazil, REVIZEE
station D1 (20°48’S, 41°09’33"W, 69 m dep).
lopesi, Mitra [Subcancilla, Mitridae]. Matthews & Coelho, 1969: 3, figs. 1−3. Type locality: Off Mucuripe,
Fortaleza, Ceará state, Brazil.
Holotype. MNRJ 3490 [sh]; paratypes: MNRJ 3491 [1 sh], Cabo Orange, Amapá state (04°28’N, 50°,16’W,
100 m dep), Brazil; MNRJ 3492 [1 sh], off Belmonte, Bahia state, Brazil (15°53’S, 38°22W, 48 m dep); MNRJ
HSL8060 [1 spm], type locality.
lopesi, Protoglyptus [Naesiotus, Bulimulidae]. Resende, Lanzieri & Inada, 1972: 374, figs. 1−47. Type locality:
Calcareous region of Fercal, Brasília (Federal District), Brazil.
Holotype. MNRJ 3555 [spm]; paratypes: MNRJ 3556 [8 sh + spms without sh], type locality; MNRJ 3557
[fragments of mummified soft parts and dissected spm], type locality; MNRJ 3558 [22 shells + 4 spms without sh +
fragments of a dissected spm, mummified], type locality; MNRJ 3559 [8 sh], type locality; MNRJ 3560 [7 sh], type
locality; MNRJ 30822, type locality [1 spm].
Remarks. The original publication referred to MNRJ as Col. Mol. M. N.; paratype MNRJ 3561 was not
located.
The original publication listed MNRJ 3555 as the holotype, but two lots (each with a complete specimen) and
labeled as MNRJ 3555 were found; one of them is labeled “holotype” and the second shell, thus a paratype, is here
separated as MNRJ 30822. Both shells were originaly illustrated (Resende et al., 1972: figs. 1, 2), but without
indication of their type status; the shell in figure 1 corresponds to the holotype.
Holotype figured by Simone (2006a: 126, fig. 399).
lopesi, Succinea [Succineidae]. Lanzieri, 1966: 171, figs. 2−4, 8−66. Type locality: Pico do Desejado, Ilha da
Trindade, off Espírito Santo state, Brazil.
Paratypes. MNRJ 3176 [22 spm + 2 dissected spms + 82 mummified spms + 32 sh + fragments of one
dissected spm studied in microanatomy], type locality (Fig. 30); MNRJ 3175 [5 spms + 2 sh + 2 fragments of
dissected spms studied for microanatomy], type locality; MNRJ 3174 [57 mummified spm], type locality.
Remarks. The holotype (published as MNRJ 3176) was not located; it is probably mixed-up with the
paratypes. It is not possible to identify the holotype based on information provided in the original publication or the
catalogue of the collection. The figured shell in Simone (2006a: 236, fig. 904) referred to as holotype MNRJ 3175,
is actually one of the paratypes and cannot be identified amongst the specimens in the lot. Original publication
states that MNRJ 3175 with 5 exemplars + 12 dissected + 2 used for microanatomic study + 7 shells.
lundi, Thaumastus [Bulimulidae]. Pena, Salgado & Coelho, 2005: 21, figs. 1−15. Type locality: Lapinha Cave,
Lagoa Santa, Minas Gerais state, Brazil.
Holotype. MNRJ 8092 [sh]; paratypes: MNRJ 8093 [1 sh], type locality; MNRJ 8094 [1 sh], type locality;
MNRJ 8095 [24 sh + dissected soft parts + 16 histological laminae], type locality; MNRJ 8096 [1 spm], type
locality; MNRJ 9807 [10 sh + mounted stub with protoconch], type locality; MNRJ 9808 [1 spm], type locality;
MNRJ 9809 [1 sh + 3 spm + soft parts dissected + mounted stub with radula + 23 histological laminae], type
locality; MNRJ 4179 [1 sh], Pedro Leopoldo Municipality, “Grande Abrigo da Lapa Vermelha”, Minas Gerais
state, Brazil; MNRJ 9983 [1 fragmented sh], Lapa Vermelha, Minas Gerais state, Brazil.
Remarks. Three shells from MNRJ 8095 and one shell from MNRJ 9809 were not located.
Simone (2006a) did not mention this species in the catalogue of land and freshwater molluscs from Brazil.
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macaensis, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2001: 83, 85, figs. 34−41. Type locality:
Arquipélago de Santana, Macaé, Rio de Janeiro state, Brazil (22°24’30"S, 41°42’30"W).
Holotype. MNRJ 8434 [sh]; paratype: MNRJ 8435 [1 sh], Rio de Janeiro state, Brazil CFVII station 6147
(22°53.7’S, 41°50.5’30"W, 50 m dep).
madrediosensis, Stephadiscus [Charopidae]. Miquel & Ramírez, 2011: 51, figs 2−6. Type locality: Manu province,
Los Amigos Conservation Concession (LACC), Los Amigos Biological station (CICRA), (12°34’07"S,
70°05’57"W, 268m alt), Madre de Dios Department, Peru.
Paratypes. MNRJ 15467 [2 sh], Inkaterra Reserva Amazónica, Canopy station: Tower 1 near Platform 1
(12°31'53.71"S, 69°02'47.76"W), Lower Madre de Dios River, Tambopata province, Madre de Dios Department,
Peru.
maese, Fusiturricula [Turridae]. Rios, 1985: 133, Pl. 46, fig. 604. Type locality: “off Rio de Janeiro to São Paulo
state”, Brazil.
Paratype. MNRJ 4545 [1 sh], off Juatinga, Rio de Janeiro state, Brazil.
Remarks. The original publication listed only the holotype, deposited in the MORG. However, the description
provided a range of shell sizes (35−50 mm length), indicating that other shells were present for the description of
this taxon. The label of MNRJ 4545 was written by the author, who donated the shell to the MNRJ, as a paratype.
We interpret this evidence in accordance with ICZN (1999: article 72.4.1.1).
maestratii, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2004a: 128, figs. 79−84. Type locality: Praia do
Despacho, Itaparica, Bahia state, northeast coast of Brazil.
Holotype. MNRJ 8934 [sh]; paratypes: MNRJ 8933 [2 sh], type locality; MNRJ 8927 [2 sh], type locality.
marcusi, Acmaea [Lottia, Lottiidae]. Righi, 1966: 269, figs. 5−6, 9, 14, 16, 23. Type locality: Tartarugas Beach,
Trindade Island, Espírito Santo state, Brazil.
Holotype. MNRJ 3200 [spm] (Fig. 1).
Remarks. The original publication figured an empty shell of Acmaea marcusi in dorsal and ventral views
(Righi, 1966: 269, figs. 5−6), without mentioning its type status. It is not the holotype, since the original
publication indicates that the holotype is “a complete limpet in the collection of the Museum Nacional, Rio de
Janeiro, No. 3,200”, i.e. MNRJ 3200 (Fig. 1).
massambabensis, Caecum [Caecidae]. Absalão, 1994: 138, figs. 1A−C. Type locality: Off Massambaba Beach,
Rio de Janeiro state, Brazil (22°57’30"S, 41°19’30"W, 31 m dep).
Paratypes. MNRJ 6882 [2 sh], type locality; MNRJ 6883 [1 sh], type locality; MNRJ 7160 [1 sh], type
locality.
Remarks. Paratype MNRJ 7160 was not cited in the original publication, but contains a label by the author
indicating “paratype 15”. MNRJ 6882, cited in the original publication as “paratype 8”, contains two shells, both
labeled by the author as “paratype 8”.
matthewsi, Tomigerus (Tomigerus) [Bulimulidae]. Salgado & Leme, 1991: 1, figs. 1−4. Type locality: Martins,
Serra do Martins, 4 km from the city, Rio Grande do Norte state, Brazil.
Holotype. MNRJ 6182 [sh]; paratypes: MNRJ 6183 [1 dissected spm + 1 reproductive system dissected,
mummified], type locality; MNRJ 6184 [4 spms without sh], Serra do Mossoró, Mossoró, Rio Grande do Norte
state; MNRJ 6185 [6 spms without sh + 4 spm], Jaguaruana, Serra Dantas, Ceará state; MNRJ 6187 [16 spms
without sh + 32 eggs + reproductive system dissected], type locality; MNRJ 6188 [3 sh], oasis on caatinga in
Umarizal, south of Rio Grande do Norte state.
Remarks. The original publication mentioned several specimens in the paratype lots MNRJ 6183, MNRJ
6184, MNRJ 6185, but only those listed above were found in the MNRJ Malacological Collection. Holotype
figured by Simone (2006a: 176−177, fig. 629).
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mauricioi, Conus [Conidae]. Coltro, 2004: 6, pl. 6, fig. A, pl. 14, figs. H1, P1−P11, pl. 16, fig. E.Type locality: Off
Rio do Fogo, Rio Grande do Norte state, Brazil.
Paratypes. MNRJ 10220 [1 sh], type locality; MNRJ 10221 [1 sh], type locality.
mauryi, Epitonium [Epitoniidae]. Tursch & Pierret, 1964: 36, fig. 5. Type locality: Off Punta de Juatinga, Brazil
(23°22’S, 48°28’W, 50 m dep).
Paratype. MNRJ 3117 [1 sh], type locality (Fig. 14).
Remarks. The original publication stated that paratypes I and III were deposited in the MNRJ collection, but
only paratype III was found.
megaumbilicata, Brookula [Seguenzioidea]. Absalão & Pimenta, 2005: 194, figs. 11−16. Type locality: Bacia de
Campos, station 2, (21°57'S, 39°50'03"W, 1200 m dep), Rio de Janeiro state, Brazil.
Holotype. MNRJ 9626 [sh]; paratypes: MNRJ 9627 [2 sh], Bacia de Campos station 41 (22°39'28.1"S,
40°08'27"W, 1222 m dep); MNRJ 9844 [7 sh], Bacia de Campos station 41 (22°39'28.1"S, 40°08'27"W, 1222 m
dep).
mermeroglaphyra, Turbonilla [Pyramidellidae]. Absalão, Santos & Tenório, 2003: 7, figs. 11−14. Type locality:
Off Alagoas state, northeast coast of Brazil, REVIZEE station 2 (10°38’01"S, 36°11’05"W, 255 m dep).
Holotype. MNRJ 9825 [sh].
mesoatlantica, Fissurella [Fissurellidae]. Simone, 2008: 293, figs. 1−40. Type locality: Belmonte Island, São
Pedro e São Paulo Archipelago (00°51’01"N, 29°20’44"W), Brazil.
Paratypes. MNRJ 12476 [3 spm], type locality.
meta, Ampullaria [Pomacea, Ampullariidae]. Ihering, 1915: 12, figs. 6−7. Type locality: Rio São Francisco,
Cidade da Barra, Bahia state, Brazil.
Holotype. MNRJ 12861 [sh] (Fig. 6).
Remarks. The original publication did not mention type material. The shell in MNRJ 12861 contains a label
listing it as a type, indicating the same locality and collector stated by Ihering (1915), and is identical to the shell
illustrated. Simone (2006a: 55, fig. 93) referred to this shell as “probable type” and used the collection number
MNRJ 34350, which corresponds to the old numbering series of the MNRJ Molluca Collection (see introduction).
metamorphosicum, Caecum [Caecidae]. Lima, Santos & Absalão, 2013: 769, figs. 3−6. Type locality: Off the
coast of the Rio de Janeiro state, southeastern Brazil, REVIZEE Central VI sta R1 (21°38’S, 40°10’W, 140 m dep).
Paratypes. MNRJ 31247 [1 sh], continental shelf of Campos Basin, Rio de Janeiro (23°04’S, 40°59’W, about
100 m dep); MNRJ 31248 [4 sh], continental shelf of Campos Basin, Rio de Janeiro (22°07’43.55"S,
40°18’47.00"W, 72 m dep); MNRJ 31249 [10 sh], continental shelf of Campos Basin, Rio de Janeiro
(22°03’39.04"S, 40°06’59.81"W,90 m dep); MNRJ 31250 [4 sh], continental shelf of Campos Basin, Rio de
Janeiro ( 22°17’42.21"S, 40°27’00.05"W, 103 m dep); MNRJ 31251 [7 sh], continental shelf of Campos Basin, Rio
de Janeiro (22°52’02.05"S, 40°57’29.00"W, 91 m dep).
midas, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2004a: 126, figs. 66−71. Type locality: Off Santos (45 m
dep), São Paulo state, southeast coast of Brazil.
Paratype. MNRJ 8925 [1 sh], type locality.
minor, Bostryx (Scansicohlea) haasi [Orthalicidae]. Weyrauch, 1960a: 35, taf. 5, fig. 34. Type locality: road Limala Oroya, 90 km NE Lima, right bank Río Rimac, above San mateo (11°45’34"S, 76°18’02"W, 3300m alt), Peru,
Deparatamento Lima.
Paratype. MNRJ 4452 [5 sh], Central Peru, on the right side of Rimac River (3300 m alt).
Remarks. The paratype MNRJ 4452 bears a label indicating “WW 2035/5 paratypes”; it was part of the
paratype lot from Weyrauch private collection originally published as “WW 2035/146”, and was subsequently
donated to the MNRJ collection.
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mirim, Acteon [Acteonidae]. Cunha, 2011: 230, figs. 2−8. Type locality: Off Fortaleza, Canopus Bank (2°14´25"S,
38°22´50"W, 240 m dep), Ceará state, Brazil.
Paratype. MNRJ 10529 [1 sh], type locality.
Remarks. The original publication mentioned three shells in the lot MNRJ 10529, but only one shell was sent
to the MNRJ collection.
moisei, Mitra [Vexilum, Costellariidae]. McGinty, 1955: 77, pl. 1, figs. 4−4a. Type locality: Triton station 820, off
Palm Beach, Florida state (25fathoms, 45 m dep), USA.
Paratypes. MNRJ HSL6138 [2 sh], type locality.
Remarks. MNRJ HSL6134 was not listed in the original publication, which mentioned “other paratypes from
various Triton stations in the McGinty collection”. The lot at MNRJ contains a label indicating that it originated
from the same locality and collecting date as the holotype. T. McGinty exchanged molluscan lots, including types,
with H. S. Lopes at the Instituto Oswaldo Cruz (IOC).
mulata, Eulima [Eulimidae]. Rios & Absalão, 1990: 62, fig. 1. Type locality: Off Santa Mônica Beach, Guarapari,
Espírito Santo state, Brazil (1 m dep).
Paratype. MNRJ 5802 [1 sh], type locality (Fig. 16).
multicostata, Benthonellania [Rissoidae]. Absalão & Santos, 2004: 334, figs. 5−6. Type locality: REVIZEE
station 504, off Bahia state, Brazil (14°28’50"S, 38°54’01"W, 278 m dep).
