HAWAIIAN MARINE MOLLUSCS

- an update to Kay, 1979

compiled by
Fabio Moretzsohn
and E. Alison Kay

(unpublished manuscript distributed at the

61st American Malacological Union Congress,
held at the University of Hawai‘i at Hilo, Hawai‘i)

University of Hawai‘i at Manoa

Honolulu, Hawai‘i
June 1995
 Hawaiian Marine Molluscs 1

Hawaiian Marine Molluscs - an update to Kay, 1979

Fabio Moretzsohn1 and E. Alison Kay2

1 Graduate student, Department of Zoology, University of Hawai’i at Manoa, 2538 The Mall, Honolulu,
Hawaii, 96822, USA
2 Professor of Zoology, Department of Zoology, UHM

Class GASTROPODA Nerita turrita Chemnitz

Subclass PROSOBRANCHIA Neritilia hawaiiensis Kay, 1979
ORDER DOCOGLOSSA Neritina granosa Sowerby, 1825
PATELLOIDAE Smaragdia bryannae Pilsbry, 1917 in Smaragdiinae
PATELLIDAE Theodoxus cariosus (Wood, 1828)
Cellana exarata (Reeve, 1854) Theodoxus neglectus (Pease, 1868)
Cellana melanostoma (Pilsbry, 1891). Pleistocene fossils, Theodoxus vespertinus (Sowerby, 1849)
Kaena Point, Oahu: James, 1989
Cellana sandwicensis (Pease, 1861) SMARAGDIINAE
Cellana talcosa (Gould, 1846) Smaragdia bryanae Pilsbry, 1917

NERITOIDEA PHENACOLEPADIDAE
NERITOPSIDAE Phenacolepas aculeata (Pease, 1868)
Neritopsis radula (Linnaeus, 1758) Phenacolepas granocostata (Pease, 1868)
Phenacolepas scobinata (Gould, 1859)
NERITIDAE
NERITINAE ORDER ARCHAEOGASTROPODA (DIOCARDIA)
Nerita albicilla Linnaeus, 1758 PLEUROTOMARIOIDEA
Nerita picea (Recluz, 1841). Pleistocene fossils, Kaena Point, SCISSURELLIDAE
Oahu: James, 1989
Scissurella lamellata (A. Adams, 1862)
Nerita plicata Linnaeus, 1758
Scissurella pseudoequatoria Kay, 1979
Nerita polita Linnaeus, 1758
Sinezona insignis (Smith, 1910)
Calliotropis (Solaricida) reticulina (Dall, 1895)
FISSURELLOIDEA
FISSURELLIDAE ENCYCLINAE
DIODORINAE Danilia eucheliformis (Nomura and Hatai, 1940)
Diodora granifera (Pease, 1861) Euchelus corrugatus Pease, 1861
Diodora octagona (Reeve, 1850) Euchelus fimbriatus (Pease, 1861)
Diodora ruppelli (Sowerby, 1834) Euchelus gemmatus (Gould, 1845)
Gibbula marmorea (Pease, 1861)
EMARGINULINAE
Emarginula dilecta A. Adams, 1852 TROCHINAE
Emarginula hawaiiensis Dall, 1895 Clanculus clanguloides (Wood, 1818)
Tugali oblonga (Pease, 1861) Thalotia ocellata (A. Adams, 1861) = Alcyna ocellata : Hickman
and McLean, 1990
TROCHOIDEA Thalotia subangulata (Pease, 1861) = Alcyna subangulata:
Hickman and McLean, 1990
TROCHIDAE
Tristichotrochus margaritissimus Habe and Okutani, 1968
CALLIOSTOMINAE
Trochus intextus Kiener, 1850
Calliostoma (Tristichotrochus) doncorni Kay, 1979
2 Moretzsohn and Kay

Cerithium placidum Gould, 1861

TURBINIDAE Cerithium rostratum Sowerby, 1855
COLLONINAE Plesiotrochus luteus (Gould, 1861) in Plesiotrochidae: Wilson,
Leptothyra candida (Pease, 1861) = Collonista candida: Hickman 1993.
and McLean 1990 Rhinoclavis articulata (Adams and Reeve, 1850)
Leptothyra rubricincta (Mighels, 18 Rhinoclavis fasciata (Bruguière, 1792)
45) Rhinoclavis sinensis (Gmelin, 1791)
Leptothyra verruca (Gould, 1845)
DIALIDAE
TURBININAE Argyropeza leucocephala (Watson, 1886) changed to Cerithidae:
Galleoastrea midwayensis Habe and Kosuge, 1970 Houbrick
Turbo sandwicensis Pease, 1861. Pleistocene fossils, Kaena Point, Cerithidium diplax (Watson, 1886)
Oahu: James, 1989 Cerithidium perparvulum (Watson, 1886)
Turbo setosus Gmelin, 1791 Diala flammea ? Couturier, 1907 (not of Pease, 1867) = Diala
albugo (Watson, 1886): Ponder & de-Keyzer, 1992
SKENEIDAE Diala scopulorum (Watson, 1886)
Brookula iki Kay, 1979 Diala sulcifera (A. ADAMS, 1862): Ponder & de-Keyzer, 1992
Cyclostremiscus emery (Ladd, 1966) in Vitrinellidae: Ponder, Diala varia A. Adams, 1861 = Diala semistriata (Philippi, 1849):
1985 Ponder & de-Keyzer, 1992
Cyclostremiscus striatus Kay, 1979 in Vitrinellidae: Ponder, 1985 Finella pupoides A. Adams, 1860 changed to Obtortionidae:
Lophocochlias minutissimus (Pilsbry, 1921) Ponder
Nannoteretispira habei Kay, 1979 = Pelycideon habei (Kay, Scaliola spp. changed to Scaliolidae: Ponder
1979): Ponder, 1985
FOSSARIDAE: not in Hipponoidea: Wilson, 1993
STOMATELLIDAE Fossarus cumingii (A. Adams, 1855)
Synaptocochlea concinna