Paratypes. MNRJ 9959 [2 sh], type locality.
myrakeenae, Petalochonchus [Vermetidae]. Absalão & Rios, 1987: 415, figs. 1−5. Type locality: Ponta de Itaipu,
Niterói, Rio de Janeiro state, Brazil.
Paratypes. MNRJ 5820 [3 spm], type locality (Fig. 9).
Remarks. The original publication stated that “paratype number 20 is deposited in the malacological
collection of the Museu Nacional do Rio de Janeiro”, without providing a catalogue number. The lot (MNRJ 5820)
contains three specimens, labeled by the author as “paratypes 19−21”.
mystica, Gemmula [Turridae]. Simone, 2005: 7, figs. 1−8. Type locality: Off Ubatuba, 24°41’01"S, 44°18’05"W,
510 m dep).
Types not located.
Remarks. The original publication listed two paratypes (shells), from the type locality, in the MNRJ
collection, without providing a catalogue number. However, they were not located in the MNRJ Mollusca
Collection.
natalensis, Bursa (Colubrellina) [Bursidae]. Coelho & Matthews, 1970: 1, figs. 1−3. Type locality: Off Pirangi,
Rio Grande do Norte state, Brazil (40−50 m dep).
Holotype. MNRJ 3527 [sh] (Fig. 11).
noronhensis, Malea [Tonnidae]. Kempf & Matthews, 1969: 57, figs. 1−5. Type locality: Atol das Rocas, Brazil
(03°52’S, 33°49’W).
Holotype. MNRJ 3493 [sh] (Fig. 10); paratype: MNRJ 3494 [1 sh], off Fernando de Noronha Archipel,
Pernambuco state, Brazil.
Remarks. Gomes et al. (2006: 194, fig. A3) figured the holotype, without type status or collection number.
notialis, Oscilla [Pyramidellidae]. Pimenta, Santos & Absalão, 2008: 175, fig.2 A−F. Type locality: Off Sergipe
state coast (11°23’21"S, 37°04’30"W, 99 m dep).
Holotype. MNRJ 10692 [sh]; paratypes: MNRJ 10942 [8 sh], REVIZEE station 6669 (24°7.42’S, 44°42.22’W,
101 m dep), São Paulo state, Brazil.
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noxia, Caelatura [Barleeidae]. Santos & Absalão, 2006a: 122, figs. 20−22, 24−27. Type locality: REVIZEE
Central station vv38, Brazil (22°00’24"S, 40°05’15"W, 100 m dep).
Holotype. MNRJ 10311 [sh].
oblatogyra, Granigyra [Seguenzioidea]. de Souza & Pimenta, 2002: 299, figs. 1−4. Type locality: Off Rio Doce,
Espírito Santo state (JOPS station 3228: 19°45.5’S, 38°45.8’W, 1100 m dep), Brazil.
Holotype. MNRJ 8433 [sh].
olearia, Brookula [Seguenzioidea]. Absalão & Pimenta, 2005: 198, figs. 28−35. Type locality: Bacia de Campos
station 61 (21°52'51.9"S, 39°48'11.680"W, 1350 m dep), Rio de Janeiro state, southeast of Brazil.
Holotype. MNRJ 9958 [sh]; paratype: MNRJ 9957 [1 sh], Bacia de Campos station 60 (21°52'50.445"S,
39°51'42.6"W, 1050 m dep), Brazil.
onca, Isotriphora Fernandes, Pimenta & Leal, 2013: 10: figs. 9−10, 45−50. Type locality: Trindade Island, VitóriaTrindade Chain, Brazil, REVIZEE Central sta C5−41F (20°30’S, 29°16’W, 360 m dep).
Holotype. MNRJ 16236 [sh]; paratypes: MNRJ 16230 [8 sh], Columbia seamount, Vitória-Trindade Chain,
Brazil, REVIZEE Ceantral sta C5−42R (22°44’'S, 31°50’W, 85 m dep).
omissa, Temesa (Temesa) [Clausiliidae]. Weyrauch, 1957: 18, pl 1, fig. 4. Type locality: road Tarma-Jauja, above
Tarmatambo (11°28’15"S, 75°41’12"W, 3400−3700 m alt), Peru, Departamento Junín.
Paratypes. MNRJ 4460 [5 sh], type locality.
Remarks. The paratype MNRJ 4460 bears a label indicating “WW 277-A/5 paratypes”; it was part of the
paratype lot from Weyrauch private collection originally published as “WW 277-A/zahlreich”, and was
subsequently donated to the MNRJ collection.
ornata, Rissoella [Rissoellidae]. Simone, 1995b: 561, figs. 1−18. Type locality: Vitória Island, São Sebastião, São
Paulo state, Brazil (23°45'S, 45°01'W).
Paratypes. MNRJ 7151 [2 spms mummified], type locality.
Remarks. Paratypes MNRJ 7151 were erroneously published under collection number MNRJ 6934, which
corresponds to the paratype of Thala crassa Simone, 1995.
pacamoni, Bursa [Bursidae]. Matthews & Coelho, 1971: 2, figs. 1−5. Type locality: Off Mucuripe Beach,
Fortaleza, Ceará state, Brazil.
Holotype. MNRJ 3528 [sh] (Fig. 12); paratypes: MNRJ 3529 [1 sh], type locality; MNRJ 3530 [1 sh], type
locality; MNRJ 3531 [1 sh], type locality; MNRJ 3532 [10 sh], type locality.
Remarks. Paratypes MNRJ 3532 were not cited in the list of material in the original publication, but one of
them was illustrated in Figure 5. One paratype from this lot was later exchanged for a paratype of Latirus
frenguellii (MACN 24442) (see also Tablado & Mantinian 2004).
pankakare, Palazzia [Turbinidae]. Absalão, 2009: 136, figs. 1E−H. Type locality: Campos Basin, BC Sul I #77
(22°36´03"S, 39°57´54"W), 1650 m.
Types not located.
Remarks. The original publication listed MNRJ 12849 as a paratype lot from Campos Basin, Brazil, BC Sul I
#85 (22°29´33"S, 39°56´17"W), 1350 m dep, without providing the number of specimens. The specimens were not
sent to the MNRJ Mollusca Collection.
papaveroi, Physa [Physidae]. Leme, 1966 : 270, figs. 2, 5 (p. 271), figs. 1−5 (p. 274), figs. 6−8 (p. 276). Type
locality: Parque D. Pedro II, São Paulo, São Paulo state, Brazil.
Paratypes. MNRJ 3514 [10 sh], type locality.
Remarks. Paratypes MNRJ 3514 were originally published under the collection number DZ 16.618 (currently
MZSP); and was subsequently donated to the MNRJ collection.
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parajeffreysi, Vetulonia [Seguenzioidea]. Absalão & Pimenta, 2005: 194, figs. 3−8. Type locality: Bacia de
Campos (22°38'46.64"W, 40°60'13.05'S, 1290 m dep), Rio de Janeiro state, southeast of Brazil.
Holotype. MNRJ 9845 [sh]; paratype: MNRJ 8912 [1 sh], REVIZEE station 50 (20°50'W, 39°14'S, 1600 m
dep), Brazil.
parviscymna, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2004a: 120, figs. 27−30. Type locality: Off Pará
state, north coast of Brazil, AMASSEDS station 3209 (01°20.93’S, 48°00.20’W, 53 m dep).
Holotype. MNRJ 8922 [sh].
parvus, Thaumastus (Thaumastus) [Bulimulidae]. Pena, Salgado & Coelho, 2011: 532, figs. 1−10. Type locality:
Serra do Curral (19°57’15.00"S; 43°55’21.77"W), Mangabeiras Park, Belo Horizonte, Minas Gerais state, Brazil.
Holotype. MNRJ 8107 [sh + fragments of dissected reproductive system]; paratypes: MNRJ 8106 [4 sh +
MEV stub with protoconch + MEV stub with jaws + fragments of dissected soft parts + 28 histological laminae],
type locality; MNRJ 8108 [32 sh], type locality; MNRJ 8103 [8 sh + fragments of dissected soft parts], type
locality; MNRJ 8104 [2 sh], type locality; MNRJ 8105 [9 sh], type locality; MNRJ 9810 [3 sh + fragments of
dissected soft parts], Municipality of Nova Lima, Belo Horizonte Peak, Minas Gerais state, Brazil; MNRJ 8109 [2
sh], Rola Moça Park, Municipality of Belo Horizonte, Minas Gerais state, Brazil; MNRJ 8102 [3 fragments of
dissected soft parts], Rola Moça Park, Municipality of Belo Horizonte, Minas Gerais state, Brazil; MNRJ 9811 [4
spm], Rola Moça Park, Municipality of Belo Horizonte, Minas Gerais state, Brazil.
pataxo, Calliotropis [Chilodontidae]. Absalão, 2009: 135, figs. 1C−D. Type locality: Brazil, Campos Basin, BC
Sul I #73 (22°41’35"S, 40°00’45"W), 1950 m.
Types not located.
Remarks. The original publication listed MNRJ 12848 from Campos Basin, Brazil, BC Sul II #77
(22°36´12"S, 39°58´22"W), 1670 m dep, as a paratype lot, without providing the number of specimens. The
specimens were not sent to MNRJ Mollusca Collection.
paulinoi, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2004a: 135, figs. 116−120. Type locality: Off Pará
state, Brazil AMASSED station 3209 (01°20.9’S, 48°00.2’W, 53 m dep), north coast of Brazil.
Holotype. MNRJ 8941 [sh].
pernambucoensis, Cerithiella [Newtoniellidae]. Lima & Barros, 2007: 64, figs. 1−9. Type locality: Continental
slope of Pernambuco, Brazil (08°46.5’00"S, 34°44.5’00"W, 690 m dep).
Holotype. MNRJ 10834 [sh]; paratypes: MNRJ 10835 [3 sh], type locality; MNRJ 10836 [1 sh], type locality.
petiti, Gergovia [Cancellariidae]. Barros & Lima, 2007: 65, figs. 12−17. Type locality: along the state of Rio
Grande do Norte, Brazil, station D−29, 06°13’22"S, 34°52’20"W, 223m.
Paratypes. MNRJ 10572 [1 sh], along the state of Alagoas, Brazil, D−31, 10°06’35"S, 35°46’41"W, 720m;
MNRJ 10573 [1 sh], along the state of Alagoas, Brazil, D−31, 10°06’35"S, 35°46’41"W, 720m.
phrix, Caelatura [Barleeidae]. Santos & Absalão, 2006a: 124, figs. 28−31, 33−34. Type locality: REVIZEE
Central V station 13B, Bahia state, Brazil (16°47’14"S, 38°41’14"W, 50 m dep).
Holotype. MNRJ 10309 [sh].
pichi, Xanthodaphne [Raphitomidae]. Figueira & Absalão, 2012: 11, figs. 16−19. Type locality: Campos Basin,
Southeast Brazil, Southwestern Atlantic (22°26’28"S, 39°58’53"W, 1046 m dep).
Holotype. MNRJ 30289 [sh].
pontesi, Volvarina [Marginellidae]. Rios & Leal, 1993: 28, figs. 1−8. Type locality: Off Itajaí, Santa Catarina state,
Brazil (27°00’S, 46°10’W, 1140 m dep).
Paratype. MNRJ 4850 [1 sh], type locality.
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potiguara, Cerberilla [Aeolididae]. Padula & Delgado, 2010: 176, figs. 1−6, 10. Type locality: Praia de Búzios,
Rio Grande do Norte state, northeastern Brazil (06°00´13"S, 35°06´24"W, intertidal).
Holotype. MNRJ 15111 [spm, dissected].
proseila, Brookula [Seguenzioidea]. Absalão & Pimenta, 2005: 196, figs. 19−24. Type locality: REVIZEE station
5 (15°34.08'S, 38°49.81'W, 50 m dep), Bahia state, northeast of Brazil.
Holotype. MNRJ 9956 [1 sh].
pseudocardinalis, Conus [Conidae]. Coltro, 2004: 7, pl. 7, fig. A, pl. 15, figs. H1, P1−P3. Type locality: 125 km
NE Abrolhos Archipelago, off Alcobaça, Bahia state, Brazil (15°57’S, 38°01’W).
Paratype. MNRJ 10222 [1 sh], type locality.
pterocladica, Tricolia affinis [Eulithidium, Phasianellidae]. Robertson, 1958: 264, pl. 144, fig 3, pl. 145, fig. 3.
Type locality: Boynton Beach, Florida.
Paratypes. MNRJ HSL5912 [5 sh], type locality (Fig. 5).
Remarks. The paratypes in the MNRJ Mollusca Collection were originally published as belonging to the
“private collection of T. McGinty”; part of that type series was later donated to the HSL collection.
puncata, Spiripockia [Pomatiopsidae]. Simone, 2012c: 516, figs. 1–33. Type locality: Serra Ramalho Karst area,
Middle São Francisco River basin, Lapa dos Peixes cave, Bahia state, Brazil (13°49’21.78"S, 43°57’24.39"W).
Paratypes. MNRJ 30503 [1 spm], type locality.
puncturina, Pseudodiala [Barleeidae]. Santos & Absalão, 2006b: 65, figs. 12−18. Type locality: REVIZEE
Central V station 30, Espírito Santo state, Brazil (20°08'S, 37°29'W, 60 m dep).
Holotype. MNRJ 10502 [sh].
pyrrocincta, Protobarleeia [Barleeidae]. Absalão, 2002: 5, figs. 9−12. Type locality: Abrolhos Bank, off Bahia
state, Brazil (about 20 m dep).
Holotype. MNRJ 8622 [sh]; paratype: MNRJ 8624 [1 sh], type locality.
quinni, Solariella [Solariellidae]. Barros & Pereira, 2008 In: Barros, Santos & Francisco, 2008: 13, figs. 15−21.
Type locality: Pernambuco state, Brazil (08°09'S, 34°34'W, 69−71 m dep).
Paratypes. MNRJ 10578 [1 sh], type locality; MNRJ 10579 [1 sh], type locality.
Remarks. Barros (2010: 60) published the description of the same name, based on the same type material.
rehderi, Bostryx (Elatibostryx) [Orthalicidae]. Weyrauch, 1960a: 35, taf. 3, figs. 4−5. Type locality: Río Huaura,
2−3 km NNE Churin (10°48’39"S, 76°52’36"W, 2400−2450 m alt), on the road to Oyón, Peru, departamento Lima.
Paratypes. MNRJ 4447 [5 sh], type locality.
Remarks. The paratype MNRJ 4447 bears a label indicating “WW 3207/5 paratypes”; it was part of the
paratype lot from Weyrauch private collection originally published as “WW 3207”, and was latter donated to
MNRJ collection. Cited in Barbosa et al. (2008) as NRJ 4447 (sic).
reticulata, Terebra [Terebra crassireticula Simone, 1999b, Terebridae]. Simone & Verissimo, 1995: 462, figs. 1−8.
Type locality: slope off Ubatuba, Station 5361, São Paulo state, Brazil (24°42’0"S, 44°30’5"W, 320 m dep).
Paratypes. MNRJ 6935 [2 spms mummified], type locality.
Remarks. MNRJ 6935 was not listed in the original publication, but contains a label by the author indicating
“sta 5361”. This corresponds to the type locality, and therefore we consider that it was part of the type material.
rhachialis, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2004a: 137, figs. 121−126. Type locality: Prainha,
Arraial do Cabo, Rio de Janeiro state, southeast coast of Brazil.
Holotype. MNRJ 8918 [sh].
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Remarks. As result of a lapsus calami, this name was spelled in more than one way in the original work. In the
abstract and figure legends, it appears as Turbonilla rhachialis, and in the description section as Turbonilla
rachialis. The etymology provided indicates that the name alludes to the Latin word “rachia”. This was an
inadvertent error by the authors, since their intention was to derive the name of the species from the Greek word
“rhachia”, meaning beach. Thus, following, ICZN (1999: Article 24.2.3 and 32.2.1), we act as the First Reviser and
select Turbonilla rhachialis as the correct original spelling.
ribeiroi, Corona [Bulimulidae]. Ihering, 1915: 8−9, figs. 1−2. Type locality: Caceres, Mato Grosso state, Brazil.
Holotype. MNRJ 3444 [sh] (Fig. 32).
Remarks. The original publication did not mention types, nor indicated the name of the collection where the
specimen was deposited, but stated that the single exemplar was found in Caceres, Mato Grosso, and provided a
color drawing of the shell in dorsal and apertural views. These images correspond to the shell in MNRJ 3444,
which bears a label written by Ihering, with the word “type”. Subsequently to the original description, this lot
received two collection numbers: 34332, corresponding to the old numbering series of the MNRJ collection, and
3444 (the current number); both numbers are written in ink on the shell.
Simone (2006a: 160, fig. 549) figured a shell referred to as holotype MZSP 34332. This collection number
(34332) is the same as the old numbering system at MNRJ, and actually MZSP 34332 is a different taxon (non-type
specimen of the family Rissoidae).
Dornellas & Simone (2011) referred to the shell illustrated by Simone (2006a) as holotype MZSP 7639, but did
not mention the erroneous number given by Simone (2006a), nor did they clarify the situation. The shell illustrated
by Simone (2006a) does not correspond to the original drawing. However, MZSP 7639 bears a label by Ihering
indicating “cotypes”. Since the original publication explicitly indicates that only one exemplar was studied and that
it corresponds to the shell of MNRJ 3444, we conclude that this shell must be considered the holotype.
The shell at MZSP 7639 could be considered a paratype, following ICZN (1999: Article 72.4.1.1); although it
was not cited in the original publication, its label indicates that it was considered by Ihering as a cotype, and is here
considered as a paratype.
ribeiroi, Drymaeus nigrogularis [Bulimulidae]. Ihering, 1915: 6. Type locality: Salto Alegre do Rio Jaurú, Juruti,
Pará state, Brazil.
Holotype. MNRJ 3454 [sh].
Remarks. The shell is marked on the internal face of the aperture with “34325”, which corresponds to the old
numbering of the MNRJ collection. The original publication did not mention the location of deposition.
The holotype was figured by Simone (2006a: 142, fig. 470).
rimula, Pseudococculina [Pseudococculinidae]. Simone & Cunha, 2003: 70, figs. 1−30. Type locality: Off
southern Rio de Janeiro state (350−400 m dep), Brazil.
Paratypes. MNRJ 8965 [2 spm], type locality.
Remarks. Page 69 of the original publication erroneously stated that the material was collected off São Paulo
state. The original publication stated that paratype MNRJ 8965 contains three specimens, but only two specimens
were found.
riosi, Conus [Conidae]. Petuch, 1986: 10, figs. 5−6. Type locality: Off Salvador, Bahia state, Brazil (50 m dep).
Paratype. MNRJ 8460 [1 sh], type locality.
Remarks. Originally published under collection the number MORG 14242; later donated to the MNRJ
collection.
riosi, Typhina [Siphonochelus, Muricidae]. Bertsch & D’Áttilio, 1980: 135, figs. 6−7. Type locality: Off
Tramandaí, Rio Grande do Sul state, southern Brazil (30°04’S, 47°55’W, 280 m dep).
Paratype. MNRJ 4863 [1 sh], type locality (Fig. 17).
riosi, Ketosia [Barleeidae]. Santos & Absalão, 2006b: 61, figs. 1−5. Type locality: Povoação, Espírito Santo state,
Brazil (21°36'32.537"S, 40°39'18.846"W, 10−30 m dep).
Holotype. MNRJ 10503 [sh]; paratypes: MNRJ 10504 [2 sh], type locality.
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rodriguezae, Bostryx (Bostryx) [Orthalicidae]. Weyrauch, 1967b: 475, figs. 33−48. Type locality: left margin of
Mayo Creek (Río Laraos), tributary to Río Cañete, 4 km from Laraos on the trail to (Cerro) Quichao (12°20’39"S,
75°48’47"W, 3500 m alt), Peru, departamento Lima.
Paratypes. MNRJ 4439 [15 sh], type locality.
Remarks. The paratype lot MNRJ 4439 bears a label indicating “WW 3331/15 paratypes”; it was part of the
paratype lot from Weyrauch private collection originally published as “WW 3331/1612”, and was later donated to
the MNRJ collection.
rudigerbieleri, Solatisonax [Architectonicidae]. Tenório, Barros, Francisco & Silva, 2011: 175, fig. 1. Type
locality: Off Pernambuco state, Brazil (08°46.5'S, 34°44.5'W, 690 m dep).
Holotype. MNRJ 13021 [sh]; paratypes: MNRJ 13022 [1 sh], off Alagoas state, Brazil, (10°06'S, 35°46'W,
720 m dep); MNRJ 13023 [1 sh], off Pernambuco state, Brazil (06°25.6'S, 34°43.9'W, 425 m dep); MNRJ 13024 [1
sh], off Pernambuco state, Brazil (08°45.1'S, 35°44.9'W, 500 m dep); MNRJ 13025 [1 sh], off Pernambuco state,
Brazil (09°04.7'S, 34°51.2'W, 520 m dep); MNRJ 13026 [1 sh], type locality.
saldanha, Mitra (Cancilla) [Mitridae]. Matthews & Rios, 1970: 61, figs. 1−2. Type locality: Off Cabo Cassiporé,
Amapa state, Brazil (04°30’N, 50°03’W, 96 m dep).
Paratype. MNRJ 3503 [1 sh], type locality (Fig. 25).
sanjosensis, Pomacea cumingii [Ampullariidae]. Morrison, 1946: 6, pl. 1, fig. 1. Type locality: weast side San
Jose Islands, Peal Islands, Panama.
Paratype. MNRJ 3488 [1 sh], type locality (Fig. 7).
Remarks. The original description described the holotype, deposited at USNM, and mentioned that “the
largest of many species seen…”, which indicates that paratypes were present, although not explicitly discriminated.
Lot MNRJ 3488 contains a label giving the same locality and date as the holotype. This specimen was donated to
MORG and then to MNRJ.
santosi, Mitromorpha [Turridae]. Lima, Barros & Francisco, 2010: 600, figs. 1A−H. Type locality: continental
slope off Rio Grande do Norte state, Brazil (4°51’40"S, 35°06’01"W, 384 m dep).
Paratypes. MNRJ 10717 [2 sh], off Rio Grande do Norte state, Brazil Natureza station 22 (4°50’60"S,
35°06’46"W, 375 m dep).
scapulata, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2004a: 135, figs. 111−115. Type locality: Off São
Paulo state, PADCT station 6579 (24°42.3’S, 45°18.8’W, 84 m dep), southeast coast of Brazil.
Paratype. MNRJ 8920 [1 sh], type locality.
schirrmeisteri, Conus [Conidae]. Coltro, 2004: 4, pl. 3, fig. C, pl. 11, figs. H1, P1−P11. Type locality: Sulfur Bank,
125 km NE Abrolhos Archipelago, off Alcobaça, Bahia state, Brazil (15°50’S, 37°57’W).
Paratypes. MNRJ 10192 [1 sh], type locality; MNRJ 10193 [1 sh], type locality.
scopulorum, Conus [Conidae]. Van Mol, Tursh & Kempf, 1971: 162, figs. 3−6. Type locality: Fernando de
Noronha Archipelago, Pernambuco state, Brazil (3°53’N, 32°37.2’W, 60−90 m dep).
Types not located.
Remarks. The original publication listed a paratype lot in the MNRJ collection, with one shell, without
indicating the collection number. This shell was probably never received by the MNRJ collection.
scymnocelata, Boonea [Pyramidellidae]. Pimenta, Absalão & Miyaji, 2009: 48, figs. 15−20. Type locality: Off
Paraná state, PADCT station 6595 (26°23.55’S, 46°39.49’W, 175 m dep).
Paratypes. MNRJ 10957 [8 sh], type locality.
seraphina, Oenopota [Turridae]. Figueira & Absalão, 2010b: 463, figs. 1A−C. Type locality: Campos Basin,
southeast Brazil, southwestern Atlantic (22°26’28"S, 39°58’53"W, 1046 m dep).
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Paratypes. MNRJ 13552 [5 sh], Campos Basin, southeast Brazil (22°10’43"S, 39°54’46"W, 750 m dep);
MNRJ 13553 [4 sh], Campos Basin, southeast Brazil (21°52’59"S, 39°55’30"W, 750 m dep).
simonei, Terebra [Terebridae]. Lima, Tenório & Barros, 2007: 64, figs. 1−7. Type locality: Brazil, continental
slope off Pernambuco #D-1 (06°25'36"S, 34°43'54"W, 425 m dep.).
Paratypes. MNRJ 11138 [2 sh], type locality.
speculabunda, Caelatura [Barleeidae]. Absalão, 2002: 4, figs. 6−8. Type locality: Dogaressa Seamount, MD 55
station 43 (20°51'S, 33°45'W, 63 m dep), Vitória-Trindade Seamount Chain, off Southeastern Brazil.
Holotype. MNRJ 8621 [sh].
spinulata, Brookula [Seguenzioidea]. Absalão, Miyaji & Pimenta, 2001: 685, fig. 5 a−g. Type locality: South of
Abrolhos Bank, JOPS station 3229, (19°38.5’S, 38°43’W, 775 m dep), Brazil.
Holotype. MNRJ 8424 [sh]; paratypes: MNRJ 8425 [2 sh], type locality.
spirocordata, Caelatura [Barleeidae]. Absalão & Rios, 1995: 87, figs. 1−3. Type locality: Off Atafona,
northeastern Rio de Janeiro state, southeastern Brazil (21°35’40"S, 40°44’35"W, 16.8 m dep).
Paratype. MNRJ 6983 [1 sh], off northeastern Rio de Janeiro state, Brazil (21°31’35"S, 40°19’00"W, 41 m
dep).
spirosulcata, Terebra [Terebridae]. Simone & Costa in Simone, 1999b: 237, figs. 1E−F, 3H, 6H−I, 7E, 8D−E, 13C,
25−26. Type locality: Off Ilha do Pai, Rio de Janeiro state, Brazil (30−40).
Holotype. MNRJ 3657 [sh]; paratypes: MNRJ 3655 [1 sh], off Cagarras Islands, Rio de Janeiro, Brazil (30−40
m dep); MNRJ 7570 [sh], type locality; MNRJ 7571 [2 sh], off Cagarras Islands, Rio de Janeiro state, Brazil
(30−40 m dep).
spongiosa, Jorunna [Discodorididae]. Alvim & Pimenta, 2013: 182, figs. 2A−B, 19−20. Type locality: Canal de
Itajurú, Cabo Frio, Rio de Janeiro state, Brazil (22°53’13.73"S; 42°00’60"W, 2 m dep).
Holotype. MNRJ 13781 [spm, dissected]; paratypes: MNRJ 14971 [2 spm, dissected], type locality; MNRJ
13203 [2 spm, dissected], type locality; MNRJ 13197 [1 spm], type locality; MNRJ 13996 [1 spm], type locality;
MNRJ 30823 [1 spm], type locality.
straubei, Thaumastus [Bulimulidae]. Colley, 2012: 44, figs. 1−9. Type locality: Área de Proteção Ambiental de
Guaratuba, Paraná state, Brazil (Usina Hidrelétrica de Guaricana).
Holotype. MNRJ 11890 [spm]; paratypes: MNRJ 17272 [1 spm + 2 soft parts + 2 dissected soft parts + 6 sh],
type locality.
Remarks. Colley (2012) mentioned the same collection number for the holotype and to the paratypes, being
seven exemplars (one young) and seven shells. The lot MNRJ 11890 includes six specimens (shells + soft parts)
and three empty shells. The holotype shell and its dissected soft parts could be recognized, following the original
illustrations and is kept under the collection number MNRJ 11890. The paratypes are here split to a separate lot
(MNRJ 17272).
tanata, Taranis [Turridae]. Figueira & Absalão, 2010a: 736, figs. 1G, H, I. Type locality: Campos basin, southeast
Brazil (22°33’08"S, 39° 54’21"W, 1934 m dep).
Paratypes. MNRJ 13557 [2 sh], (21°52’44"S, 39° 40’45"W, 1950 m dep), Campos Basin, southeast Brazil.
teaguei, Olivancillaria [Olividae]. Klappenbach, 1965: 132, pl. 8, figs. 1, 3−5, 8. Type locality: La Coronilla,
Departament of Rocha, Uruguay.
Paratype. MNRJ 3399 [1 sh], type locality.
Remarks. Published as Mus. Nac. Hist. Nat. Montevideo No. 1218; later donated to the MNRJ collection.
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tephrina, Fasciolaria [Fasciolariidae]. de Souza, 2002: 2, figs. 1−5. Type locality: North of Quita Sueño Bank,
Colombia, off Nicaragua (14°40'N, 81°25'W, 480 m dep).
Paratype. MNRJ 8668 [1 sh], type locality.
tessa, Anostoma [Orthalicidae]. Simone, 2012a: 435, figs. 15–20. Type locality: Santa Maria da Vitória, Bahia
state, Brazil (~13°24’S, 44°12’W, ~460 m alt).
Paratype. MNRJ 30379 [2 sh], type locality.
Remarks. Original description did not provide the catalogue number for MNRJ paratypes.
thetys, Nassarina [Columbellidae]. Costa & Absalão, 1998: 279, figs. 1, 4−5, 8−9, 11. Type locality: Off
Guarapari, Espírito Santo state, Brazil (20°41’S, 40°22’W, 20−30 m dep).
Paratype. MNRJ 5727 [1 sh], between Rio de Janeiro (19°30’S, 39°40’) and Espírito Santo (22°30’, 41°),
Brazil, 50m.
thomei, Ketosia [Barleeidae]. Santos & Absalão, 2006b: 62, figs. 6−11. Type locality: GEOMAR XVI station 17
Am.19, off Rio de Janeiro state, southeast of Brazil (22°27'S, 40°58'W).
Holotype. MNRJ 10505 [sh].
tigrina, Caelatura [Barleeidae]. Absalão, 2002: 2, figs. 1−5. Type locality: Davis Seamount, MD 55 station DC41
(20°39'S, 34°43'W, 58−70 m dep). Vitória-Trindade Seamounts chain, off Southeastern Brazil.
Holotype. MNRJ 8619 [sh]; paratypes: MNRJ 8620 [5 sh], type locality.
tigrina, Isotriphora Fernandes, Pimenta & Leal, 2013: 8, figs 7−8, 39−44. Type locality: Trindade Island, VitóriaTrindade Chain, Brazil, MD55 sta 35−DC59 (20°30’ S, 29°19’ W, 52–60 m dep).
Holotype. MNRJ 25992 [sh]; paratypes: MNRJ 16227 [2 sh], Trindade Island, Vitória-Trindade Chain, Brazil,
REVIZEE C5−49R (20°31’S, 29°21’W, 57 m dep); MNRJ 25993 [7 sh], type locality.
tiradentensis, Thaumastus (Thaumastus) [Bulimulidae]. Pena, Coelho & Salgado, 1996: 3, figs. 2, 12, 14−20.
Type locality: “Casa de Pedra”, Tiradentes, Minas Gerais state, Brazil (21°06’32"S, 44°11’00"W).
Holotype. MNRJ 6923 [sh + dissected soft parts]; paratypes: MNRJ 2852 [11 sh], type locality; MNRJ
HSL7814 [10 sh], type locality; MNRJ 6922 [12 sh + 5 young sh + 2 empty eggs + 8 soft parts, dissected or not +
20 laminae], type locality.
Remarks. As result of a lapsus calami, the original publication inverted the photographs of the dorsal views of
the holotype of this species and the other species described in the same paper. Pena et al. (1996: page 10, fig. 13) is
actually a photograph of the holotype of Thaumastus baixoguanduensis, while Pena et al. (1996: page 7, fig. 2) is
the photograph of the holotype of T. tiradentensis in dorsal view.
Holotype figured by Simone (2006a: 155, fig. 528).
torquata, Eulimella [Pyramidellidae]. Pimenta, Santos & Absalão, 2011: 28, figs. 19−25. Type locality: Off Bacia
de Campos (1,084 m dep), Rio de Janeiro state, southeast of Brazil.
Holotype. MNRJ 13251 [sh]; paratypes: MNRJ 13095 [1 sh], off Bacia de Campos (1,080 m dep),
southeastern Brazil; MNRJ 13096 [1 sh], off Bacia de Campos (1,080 m dep), southeastern Brazil; MNRJ 13252 [1
sh], type locality; MNRJ 13506 [1], off Bacia de Campos (1170 m dep), southeastern Brazil.
tostesi, Conus [Conidae]. Petuch, 1986: 11, figs. 9−10. Type locality: Off Cabo Frio, Rio de Janeiro state, Brazil
(100 m dep).
Paratype. MNRJ 8461 [1 sh], type locality.
Remarks. Published under the collection number MORG 14247; later donated to the MNRJ collection.
trindadense, Caecum [Caecidae]. Lima, Santos & Absalão, 2013: 773, figs. 7−10. Type locality: Off the coast of
Rio de Janeiro state, southeastern Brazil (21°38’S, 40°10’W, 140 m dep).
Paratypes. MNRJ 31260 [1 sh], Trindade Island, Trindade and Martim Vaz Archipelago, Espírito Santo state,
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sotheastern Brazil; MNRJ 31261[5 sh], continental shelf of Campos Basin, Rio de Janeiro (22°23’22.23"S,
40°34’59.11"W, 110 m dep); MNRJ 31262 [4 sh], continental shelf of Campos Basin, Rio de Janeiro
(22°17’42.21"S, 40°27’00.05"W, 103 m dep); MNRJ 31263 [1 sh], continental shelf of Campos Basin, Rio de
Janeiro (22°23’39.34"S, 40°20’41.32"W, 149 m dep).
trindadensis, Bulimulus [Vegrandinia, Bulimulidae]. Breure & Coelho, 1976: 3−5, figs. 1−4, tab. 1. Type locality:
Praia dos Cabritos, Ilha da Trindade, Espírito Santo state, Brazil.
Holotype. MNRJ 3711 [sh] (Fig. 33); paratypes: MNRJ 3712 [151 sh], type locality.
Remarks. The original publication listed “many” paratypes in the lot MNRJ 3712.
Simone (2006a: 120, fig. 372) figured the holotype, but indicated a wrong collection number (MNRJ 5711). Gomes
et al. (2006: 195, fig. C4) also figured the holotype, but neither provided the type status nor the collection number
of the illustrated shell. Holotype figured by Salvador et al. (2013).
troglobius, Pothamolithus [Hydrobiidae]. Simone & Morachioli, 1994: 456, figs. 2g−i, 3, 4c, 5c, 7c, 8c, 10c, 11c.
Type locality: Areias stream in Areias Caves, Iporanga, São Paulo state, Brazil (24°35’20"S, 48°42’05"W).
Paratypes. MNRJ 6937 [2 spm], type locality.
tunicata, Volvarina [Marginellidae]. Boyer, 2000: 47, figs. 24−27. Type locality: Guarapari, Espírito Santo state,
(under rocks, 20−25 m dep), Brazil.
Paratype. MNRJ 8411 [1 sh], type locality.
Remarks. The original publication did not provide the collection number.
tupi, Caelatura [Barleeidae]. Santos & Absalão, 2006a: 125, figs. 35−39. Type locality: REVIZEE Central V
station 13B, Bahia state, northeast of Brazil (16°47’14"S, 38°41’14"W, 40 m dep).
Holotype. MNRJ 10310 [sh].
tupinamba, Calliostoma. Dornellas, 2012: 91, figs. 1–9, 16–28. Type locality: Angra dos Reis, Jorge Grego Island,
Rio de Janeiro state, Brazil (23°13′S, 44°08′W, 5–7 m dep.).
Paratype. MNRJ 30602 [2 spm], Vitória Island, Saco do Paiá, São Paulo state, Brazil (23°44.658′S,
45°01.343′W, 6–9 m dep).
tupinamba, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2002: 11−13, figs. 33−37. Type locality: Off Prainha
(22°58'S, 042°00'W), Arraial do Cabo, Rio de Janeiro state, Southeast of Brazil.
Holotype. MNRJ 8944 [sh].
uaca, Turbonilla [Pyramidellidae]. Pimenta & Absalão, 2004a: 137, figs. 127−132. Type locality: Off Amapá
state, north coast of Brazil.
Paratype. MNRJ 8942 [1 sh], type locality.
urueuauau, Mirachelus [Chilodontidae]. Absalão, 2009: 134, figs. 1A−B. Type locality: Brazil, Campos Basin,
BC Sul I #75 (22°31´28"S, 40°03´50"W), 1050 m.
Types not located.
Remarks. The original publication listed MNRJ 12847 from Campos Basin, Brazil, BC Sul II #80
(22°24´30"S, 39°57´28"W), 1044 m dep, as a paratype lot, without providing the number of specimens. The
specimens were not sent to MNRJ Mollusca Collection.
valeriae, Alvania [Rissoidae]. Absalão, 1993: 104, figs. 1−3. Type locality: Off southeastern of Espírito Santo
state, southeastern of Brazil (20°38’12"S, 40°16’12"W, 38 m dep).
Paratypes. MNRJ 5978 [2 sh], off Rio de Janeiro state, Brazil (23°16’50"S, 43°02’40W, dep 92 m dep),
Almirante Saldanha station 6174.
Remarks. The original publication listed one shell as paratype 4, but MNRJ 5978 contains two shells from the
same locality, both labeled by author indicating “paratype 4”.
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verheckeni, Brocchinia [Cancellariidae]. Barros & Lima, 2007: 60, figs. 1−5. Type locality: Along the state of Rio
Grande do Norte, Brazil (“Natureza”, station D−27, 06°14’24"S, 34°52’06"W), 500 m dep.
Paratypes. MNRJ 10575 [1 sh], along the state of Ceará, Brazil, station D−20, 03°30’51"S, 37°59’28"W, 384
m dep; MNRJ 10576 [1 sh], along the state of Alagoas, Brazil, station D−32, 09°20’30"S, 34°59’11"W, 452 m dep;
MNRJ 10577 [1 sh], along the state of Sergipe, Brazil, station D−06, 10°41.4’00"S, 36°18’07"W, 365 m dep.
vesica, Bulla [Olivancillaria, Olividae]. Gmelin, 1791: 3433. Type locality: Maricá, Rio de Janeiro state, Brazil.
Neotype: MNRJ 18979 [sh].
Remarks. The neotype was designated and illustrated by Teso & Pastorino (2011).
vilchezi, Bostryx (Bostryx) [Orthalicidae]. Weyrauch, 1960a: 32, taf. 3, figs. 8−10. Type locality: Río Guineamaya,
20 km NE Cutervo, Sócota (06°18’59"S, 78°44’37"W, 1820 m alt), Peru, Departamento cajamarca.
Paratypes. MNRJ 4441 [11 sh], type locality.
Remarks. The paratype MNRJ 4441 bears a label indicating “WW 2009/10 paratypes”; it was part of the
paratype lot from Weyrauch private collection originally published as “WW 2009/zahlreich”, and was later donated
to MNRJ.
viperinus, Nanomelon [Volutidae]. Leal & Bouchet, 1989: 8, figs. 9−10, 15−18, 23−25, 28−33. Type locality:
Continental slope off coast of Rio de Janeiro state, Brazil (23°47’S, 42°10’W, 610 m dep).
Paratypes. MNRJ 5762 [2 sh], MD 55 station CB104 (23°42’S, 42°07’W, 430−450 m dep), Continental slope
off coast of Rio de Janeiro state, Brazil.
virgulinoi, Turbonilla [Pyramidellidae]. Absalão, Santos & Tenório, 2003: 3, figs. 1−3. Type locality: Off
Pernambuco state, northeast coast of Brazil REVIZEE station 23 (08°46’01"S, 34°44’02"W, 690 m dep).
Holotype. MNRJ 9823 [sh].
vossi, Nanomelon [Volutidae]. Leal & Rios, 1990: 317, figs. 1−5. Type locality: Continental slope off coast of Rio
Grande do Sul state, Brazil (32°25’S, 50°11’W, 460 m dep).
Paratype. MNRJ 5767 [1 spm], type locality.
willianseverii, Heliacus [Architectonicidae]. Tenório, Barros, Francisco & Silva, 2011: 186, fig. 5). Type locality:
Off Pernambuco state, Brazil (09°07'S, 34°53'W, 104 m dep).
Holotype. MNRJ 13033 [sh]; paratypes: MNRJ 13034 [1 sh], off Alagoas state, Brazil, (09°28'S, 35°04'W,
175 m dep); MNRJ 13035 [1 sh], off Alagoas state, Brazil, (09°28'S, 35°04'W, 175 m dep); MNRJ 13036 [1 sh], off
Alagoas state, Brazil, (09°28'S, 35°04'W, 175 m dep); MNRJ 13037 [1 sh], off Alagoas state, Brazil, (09°28'S,
35°04'W, 175 m dep); MNRJ 13038 [1 sh], off Alagoas state, Brazil, (09°28'S, 35°04'W, 175 m dep); MNRJ 13039
[1 sh], type locality; MNRJ 13040 [1 sh], type locality; MNRJ 13041 [1 sh], MNRJ 13042 [1 sh], off Rio Grande
do Norte state, Brazil (04°15'S, 37°53'W, 177 m dep).
xacriaba, Ponderinella [Tornidae]. Absalão, 2009: 138, figs. 2E−H. Type locality: Campos Basin, BC Sul I, #73
(22°41´35"S, 40°00´45"W), 1950 m.
Types not located.
Remarks. The original publication listed MNRJ 12851 from Campos Basin, Brazil, BC Sul I #77
(22°36´03"S, 39°57´54"W), 1650 m dep, as a paratype lot, without providing the number of specimens. The
specimens were not sent to MNRJ Mollusca Collection.
xaioca, Gymnobela [Raphitomidae]. Figueira & Absalão, 2012: 20, figs. 33−34. Type locality: Campos Basin,
Southeast Brazil, Southwestern Atlantic (22°35’S, 39°58’−22°32’S, 39°56’W, 1620−1623 m dep).
Holotype. MNRJ 30287 [sh].
xerente, Adeuomphalus [Turbinidae]. Absalão, 2009: 136, figs. 2A−D. Type locality: Campos Basin, BC Sul II
#85 (22°30´21"S, 39°56´53"W), 1353 m.
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Types not located.
Remarks. The original publication listed MNRJ 12850 from Campos Basin, Brazil, BC Sul II #86
(22°31´37"S, 39°55´14"W), 1630 m dep, as a paratype lot, without providing the number of specimens. The
specimens were not sent to MNRJ Mollusca Collection.
ybessa, Pleurotomella [Raphitomidae]. Figueira & Absalão, 2012: 24, figs. 51−55. Type locality: Campos Basin,
Southeast Brazil, Southwestern Atlantic (22°45’S, 40°19’−22°44’S, 40°16’W, 1118−1130 m dep).
Holotype. MNRJ 30285 [sh].
zelindae, Oliva (Plicoliva) [Plicoliva, Volutidae]. Petuch, 1979: 521, figs. 3D−E, 4A−B. Type locality: south side
of Guaratibas Reefs, North end of Abrolhos Reef complex, Bahia state, Brazil (17°25’S, 39°8’W).
Paratype. MNRJ 3734 [1 sh], Santa Barbara Island, Abrolhos Archipelago, Bahia state, Brazil (17°58’S,
38°42’W).
Remarks. The original publication reffered to MNRJ as “Museu Nacional do Brasil”.
zumbii, Ancistrobasis [Seguenziidae]. Lima, Christoffersen & Barros, 2013: 104, figs 1A−G. Type locality: Off
Alagoas, northeast Brazil (10°06’35"S, 35°46’41"W, 720 m dep).
Paratype. MNRJ 29764 [1 sh], type locality.
Class Cephalopoda
gaucha, Eledone [Octopodidae]. Haimovich, 1988: 82, figs. 2−14, tab. 1. Type locality: south Rio Grande, Rio
Grande do Sul state, Brazil (32°58’S, 51°19’W, 56 m dep).
Paratypes. MNRJ 5626 [1 spm, male]; MNRJ 5627 [1 spm, female], (33°02’S, 51°30’W, 130 m dep).
insularis, Octopus [Octopodidae]. Leite & Haimovici in Leite, Haimovici, Molina & Warnke, 2008: 66, figs. 2−5.
Type locality: behind Belmont Island, St. Peter and St. Paul Archipelago, Brazil (0°55’00"S, 29°20’76"W, 10 m
dep).
Paratypes. MNRJ 10368 [1 spm, female], Buraco do Inferno, behind Rata Island, Fernando de Noronha
Archipelago (03°48’36"S, 32°23’00"W, 10 m dep); MNRJ 10369 [1 spm, male], Buraco da Rachel, behind
Fernando de Noronha Island, Fernando de Noronha Archipelago (03°50’12"S, 32°24’00"W, 2 m dep).
jurujubai, Sepia officinalis [Sepiidae]. Oliveira, 1940: 29, one figure (not numbered). Type locality: Praia de
Jurujuba, Niterói, Rio de Janeiro state, Brazil.
Holotype. MNRJ 3500.
Remarks. The original description was based on a single specimen, without reference to a location where the
collection is deposited. Lot MNRJ 3500 contains a label on which is written “holotype”. This specimen is
deteriorated; the gladius, jaws and otolith remains.
Class Bivalvia
arariboia, Yoldiella [Yoldiidae]. Benaim & Absalão, 2011b: 521, fig. 6. Type locality: Bacia de Campos, station 60
(21°52′50 45′′S, 39°51′41 67′′W, 1,050 m dep), Rio de Janeiro state, Brazil.
Holotype. MNRJ 15202 [valve]; paratypes: MNRJ 15206 [20 valves], type locality.
aurora, Semele [Semelidae]. Tursch & Pierret, 1964 : 35, figs. 1−3. Type locality: Off Rio de Janeiro, Rio de
Janeiro state, Brazil; 30 fathoms.
Paratype. MNRJ 3116 [1 sh], type locality.
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besnardi, Americuna [Warrana, Condylocardiidae]. Klappenbach, 1963: 14, figs. 1−3, pl 1. Type locality: La
Paloma Beach, Departamento de Rocha, Uruguay.
Paratypes. MNRJ 2810 [13 valves], Ponta dos Anjos (5 m dep), Arraial do Cabo, Rio de Janeiro state.
Remarks. Published under collection number 0999 Col. Malac., M.N.H.N. (currently MNHNM);
subsequently donated to the MNRJ collection.
brasiliana, Echinochama [Arcinella, Chamidae]. Nicol, 1953: 386, figs. 1−4. Type locality: Off Ilha do Francês,
Santa Catarina state, Brazil.
Paratype. MNRJ HSL2366 [1 valve], type locality.
Remarks. The original publication stated that one paratype was deposited at “Instituto Oswaldo Cruz (Rio de
Janeiro)”, later donated to the MNRJ collection.
celeste, Lyonsia [Lyonsiidae]. Pimenta & Oliveira, 2013: 79, figs. 3−4. Type locality: Maldonado Bay, Uruguay.
Paratype. MNRJ 18565 [1 sh]. off Rio de la Plata, Albatross sta 2765 (36Σ43͛S, 56Σ23͛W, 19.2 m dep).
chodo, Triplodon [Hyriidae]. Mansur & Pimpão, 2008: 112, figs 1−6. Type locality: Rio Aripuanã, Amazonas
state, Brazil.
Paratype. MNRJ 5811 [1 valve], Oriximinã, Tabuleiro do Jacaré, Trombetas River, state, Brazil.
curupira, Yoldiella [Yoldiidae]. Benaim & Absalão, 2011b: 523, fig. 8A−H. Type locality: Bacia de Campos,
station 38 (22°41′19′′S, 40°14′06′′W, 1100 m dep), Rio de Janeiro state, Brazil.
Holotype. MNRJ 15203 [valve]; paratype: MNRJ 15207 [1 valve], Bacia de Campos, station 82 (22°28′46
5′′S, 39°53′27 9′′W, 1650 m dep), Rio de Janeiro state, Brazil.
Remarks. Paratype MNRJ 15207 was originally published as MNRJ 15205, same collection number for the
paratype of Yoldiella paranapuensis; after examination of the labels on the lots, we consider the duplication of the
number was a lapsus calami.
flemingi, Gaimardia trapesina [Cyamiidae]. Powell, 1955: 28, pl. 1, fig. 4. Type locality: Auckland Islands, main
island, opposite Rose Island, New Zealand.
Paratypes. MNRJ HSL4060 [2 valves], opposite Rose Island, Auckland Island, New Zealand (Fig. 36).
Remarks. The original publication mentioned the holotype and three paratypes, giving measurements of
length, height and diameter (both valves) for each type. Lot MNRJ HSL4060 was not cited in the original
publication. This lot contains two loose valves, probably corresponding to the same shell; its length, height and
diameter are somewhat different from the measurements provided in the original publication for the three paratypes
mentioned. The catalogue by H. S. Lopes states: “Powell det et leg. (paratypes)”. Therefore, we consider that,
despite the minor differences in the measurements (probably due to a different measurement method), the shell in
MNRJ HSL4060 was part of the type series, later exchanged by Powell and H. S. Lopes.
guidoi, Mactra [Mactridae]. Signorelli & Scarabino, 2010: 34, figs. 13−24. Type locality: 60 km ENE of Cabo
Polonio, Uruguay (34°10’S, 53°04’W, 29 m dep).
Paratypes. MNRJ 9261 [2 spm], 23 km southeast of Chuy, Uruguay (33°52’S, 53°10’6"W, 22 m dep).
jarauara, Protocuspidaria (Bidentaria) [Protocuspidariidae]. Oliveira & Absalão, 2009: 153, figs.5G−M. Type
locality: Campos Basin (21°58’36"S, 39°46’30"W, 1700 m dep), Rio de Janeiro state, Brazil.
Paratypes. MNRJ 12860 [2 valves], Campos Basin (22°10’54"S, 39°48’59"W,1350 m dep), Rio de Janeiro
state, Brazil.
kaiwa, Myonera [Cuspidariidae]. Oliveira & Absalão, 2009: 145, figs. 3A−I. Type locality: 21°52’41"S,
39°46’17"W, 1650 m dep, Campos Basin, Rio de Janeiro state, Brazil.
Paratypes. MNRJ 12859 [2 valves], Campos Basin (22°36’03"S, 39°57’54"W, 1650 m dep), Rio de Janeiro
state, Brazil.
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lapernoi, Yoldiella [Yoldiidae]. Benaim & Absalão, 2011b: 515, fig. 2A−G, J. Type locality: Bacia de Campos,
station 41 (22°39′34 36′′S, 40°08′22 27′′W, 1200 m dep), Rio de Janeiro state, Brazil.
Holotype. MNRJ 15200 [valve].
lepidoformis, Acar [Arcidae]. Francisco, Barros & Lima, 2012: 1144, figs. 13−18. Type locality: Off the state of
Rio Grande do Sul, Brazil, ‘Almirante Saldanha’ sta 2233A (30°41’S, 49°31’W, 141 m dep).
Holotype. MNRJ 11124 [sh]; paratypes: MNRJ 11129 [1 v], type locality.
Remarks. The original publication mentioned that a paratype from MNRJ 11129 was illustrated in fig. 14,
however it actually corresponds to the internal view of a holotype’s valve.
lopesi, Mytilopsis [Dreissenidae]. Alvarenga & Ricci, 1989: 27, figs. 1−9. Type locality: Rio Tocantins, Tucuruí,
Pará state, Brazil.
Holotype. MNRJ 5930 [sh]; paratypes: MNRJ 5931 [22 sh], type locality; MNRJ 5932 [388 spm], type
locality; MNRJ 5933 [1 sh, metalized], type locality; MNRJ 5934 [1256 spm], Vila Pederneira, Rio Tocantins,
Tucuruí, Pará state, Brazil; MNRJ 5935 [1026 spm], Nazaré dos Patos, Tucurui, Rio Tocantins; MNRJ 5936 [120
sh], Nazaré dos Patos, Tucurui, Rio Tocantins.
macaron, Microgloma [Yoldiidae]. Benaim & Absalão, 2011a: 10, figs. 14−22. Type locality: Bacia de Campos,
station 54 (21°57′17.5′′S, 39°56′01.1′′W, 750 m dep), Rio de Janeiro state, Brazil.
Holotype. MNRJ 19112 [sh].
mattogrosensis, Fossula balzani [Mycetopodidae]. Ihering, 1915: 13, figs. 4−5. Type locality: Paraguay River,
Mato Grosso (do Sul) state, Brazil.
Holotype. MNRJ 34322 [sh].
Remarks. The original publication did not provide the catalogue numbers of malacological collections. The
shell was later catalogued in the MNRJ Collection. Holotype figured by Simone (2006a: 286, fig. 1015B).
micella, Cardita (Carditamera) [Cardites, Carditidae]. Penna, 1971: 158, fig. 2, est. 2. Type locality: Bay of Ilha
Grande, Rio de Janeiro state, Brazil; est. 302.
Paratype. MNRJ 3547 [1 spm], Bay of Ilha Grande, Rio de Janeiro state, Brazil est. 132 (24 m dep).
Remarks. Published under the collection number MZUSP 18471; later donated to the MNRJ collection.
naturezae, Acar [Arcidae]. Francisco, Barros & Lima, 2012: 1143, figs. 7−12. Type locality: Off the state of
Sergipe, Brazil, ‘Natureza’ sta D-08 (11°58’S, 36°49’W, 100 m dep).
Paratypes. MNRJ 11128 [1 v], type locality; MNRJ 11135 [2 v], type locality; MNRJ 11136 [3 v], type
locality.
Remarks. Original publication mentioned that the holotype is in the lot MNRJ 11128, which indeed bears a
label, written by the authors as holotype; however, the valve in this lot does not correspond to the holotype’s figure
in the original publication and thus must be considered a paratype. The holotype itself was not sent to MNRJ
collection.
nhanduti, Microgloma [Yoldiidae]. Benaim & Absalão, 2011a: 13, figs. 23−31. Type locality: Bacia de Campos,
Station 54 (21°57′17.5′′S, 39°56′01.1′′W, 750 m dep), Rio de Janeiro state, Brazil.
Holotype. MNRJ 19113 [sh].
oliveirae, Acar [Arcidae]. Francisco, Barros & Lima, 2012: 1147, figs. 19−24. Type locality: Off the state of
Sergipe, Brazil, ‘Natureza’ sta D-06 (10°41’S, 36°18’W, 365 m dep).
Holotype. MNRJ 11125 [v]; paratypes: MNRJ 11132 [1 v], off the state of Sergipe, Brazil, ‘Natureza’ sta D-08
(11°58’S, 36°49’W, 100 m dep); MNRJ 11131 [2 v], off the state of Pernambuco, Brazil, ‘Natureza’ sta D-11
(08°46’S, 34°44’W, 690 m dep); MNRJ 11130 [1 v], off the state of Rio Grande do Norte, Brazil, ‘Natureza’ sta D22 (04°15’S, 37°12’W, 206 m dep).
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ouricuri, Verticordia [Verticordiidae]. Oliveira & Absalão, 2010: 813, figs. 10−15. Type locality: Campos Basin
(22°37’54"S 40°13’36"W, 1000 m dep), Rio de Janeiro state, Brazil.
Paratypes. MNRJ 14685 [3 valves], Campos Basin (21°57’26"S, 39°40’33"W, 1950 m dep), Rio de Janeiro
state, Brazil.
paraibanensis, Crassostrea [Ostreidae]. Singarajah, 1980: 837, figs. 2−3. Type locality: Salinas, Paraíba River
Estuary, Paraíba state, Brazil.
Paratypes. MNRJ 3815 [1 sh], type locality; MNRJ 3816 [1 sh], type locality.
Remarks. The original publication referred to MNRJ as “Rio de Janeiro National Museum (RJNM)”. In the
original publication, it is indicated that the specimens of the type series were found at two locations on the Paraíba
River, Salinas and Livramento, and the locality of the holotype is not specified. In Figure 2, where the holotype is
illustrated (based on the measurements provided), it is stated that the specimen was collected from Salinas at 2.8 m,
which we consider should therefore be considered as the type locality.
paranapuaensis, Yoldiella [Yoldiidae]. Benaim & Absalão, 2011b: 517, fig. 3A, B, E−H. Type locality: Bacia de
Campos, Station 41 (22°39′34 36′′S, 40°08′22 27′′W, 1200 m dep), Rio de Janeiro state, Brazil.
Holotype. MNRJ 15201 [valve]; paratype: MNRJ 15205 [1 valve], Bacia de Campos, Station 61 (21°525′1
90′′S, 39°48′11 68′′W, 1350 m dep), Rio de Janeiro state, Brazil.
sanmartini, Pleuromeris [Carditidae]. Klappenbach, 1970: 36, figs. 1−5. Type locality: La Paloma (Cabo de Santa
María), Departamento de Rocha, Uruguay.
Paratypes. MNRJ 18305 [5 valves], type locality (Fig. 37).
Remarks. The original publication listed paratypes in MNHN 1324 (currently MNHNM), with “almost a
hundred valves”; five of them were later donated to the MNRJ collection.
taludae, Paranadara [Arcidae]. Francisco, Barros & Lima, 2012: 1141, figs. 1−6. Type locality: Off the state of
Rio Grande do Norte, Brazil, ‘Natureza’ sta D−23 (04°51’S 35°06’W, 375 m dep).
Paratypes. MNRJ 11127 [1 v], off the state of Pernambuco, Brazil, ‘Natureza’ sta D-11 (08°46’S, 34°44’W,
690 m); MNRJ 11134 [3 v], type locality.
Remarks. Original publication mentioned that the holotype is in the lot MNRJ 11127, which indeed bears a
label, written by the authors as holotype; however, the valve in this lot does not correspond to the holotype’s figure
in the original publication and thus must be considered a paratype; this lot also bears a different location data than
that indicated in the original publication. The holotype itself was not sent to MNRJ collection.
tarcylae, Asperarca [Arcidae]. Francisco, Barros & Lima, 2012: 1148, figs. 25−30. Type locality: Off the state of
Alagoas, Brazil, ‘Natureza’ sta D-32 (09°20’S, 34°59’W, 452 m dep).
Paratypes. MNRJ 11126 [1 v], off the state of Bahia, Brazil, ‘Natureza’ sta D-30 (12°02’S, 37°36’W, 500 m
dep); paratypes: MNRJ 11133 [1 v], type locality.
Remarks. Original publication mentioned that the holotype is in the lot MNRJ 11126, which indeed bears a
label, written by the authors as holotype; however, the valve in this lot does not correspond to the holotype’s figure
in the original publication and thus must be considered a paratype; this lot also bears a different location data than
that indicated in the original publication. The holotype itself was not sent to MNRJ collection.
tamandua, Cuspidaria [Cuspidariidae]. Absalão & Oliveira, 2011: 133, figs. 57−63. Type locality: 22°11’04"S,
39°47’04"W, 1650m, Campos Basin, Rio de Janeiro state, Brazil.
Paratype. MNRJ 18729 [1 valve], Campos Basin (22°31’37"S, 39°55’14"W, 1650 m dep), Rio de Janeiro
state, Brazil.
tarasconii, Corbula (Caryocorbula) [Caryocorbula, Corbulidae]. Arruda, Domaneschi, Francisco & Barros, 2007:
202, figs. 1−18. Type locality: Off Guarapari, Espírito Santo state, Brazil (20°45’S, 40°25’, 60−65 m dep).
Paratypes. MNRJ 11146 [2 sh], northeast of Bahia state (15°58.7’S, 36°49.2’W, 100 m dep); MNRJ 11147 [2
sh], northeast of Bahia state (15°58.7’S, 36°49.2’W, 100 m dep); MNRJ 11148 [10 valves], northeast of Bahia state
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(15°58.7’S, 36°49.2’W, 100 m dep); MNRJ 11149 [10 valves], northeast of Bahia state (15°58.7’S, 36°49.2’W, 100
m dep); MNRJ 11150 [10 valves], northeast of Bahia state (15°58.7’S, 36°49.2’W, 100 m dep); MNRJ 11151 [10
valves], northeast of Bahia state (15°58.7’S, 36°49.2’W, 100 m dep); MNRJ 11152 [10 valves], northeast of Bahia
state (15°58.7’S, 36°49.2’W, 100 m dep); MNRJ 11153 [10 valves], northeast of Bahia state (15°58.7’S,
36°49.2’W, 100 m dep); MNRJ 11154 [10 valves], northeast of Bahia state (15°58.7’S, 36°49.2’W, 100 m dep);
MNRJ 11155 [10 valves], northeast of Bahia state (15°58.7’S, 36°49.2’W, 100 m dep); MNRJ 11156 [10 valves],
northeast of Bahia state (15°58.7’S, 36°49.2’W, 100 m dep); MNRJ 11157 [10 valves], northeast of Bahia state
(15°58.7’S, 36°49.2’W, 100 m dep); MNRJ 11040 [2 valves], southeast of Bahia state (15°53’82"S, 38°31’09", 66
m dep); MNRJ 11812 [1 sh], off Guarapari, Espírito Santo state, Brazil (60−70 m dep).
thryptica, Lima [Limaria, Limidae]. Penna, 1971: 156, fig. 3, est. 1. Type locality: Bay of Ilha Grande, Rio de
Janeiro state, Brazil; est. 72, 8 m dep.
Paratype. MNRJ 3546 [1 spm with somewhat broken valves], Bay of Ilha Grande, Rio de Janeiro state, Brazil
est. 82 (13 m dep).
Remarks. Published under the collection number MZUSP 18457; later donated to the MNRJ collection.
transmar, Acar [Arcidae]. Simone, 2009b: 10, figs. 1−26. Type locality: Belmonte Island (00°55’01"S,
29°20’44"W), São Pedro e São Paulo Island, Brazil.
Paratypes. MNRJ 18304 [2 spm], type locality.
Remarks. The original publication did not indicate the catalogue number.
Class Scaphopoda
elegantulum, Dentalium (Antalis) [Dentaliidae]. Penna-Neme 1974: 108, figs. 1, 3. Type locality: 10°41’02"S,
36°20’25"W, 100m, off Sergipe state, northeast Brazil.
Types not located.
Remarks. The original publication listed MNRJ 3646 as a paratype lot with four shells; this lot is missing.
eliezeri, Cadulus [Gadilidae]. Caetano, Scarabino & Absalão, 2006: 35, figs. 70−71. Type locality: Off Bahia state,
Brazil (15°53'49"S, 38°31'05"W, 66 m dep).
Paratypes. MNRJ 10571 [3 sh], type locality.
meridionalis, Heteroschismoides [Entalinidae]. Scarabino & Caetano 2008: 175. Type locality: South-western
Atlantic Ocean: MD 55 stn DC 70, 18°59’1S, 37°47’8W, 1540−1550 m.
Paratype. MNRJ 12707 [1 sh], type locality.
xavante, Polyschides [Gadilidae]. Caetano & Absalão, 2005: 3, figs. 1−4. Type locality: Baia do Sancho, Fernando
de Noronha, Pernambuco state, Brazil (03°51'S, 32°26'W, 10−15 m dep).
Paratypes. MNRJ 10313 [6 sh], type locality.
longa, Gadila [Gadilidae]. Filho, Pinto & Alves, 2010: 49, fig. 2. Type locality: northeastern Brazil, Rio Grande do
Norte state (04°51’02"S, 37°12’37"W, 206 m dep).
Paratypes. MNRJ 10482 [1 sh], type locality, (according to original description); Ceará state, northeastern
Brazil (04°14’91"S, 37°12’14"W, 177 m dep), (according to lable).
Remarks. The data about locality on the label of MNRJ 10482, written by the authors, is different from that
indicated on original description.
robusta, Gadila [Gadilidae]. Filho, Pinto & Alves, 2010: 51, fig. 3. Type locality: northeastern Brazil, Rio Grande
do Norte state (04°51’0"S, 35°06’46"W, 460 m dep).
Paratypes. MNRJ 10483 [1 sh], Rio Grande do Norte, (06°14’04"S, 34°52’33"W, 510 m dep).
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FIGURES 1–7. 1. Acmaea marcusi Righi, 1966; holotype MNRJ 3200; 1a, dorsal view; 1b, ventral view; length: 15.9 mm,
width: 12.4 mm. 2. Acmaea (Colisella) abrolhosensis Petuch, 1979; paratype MNRJ 3733; 2a, dorsal view; 2b, ventral view;
length: 16.5 mm, width: 13.9 mm. 3. Calliostoma (Emerlinia) bullisi Clench & Turner, 1960; paratype MNRJ 12725; 3a,
frontal view; 3b, apical view; 3c, basal view; length: 14.8 mm, width: 17.5 mm. 4. Arene flexispina Leal & Coelho, 1985;
holotype MNRJ 5352; 5a, frontal view; 5b, apical view; 5c, basal view; length: 6.3 mm, width: 6.0 mm. 5. Tricolia affinis
pterocladica Robertson, 1958; paratype MNRJ HSL5912; 5a, frontal view; 5b, dorsal view; length: 5.5 mm, width: 3.5 mm. 6.
Ampullaria meta Ihering, 1915; holotype MNRJ 12861; 6a, frontal view; 6b, apical view; 6c, basal view; length: 28.0 mm,
width: 29.3 mm. 7. Pomacea cumingii sanjosensis Morrison, 1946; paratype MNRJ 3488; 7a, frontal view; 7b, apical view; 7c.
basal view; length: 43.4 mm, width: 42.4 mm.
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FIGURES 8–16. 8. Thericium biminiense Pilsbry & McGinty, 1949; paratype MNRJ HSL6151; length: 9.8 mm, width: 5.5
mm. 9. Petalochonchus myrakeenae Absalão & Rios, 1987; paratypes MNRJ 5820. 10. Malea noronhensis Kempf &
Matthews, 1969; holotype MNRJ 3493; 10a, frontal view; 10b, apical view; length: 69.6 mm, width: 51.0 mm. 11. Bursa
(Colubrellina) natalensis Coelho & Matthews, 1970; holotype MNRJ 3527; 11a, frontal view; 11b, dorsal view; length: 78.5
mm, width: 32.5 mm. 12. Bursa pacamoni Matthews & Coelho, 1971; holotype MNRJ 3528; 12a, frontal view; 12b, dorsal
view; length: 28.0 mm, width: 19.1 mm; 12c, operculum. 13. Bursa barcellosi Matthews, Rios & Coelho, 1973; paratype
MNRJ 3647; 13a, frontal view; 13b, dorsal view; length: 148.2 mm, width: 75.5 mm. 14. Epitonium mauryi Tursch & Pierret,
1964; paratype MNRJ 3117; 15a, frontal view; 15b, dorsal view; length: 13.5 mm, width: 5.2 mm. 15. Epitonium arnaldoi
Tursch & Pierret, 1964; paratype MNRJ 3118; 14a, frontal view; 14b, dorsal view; length: 5.8 mm, width: 2.8 mm. 16. Eulima
mulata Rios & Absalão, 1990; paratype MNRJ 5802; length: 15.2 mm, width: 3.1 mm.
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FIGURES 17–28. 17. Typhina riosi Bertsch & D’Áttilio, 1980; paratypes MNRJ 4863; 17a, frontal view; 17b, dorsal view;
17c, apical view; length: 11.1 mm, width: 8.0 mm. 18. Latirus frenguelli Carceles, 1953; paratype MNRJ 13204, 18a, frontal
view; 18b, dorsal view; length: 37.8 mm, width: 15.6 mm. 19. Latirus lacteum Matthews-Cascon, Matthews & Rocha, 1991;
holotype MNRJ 6201; 19a, frontal view; 19b. dorsal view; length: 27.0 mm, width: 10.7 mm. 20. Mitrella albovittata Lopes,
Coelho & Cardoso, 1965; holotype MNRJ HSL6502; 20a, frontal view; 20b, dorsal view; length: 2.9 mm, width: 1.5 mm. 21.
Anachis helenae Costa, 1983; paratype MNRJ 4864; 21a, frontal view; 21b, dorsal view; length: 7.0 mm, width: 3.1 mm. 22.
Pyrene (Conella) ledaluciae Rios & Tostes, 1981; paratypes: MNRJ 3900; 21a, frontal view; 21b, dorsal view; length: 25.1
mm, width: 11.0 mm. 23. Ancilla faustoi Matthews, Matthews & Dijck, 1977; holotype MNRJ 3790; 23a, frontal view; 23b,
dorsal view; length: 15.0 mm, width: 6.7 mm. 24. Olivella (Orbignytesta) formicacorsii Klappenbach, 1962; paratype MNRJ
2909; 24a, frontal view; 24b, dorsal view; length: 11.3 mm, width: 4.0 mm. 25. Mitra (Cancilla) saldanha Matthews & Rios,
1970; paratype MNRJ 3503; 25a, frontal view; 25b, dorsal view; length: 13.6 mm, width: 4.4 mm. 26. Conus carcellesi
Martins, 1945; holotype MNRJ 8462; 26a, frontal view; 26b, dorsal view; 26c, apical view; length: 28.9 mm, width: 14.8 mm.
27. Conus carioca Petuch, 1986; paratypes MNRJ 8459; 27a, frontal view; 27b, dorsal view; 27c, apical view; length: 53.0
mm, width: 28.3 mm. 28. Conus clenchi Martins, 1943; holotype MNRJ 11720; 28a, frontal view; 28b, dorsal view; 28c, apical
view; length: 36.0 mm, width: 19.2 mm.
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FIGURES 29–37. 29. Kerguelenella flemingi Powell, 1955; paratype MNRJ HSL4069; 29a, dorsal view; 29b, ventral view;
length: 21.7 mm, width: 16.2 mm. 30. Succinea lopesi Lanzieri, 1966; paratype MNRJ 3176; 30a, frontal view; 30b, dorsal
view; length: 11.2 mm, width: 6.0 mm. 31. Oxyloma beckeri Lanzieri, 1966; paratype MNRJ 3177; 31a, frontal view; 31b,
dorsal view; length: 7.9 mm, width: 4.9 mm. 32. Corona ribeiroi Ihering, 1915; holotype MNRJ 3444 ; 32a, frontal view; 32b,
dorsal view; length: 47.1 mm, width: 22.6 mm. 33. Bulimulus trindadensis Breure & Coelho, 1976; holotype MNRJ 3711; 33a,
frontal view; 33b, dorsal view; length: 7.5 mm, width: 3.2 mm. 34. Loboa brunoi Ihering, 1917; holotype MNRJ 1128; 34a,
frontal view; 34b, dorsal view; length: 19.4 mm, width: 7.5 mm. 35. Protoglyptus arnaldoi [Naesiotus] Lanzieri & Rezende,
1971; holotype MNRJ 3105; 35a, frontal view; 35b, dorsal view; length: 9.5 mm, width: 4.7 mm. 36. Gaimardia trapesina
flemingi Powell, 1955; paratype MNRJ HSL4060; length: 25.8 mm, heigth: 17.3 mm. 37. Pleuromeris sanmartini
Klappenbach, 1970; paratype MNRJ 18305; length: 2.4 mm, heigth: 2.5 mm.
Class Polyplacophora
terezae, Acanthochitona [Polyplacophora, Acanthochitonidae]. Guerra, 1983: 385, figs. 1−9. Type locality: coast
of Bahia state, close to Farol de Itapoã, Salvador.
Types not located.
Remarks. The original publication stated that the holotype and 17 paratypes were deposited in, respectively,
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the lots Col. Mol. M.N. 4584 and Col. Mol. M.N. 4585. However, these two lots were not located, and were
probably never received by the MNRJ Mollusca Collection.
Doubtful types or specimens erroneously published as types stored in the MNRJ Mollusca Collection
Some specimens published elsewhere as types belonging to the MNRJ were not confirmed. Below is a list of the
specimens that were erroneously published as types:
cajamarcana, Steeriana (Steeriana) [Clausiliidae]. Weyrauch & Zilch in Zilch, 1954: 73, taf. 5, fig. 8, Abb. 6.
Type locality: 18 km from cajamarca on the road to San marcos, Hacienda Cochambul, 2800 m alt), Peru,
Departamento Cajamarca.
Remarks. The lot MNRJ HSL1925, from the type locality, bears a label giving the information “leg. W.
Weyrauch”. This lot was cited in Barbosa et al. (2008) as MNJR HSL−1925 (sic) with doubts about its type status.
We believe that there is no enough evidence to consider it as a paratype lot.
celendinensis, Steeriana (Steeriana) [Clausiliidae]. Weyrauch & Zilch in Zilch, 1954: 70, Taf. 5, fig. 6; Abb. 3.
Type locality: km 113 road (Cajamarca) Balsas, 3 km SE celendin (06°53’12"S, 78°07’40"W, 2900 m alt), Peru,
Departamento Cajamarca.
Remarks. The paratype MNRJ HSL1263, from type locality, has a label providing the information “leg. W.
Weyrauch”. Barbosa et al. (2008) assumed they are paratypes. However, although the label of MNRJ HSL 1263
indicates that they are paratypes, there is no information on the label that can be related to a lot mentioned in the
original publication.
cardosoi, Strophocheilus (Megalobulimus) [Megalobulimidae]. Lange-de-Morretes, 1952: 119, Estampas III, IV,
fig. 3). Type locality: Muricí, Alagoas state, Brazil.
Remarks. Simone (2006a: 208, fig. 788B) figured a shell, indicated as “paratype MNRJ 2060”. However, the
collection number 2060 in the MNRJ catalogue does not consist of this taxon. The number 2060 is written in ink on
the inner wall of the shell, and corresponds to the malacological collection of Museo Nacional de Historia Natural
y Antropologia, Montevideo (MNHNM). Examination of this lot revealed three shells and a label reading “4
paratypes”, one of which was donated to the MNRJ collection.
Original publication mentioned five paratypes; all of them are currently deposited at MZSP malacological
collection (Dornellas & Simone, 2011).
nordestensis, Tropicorbis [Planorbidae]. Lucena, 1954: 330, fig. 1.
Remarks. Simone (2006) illustrated a shell from MNRJ, indicating that it corresponds to the type of T.
nordestensis. However, such shell was not found in the MNRJ collection and the original publication (Lucena,
1954) does not mention type deposited in the MNRJ.
pimentai, Gregorioiscala [Epitoniidae]. Lima, Christoffersen, Barros & Folly, 2012: 9, figs. 10a–d.
Remarks. The paper in which this name was published does not constitute a published work for nomenclatural
purposes according to amendment of Article 8.5 of the International Code of Zoological Nomenclature (2012). The
work was issued and distributed electronically only, and does not fulfil the requirements of stating the date of
publication in the work itself and neither the taxon nor the publication has been registered in the Official Register
of Zoological Nomenclature (ZooBank).
Although the name will be formally published in the future, with the same type material (Lima pers. com.), for
the moment it should be considered not available, and thus, the material cited under the number MNRJ 18306 as
holotype is not a type.
renatae, Anachis [Columbellidae, not available name]. Rios, 2009: 238, fig, 593.
Remarks. Rios (2009: 238) presented the description of a supposed new species of Columbellidae mentioning
that “…the holotype at National Museum” (sic). Rios (2009) was probably referring to the MNRJ collection, but
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PIMENTA ET AL.
since he did not explicitly refer to it, and “National Museum” could mean other museums in the world, its
establishment is not in accordance with the ICZN (1999: 16.4.2). This article requires that a new species name
published after 1999, to be available, must be accompanied by a statement indicating the name and location of the
collection where the holotype is deposited. Furthermore, the supposed holotype was never sent by the author to the
MNRJ collection.
revizee, Opalia [Epitoniidae]. Lima, Christoffersen, Barros & Folly, 2012: 8, figs. 9a–e.
Remarks. The paper in which this name was published does not constitute a published work for nomenclatural
purposes according to amendment of Article 8.5 of the International Code of Zoological Nomenclature (2012). The
work was issued and distributed electronically only, and does not fulfil the requirements of stating the date of
publication in the work itself and neither the taxon nor the publication has been registered in the Official Register
of Zoological Nomenclature (ZooBank).
Although the name will be correctly published in the future, with the same type material (Lima pers. com.), for
the moment it should be considered not available, and thus, the material cited under the number MNRJ 18307 as
holotype is not a type.
tarmensis, Naesiotus (Reclasta) [Bostryx, Orthalicidae]. Weyrauch, 1967b: 479, figs. 51−52. Type locality: Valley
Río Tarma, Cerro Huayuncayo, near La Florida (11°22’13"S, 75°40’30"W, 3000−3100 m alt), Peru, Departamento
Lima.
Remarks. MNRJ 4418, from type locality, bears a label indicating IML 12227, the abbreviation used for the
Instituto Miguel Lillo, where types for other names described by Weyrauch (1967b) were deposited. However, the
original publication did not list IML 12227 as a paratype lot. Although Barbosa et al. (2008) assumed MNRJ 4418
as paratypes, we consider that they are only topotypes.
Acknowledgments
We are grateful to Claudio Costa, for helping managing the molluscan type collection, Dr. P. M. S. Costa and Dr. C.
D. de Oliveira, for their help taking the photographs; Dr. L. R. Simone, Dr. C. M. Cunha, F. Scarabino, Dr. G.
Rosenberg, for exchanging of information about the type status of specimens, Dr. J. Reid, for revising the English
text, and Dr. Rüdiger Bieler, Dr. Frank Köhler and Dr. Alexander Nützel for their valuable critics and sugestions.
This work was partially supported by "Fundação Carlos Chagas de Amparo à Pesquisa do Estado do Rio de
Janeiro" (Faperj), processo APQ4 E-26/110.068/2014.
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APPENDIX. List of names of molluscan types deposited in the MNRJ collection.
Class Gastropoda
Lottiidae
Lottia abrolhosensis (Petuch, 1979)
Lottia marcusi (Righi, 1966)
Fissurellidae
Lucapina elisae Costa & Simone, 2006
Fissurella mesoatlantica Simone, 2008
Pseudococculinidae
Pseudococculina rimula Simone & Cunha, 2003
Scissurellidae
Sinezona insularis Simone, 2009
Seguenzioidea (unassigned to family)
Brookula megaumbilicata Absalão & Pimenta, 2005
Brookula olearia Absalão & Pimenta, 2005
Brookula proseila Absalão & Pimenta, 2005
Brookula spinulata Absalão, Miyaji & Pimenta, 2001
Granigyra oblatogyra de Souza & Pimenta, 2002
Vetulonia parajeffreysi Absalão & Pimenta, 2005
Seguenziidae
Ancistrobasis zumbii Lima, Christoffersen & Barros, 2013
Calliostomatidae
Calliostoma bullisi Clench & Turner, 1960
Calliostoma tupinamba Dornellas, 2012
Trochidae
Synaptocochlea belmonti Simone, 2009
Solariellidae
Solariella quinni Barros & Pereira, 2008
Margaritidae
Gaza compta Simone & Cunha, 2006
Areneidae
Arene flexispina Leal & Coelho, 1985
Phasianellidae
Eulithidium pterocladicum Robertson, 1958
Abyssochrysidae
Abyssochrysos brasilianus Bouchet, 1991
Ampullariidae
Pomacea meta (Ihering, 1915)
Pomacea cumingii sanjosensis Morrison, 1946
Cerithiidae
Cerithium biminiense (Pilsbry & McGinty, 1949)
Barleeidae
Caelatura albertoi Santos & Absalão, 2006
Caelatura aulakion Santos & Absalão, 2006
Caelatura barcellosi Absalão & Rios, 1995
Caelatura carinata Santos & Absalão, 2006
Caelatura noxia Santos & Absalão, 2006
Caelatura phrix Santos & Absalão, 2006
Caelatura speculabunda Absalão, 2002
Caelatura spirocordata Absalão & Rios, 1995
Caelatura tigrina Absalão, 2002
Caelatura tupi Santos & Absalão, 2006
Ketosia riosi Santos & Absalão, 2006
Ketosia thomei Santos & Absalão, 2006
Protobarleeia pyrrocincta Absalão, 2002
Pseudodiala puncturina Santos & Absalão, 2006
Rissoidae
Alvania valeriae Absalão, 1993
Benthonellania coronata Absalão & Santos, 2004
Benthonellania multicostata Absalão & Santos, 2004
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Rissoina indiscreta Leal & Moore, 1989
Anabathridae
Pisinna bicincta Santos & Absalão, 2004
Caecidae
Caecum eliezeri Absalão, 1997
Caecum massambabense Absalão, 1994
Caecum metamorphosicum Lima, Santos & Absalão, 2013
Caecum trindadense Lima, Santos & Absalão, 2013
Pomatiopsidae
Spiripockia puncata Simone, 2012
Tornidae
Episcynia itanhura Simone, 2012
Hydrobiidae
Pothamolithus karsticus Simone & Morachioli, 1994
Pothamolithus troglobius Simone & Morachioli, 1994
Calyptraeidae
Crepidula intratesta Simone, 2006
Crepidula carioca Simone, 2006
Vermetidae
Petaloconchus myrakeenae Absalao & Rios, 1987
Thylaeodus equatorialis Spotorno & Simone, 2013
Naticidae
Natica juani Costa & Pastorino, 2012
Notocochlis laurae Costa & Pastorino, 2012
Tonnidae
Malea noronhensis Kempf & Matthews, 1969
Bursidae
Bursa natalensis Coelho & Matthews, 1970
Bursa pacamoni Matthews & Coelho, 1971
Ranellidae
Ranella barcellosi (Matthews, Rios & Coelho, 1973)
Epitoniidae
Epitonium arnaldoi Tursch & Pierret, 1964
Epitonium mauryi Tursch & Pierret, 1964
Nystiellidae
Opaliopsis cearensis Andrade, Costa & Pimenta, 2011
Eulimidae
Annulobalcis aurisflamma Simone & Martins, 1995
Eulima mulata Rios & Absalao, 1990
Triphoridae
Isotriphora onca Fernandes, Pimenta & Leal, 2013
Isotriphora tigrina Fernandes, Pimenta & Leal, 2013
Metaxia gongyloskymnus Fernandes & Pimenta, 2011
Cerithiopsidae
Cerithiopsis balaustium Figueira & Pimenta, 2008
Cerithiopsis capixaba Figueira & Pimenta, 2008
Newtoniellidae
Cerithiella cepene Lima & Barros, 2007
Cerithiella pernambucoensis Lima & Barros, 2007
Muricidae
Brasityphis barrosi Absalao & Santos, 2003
Leptotrophon atlanticus Pimenta, Couto & Costa, 2008
Pazinotus bodarti (Costa, 1993)
Pazinotus gili Costa & Pimenta, 2012
Siphonochelus riosi (Bertsch & D'Attilio, 1980)
Buccinidae
Monostiolum atlanticum (Coelho, Matthews & Cardoso, 1970)
Anna goncalvesi (Coltro, 2005)
Engina janowskyi Coltro, 2005
Metula anfractura Matthews & Rios, 1968
Metula gigliottii Coltro, 2005
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Pisania bernardoi Costa & Gomes, 1998
Fasciolariidae
Fasciolaria tephrina de Souza, 2002
Fusinus frenguellii (Carcelles, 1953)
Hemipolygona beckyae (Snyder, 2000)
Latirus devyanae Rios, Costa & Calvo, 1994
Polygona lactea (Matthews-Cascon, Matthews & Rocha, 1991)
Columbellidae
Anachis carloslirai Costa, 1996
Astyris antares (Costa & de Souza, 2001)
Cosmioconcha helenae (Costa, 1983)
Conella ledaluciae (Rios & Tostes, 1981)
Mitrella cabofrioensis Costa & de Souza, 2001
Mokumea albovittata (Lopes, Coelho & Cardoso, 1965)
Nassarina thetys Costa & Absalao, 1998
Marginellidae
Bullata analuciae de Souza & Coovert, 2001
Bullata guerrinii de Souza & Coovert, 2001
Marginella cloveri Rios & Matthews, 1972
Prunum amphorale de Souza, 1992
Volvarina brasiliana Boyer, 2000
Volvarina pontesi Rios & Leal, 1993
Volvarina tunicata Boyer, 2000
Mitridae
Subcancilla lopesi (Matthews & Coelho, 1969)
Mitra saldanha (Matthews & Rios, 1970)
Costellariidae
Vexilum moisei (McGinty, 1955)
Nodicostellaria crassa (Simone, 1995)
Nodicostellaria kaicherae (Petuch, 1979)
Olividae
Ancilla faustoi Matthews, Matthews & Muniz Dijck, 1979
Olivancillaria buckuporum Thomé, 1966
Olivancillaria teaguei Klappenbach, 1964
Olivancillaria vesica (Gmelin, 1791)
Olivellidae
Olivella arionata Absalao, 2000
Olivella careorugula Absalao & Pimenta, 2003
Olivella formicacorsii Klappenbach, 1962
Olivella hyphala Absalao & Pimenta, 2003
Olivella klappenbachi Absalao & Pimenta, 2003
Volutidae
Adelomelon indigestus Ihering, 1908
Nanomelon viperinus Leal & Bouchet, 1989
Nanomelon vossi Leal & Rios, 1990
Plicoliva zelindae (Petuch, 1979)
Conidae
Conus baiano Coltro, 2004
Conus bertarollae Costa & Simone, 1997
Conus bodarti Coltro, 2004
Conus carcellesi Martins, 1945
Conus cargilei Coltro, 2004
Conus carioca Coltro, 2004
Conus clenchi Martins, 1943
Conus delucai Coltro, 2004
Conus henckesi Coltro, 2004
Conus iansa Petuch, 1979
Conus mauricioi Coltro, 2004
Conus pseudocardinalis Coltro, 2004
Conus riosi Petuch, 1986
Conus schirrmeisteri Coltro, 2004
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Conus tostesi Petuch, 1986
Terebridae
Terebra crassireticula Simone, 1999
Terebra simonei Lima, Tenório & Barros, 2007
Terebra spirosulcata Simone & Costa, 1999
Borsoniidae
Diptychophlia hubrechti Cunha, 2005
Clathurellidae
Corinnaeturris angularis Figueira & Absalão, 2010
Cochlespiridae
Cochlespira elongata Simone, 1999
Drilliidae
Fusiturricula maese Rios, 1985
Mangeliidae
Benthomangelia enceladus Figueira & Absalão, 2010
Oenopota carioca Figueira & Absalão, 2010
Oenopota seraphina Figueira & Absalão, 2010
Oenopota diabula Figueira & Absalão, 2010
Mitromorphidae
Mitromorpha santosi Lima, Barros & Francisco, 2010
Raphitomidae
Neopleurotomoides aembe Figueira & Absalão, 2012
Taranis tanata Figueira & Absalão, 2010
Xanthodaphne pichi Figueira & Absalão, 2012
Gymnobela xaioca Figueira & Absalão, 2012
Pleurotomella ybessa Figueira & Absalão, 2012
Cancellariidae
Brocchinia harasewychi Barros & Petit, 2007
Brocchinia verheckeni Barros & Petit, 2007
Gerdiella alvesi Lima, Barros & Petit, 2007
Gergovia petiti Barros & Petit, 2007
Microcancilla jonasi Barros & Petit, 2007
Pyramidellidae
Boonea scymnocelata Pimenta, Absalão & Miyaji, 2009
Chrysallida conifera Pimenta, 2012
Egila ektopa Pimenta & Absalão, 2004
Eulimastoma exiguum Pimenta, 2012
Eulimastoma franklini Pimenta, 2012
Eulimella cylindrata Pimenta, Santos & Absalão, 2011
Eulimella ejuncida Pimenta, Santos & Absalão, 2011
Eulimella torquata Pimenta, Santos & Absalão, 2011
Odostomella carceralis Pimenta, Absalão, & Alencar, 2000
Oscilla aquilonia Pimenta, Santos & Absalão, 2008
Oscilla notialis Pimenta, Santos & Absalão, 2008
Parthenina biumbilicata Pimenta, 2012
Peristichia lepta Pimenta, Santos & Absalão, 2008
Triptychus litosbathron Pimenta, Santos & Absalão, 2008
Turbonilla aquilonaria Absalão, Santos & Tenório, 2003
Turbonilla aracruzensis Pimenta & Absalão, 2004
Turbonilla capixaba Pimenta & Absalão, 2004
Turbonilla farinatiae Pimenta & Absalão, 2004
Turbonilla farroupilha Pimenta & Absalão, 2004
Turbonilla fluminensis Pimenta & Absalão, 2004
Turbonilla goytacazi Pimenta & Absalão, 2004
Turbonilla kaapor Pimenta & Absalão, 2004
Turbonilla kamayura Absalão, Santos & Tenório, 2003
Turbonilla lepta Pimenta & Absalão, 2004
Turbonilla macaensis Pimenta & Absalão, 2001
Turbonilla maestratii Pimenta & Absalão, 2004
Turbonilla mermeroglaphyra Absalão, Santos & Tenório, 2003
Turbonilla midas Pimenta & Absalão, 2004
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Turbonilla parviscymna Pimenta & Absalão, 2004
Turbonilla paulinoi Pimenta & Absalão, 2004
Turbonilla rhachialis Pimenta & Absalão, 2004
Turbonilla scapulata Pimenta & Absalão, 2004
Turbonilla tupinamba Pimenta & Absalão, 2002
Turbonilla uaca Pimenta & Absalão, 2004
Turbonilla virgulinoi Absalão, Santos & Tenório, 2003
Architectonicidae
Heliacus willianseverii Tenório, Barros, Francisco & Silva, 2011
Pseudotorinia jonasi Tenório, Barros, Francisco & Silva, 2011
Psilaxis clertoni Tenório, Barros, Francisco & Silva, 2011
Solatisonax cabrali Tenório, Barros, Francisco & Silva, 2011
Solatisonax rudigerbieleri Tenório, Barros, Francisco & Silva, 2011
Rissoellidae
Rissoella ornata Simone, 1995
Acteonidae
Acteon mirim Cunha, 2011
Crenilabium birmani Simone, 2006
Onchidorididae
Onchidoris brasiliensis Alvim, Padula & Pimenta, 2011
Discodorididae
Jorunna spongiosa Alvim & Pimenta, 2013
Taringa iemanja Alvim & Pimenta, 2013
Chromodorididae
Felimare juliae (Dacosta, Padula & Schrödl, 2010)
Aeolididae
Cerberilla potiguara Padula & Delgado, 2010
Siiphonariidae
Kerguelenella flemingi Powell, 1955
Planorbidae
Biomphalaria inflexa (Paraense & Deslandes, 1956)
Biomphalaria intermedia (Paraense Deslandes, 1962)
Physidae
Physa papaveroi Leme, 1966
Succineidae
Oxyloma beckeri Lanzieri, 1966
Succinea lopesi Lanzieri, 1966
Charopidae
Stephadiscus madrediosensis Miquel & Ramírez, 2011
Orthalicidae
Anostoma tessa Simone, 2012
Bostryx coelhoi Breure, 1978
Bostryx haasi minor Weyrauch, 1960
Bostryx imeldae costifer Weyrauch, 1960
Bostryx rehderi Weyrauch, 1960
Bostryx rodriguezae Weyrauch, 1967
Bostryx vilchezi Weyrauch, 1960
Bostryx zilchi glomeratus Weyrauch, 1960
Bulimulus brunoi (Iheringi, 1917)
Corona ribeiroi Iheringi, 1915
Drymaeus ribeiroi Iheringi, 1915
Kora corallina Simone, 2012
Naesiotus arnaldoi (Lanzieri & Rezende, 1971)
Naesiotus carlucioi (Rezende & Lanzieri, 1963)
Naesiotus lopesi (Rezende, Lanzieri & Inada, 1972)
Scutalus coraeformis debilisculptus Weyrauch, 1967
Scutalus chiletensis Weyrauch, 1967
Scutalus grandiventris Weyrauch, 1960
Spixia coltrorum Simone, 2012
Peltella iheringi Leme, 1968
Thaumastus baixoguanduensis Pena, Salgado & Coelho, 1996
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Thaumastus caetensis Pena, Salgado & Coelho, 2011
Thaumastus lundi Pena, Salgado & Coelho, 2005
Thaumastus parvus Pena, Salgado & Coelho, 2011
Thaumastus straubei Colley, 2012
Thaumastus tiradentensis Pena, Salgado & Coelho, 1996
Tomigerus esamianus Salgado & Coelho, 1990
Tomigerus matthewsi Salgado & Leme, 1991
Vegrandinia trindadensis (Breure & Coelho, 1976)
Strophocheilidae
Megalobulimus amandus Simone, 2012
Megalobulimus klappenbachi Leme, 1964
Gonyostomus insularis Leme, 1974
Clausiliidae
Temesa omissa Weyrauch, 1957
Zilchiella grandiportus Weyrauch, 1957
Class Cephalopoda
Sepiidae
Sepia jurujubai (Oliveira, 1940)
Octopodidae
Eledone gaucha Haimovich, 1988
Octopus insularis Leite & Haimovici in Leite, Haimovici, Molina & Warnke, 2008
Class Bivalvia
Yoldiidae
Microgloma macaron Benaim & Absalão, 2011
Microgloma nhanduti Benaim & Absalão, 2011
Yoldiella arariboia Benaim & Absalão, 2011
Yoldiella curupira Benaim & Absalão, 2011
Yoldiella lapernoi Benaim & Absalão, 2011
Yoldiella paranapuaensis Benaim & Absalão, 2011
Arcidae
Acar lepidoformis Francisco, Barros & Lima, 2012
Acar naturezae Francisco, Barros & Lima, 2012
Acar oliveirae Francisco, Barros & Lima, 2012
Acar transmar Simone, 2009
Asperarca tarcylae Francisco, Barros & Lima, 2012
Paranadara taludae Francisco, Barros & Lima, 2012
Carditidae
Cardites micellus (Penna-Neme, 1971)
Pleuromeris sanmartini Klappenbach, 1970
Chamidae
Arcinella brasiliana (Nicol, 1953)
Condylocardiidae
Warrana besnardi (Klappenbach, 1963)
Limidae
Limaria thryptica (Penna, 1971)
Ostreidae
Crassostrea paraibanensis Singarajah, 1980
Semelidae
Semele aurora Tursch & Pierret, 1964
Dreissenidae
Mytilopsis lopesi Alvarenga & Ricci, 1989
Hyriidae
Triplodon chodo Mansur & Pimpão, 2008
Mycetopodidae
Fossula balzani mattogrosensis Ihering, 1915
Mactridae
Mactra guidoi Signorelli & Scarabino, 2010
Corbulidae
Caryocorbula tarasconii (Arruda, Domaneschi, Francisco & Barros, 2007)
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Lyonsiidae
Lyonsia celeste Pimenta & Oliveira, 2013
Verticordiidae
Verticordia ouricuri Oliveira & Absalão, 2010
Cuspidariidae
Myonera kaiwa Oliveira & Absalão, 2009
Cuspidaria tamandua Absalão & Oliveira, 2011
Protocuspidariidae
Protocuspidaria jarauara Oliveira & Absalão, 2009
Cyamiidae
Gaimardia trapesina flemingi Powell, 1955
Class Scaphopoda
Gadilidae
Cadulus eliezeriCaetano, Scarabino & Absalão, 2006
Gadila longa Filho, Pinto & Alves, 2010
Gadila robusta Filho, Pinto & Alves, 2010
Polyschides xavante Caetano & Absalão, 2005
Entalinidae
Heteroschismoides meridionalis Scarabino & Caetano 2008
MOLLUSCAN TYPES FROM BRAZIL
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Zootaxa 3780 (1): 051–107
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ISSN 1175-5326 (print edition)
Article
Copyright © 2014 Magnolia Press
ZOOTAXA
ISSN 1175-5334 (online edition)
http://dx.doi.org/10.11646/zootaxa.3780.1.3
http://zoobank.org/urn:lsid:zoobank.org:pub:74DE381B-0185-4735-8CCB-60AD25E0DA01
Catalogue of the type specimens deposited in the Mollusca Collection of the
Museu Nacional / UFRJ, Rio de Janeiro, Brazil
ALEXANDRE DIAS PIMENTA1,4, JÚLIO CÉSAR MONTEIRO2, ANDRÉ FAVARETTO BARBOSA3,
NORMA CAMPOS SALGADO1 & ARNALDO CAMPOS DOS SANTOS COELHO1
1
Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro (UFRJ)
Instituto de Estudos do Mar “Almirante Paulo Moreira” (IEAPM)
3
Instituto Brasileiro do Meio Ambiente e dos Recursos Naturais Renováveis (IBAMA)—Coordenação-Geral de Petróleo e Gás, Rio de
Janeiro
4
Correspondent author. E-mail: alexpim@mn.ufrj.br
2
Abstract
A curatorial revision of the type specimens deposited in the Mollusca Collection of the Museu Nacional / UFRJ, Rio de
Janeiro, Brazil (MNRJ) revealed the existence of 518 lots of type specimens (holotypes, neotypes, syntypes and paratypes)
for 285 names of molluscan taxa from 88 families, including 247 gastropods, 30 bivalves, three cephalopods and five scaphopods. A total of 106 holotypes and one neotype are deposited in the MNRJ. Type material for ten nominal taxa described as being deposited in the MNRJ was not located; the probable reasons are discussed. Some previously published
erroneous information about types in the MNRJ is rectified. A total of 37 type specimens are illustrated.
Key words: types, malacological collection, mollusc, malacology, nomenclature
Introduction
Founded in 1818 by D. João of Portugal as the “Museu Real”, the Museu Nacional is the oldest museum in Brazil
and also its oldest scientific institution (Museu Nacional, 2007). Since 1946, the museum has been part of the
Federal University of Rio de Janeiro (Universidade Federal do Rio de Janeiro—UFRJ) and it has academic,
scientific and educational missions, aiming to develop and disseminate knowledge of natural history and
anthropology. As part of its scientific activities, the Museu Nacional holds the most important zoological
collections in Brazil.
Following a recommendation of the International Code of Zoological Nomenclature (ICZN, 1999: 72F.4) to
encourage the publication of lists of types housed in institutions, this paper presents a catalogue of the molluscan
types held in the Mollusca Collection of the Museu Nacional.
The Mollusca Collection of the Museu Nacional
The Mollusca Collection of the Museu Nacional (acronym MNRJ), one of the most important of its kind in South
America, holds more than 32,000 registered lots and about 10,000 unregistered lots, which are currently being
processed, including dry shells and preserved specimens from Brazilian and foreign provenance. The type material
stems from studies carried out since the end of the 19th century by Brazilian and foreign researchers.
The early beginnings of this collection are difficult to establish. The earliest reference was by Netto (1870),
who mentioned that molluscs were exhibited in the first building of the Museu Nacional; followed by Lacerda
(1905), who stated that these specimens were exhibited at the present location of the museum. Miranda-Ribeiro
(1924) referred to the mollusc collection, which at that time contained 584 Brazilian species, as determined by E.
von Martens from “Zoologisches Museum”, presently “Museum für Naturkunde”, in Berlin.
Accepted by A. Nutzel: 7 Feb. 2014; published: 20 Mar. 2014
51
since he did not explicitly refer to it, and “National Museum” could mean other museums in the world, its
establishment is not in accordance with the ICZN (1999: 16.4.2). This article requires that a new species name
published after 1999, to be available, must be accompanied by a statement indicating the name and location of the
collection where the holotype is deposited. Furthermore, the supposed holotype was never sent by the author to the
MNRJ collection.
revizee, Opalia [Epitoniidae]. Lima, Christoffersen, Barros & Folly, 2012: 8, figs. 9a–e.
Remarks. The paper in which this name was published does not constitute a published work for nomenclatural
purposes according to amendment of Article 8.5 of the International Code of Zoological Nomenclature (2012). The
work was issued and distributed electronically only, and does not fulfil the requirements of stating the date of
publication in the work itself and neither the taxon nor the publication has been registered in the Official Register
of Zoological Nomenclature (ZooBank).
Although the name will be correctly published in the future, with the same type material (Lima pers. com.), for
the moment it should be considered not available, and thus, the material cited under the number MNRJ 18307 as
holotype is not a type.
tarmensis, Naesiotus (Reclasta) [Bostryx, Orthalicidae]. Weyrauch, 1967b: 479, figs. 51−52. Type locality: Valley
Río Tarma, Cerro Huayuncayo, near La Florida (11°22’13"S, 75°40’30"W, 3000−3100 m alt), Peru, Departamento
Lima.
Remarks. MNRJ 4418, from type locality, bears a label indicating IML 12227, the abbreviation used for the
Instituto Miguel Lillo, where types for other names described by Weyrauch (1967b) were deposited. However, the
original publication did not list IML 12227 as a paratype lot. Although Barbosa et al. (2008) assumed MNRJ 4418
as paratypes, we consider that they are only topotypes.
Acknowledgments
We are grateful to Claudio Costa, for helping managing the molluscan type collection, Dr. P. M. S. Costa and Dr. C.
D. de Oliveira, for their help taking the photographs; Dr. L. R. Simone, Dr. C. M. Cunha, F. Scarabino, Dr. G.
Rosenberg, for exchanging of information about the type status of specimens, Dr. J. Reid, for revising the English
text, and Dr. Rüdiger Bieler, Dr. Frank Köhler and Dr. Alexander Nützel for their valuable critics and sugestions.
This work was partially supported by "Fundação Carlos Chagas de Amparo à Pesquisa do Estado do Rio de
Janeiro" (Faperj), processo APQ4 E-26/110.068/2014.
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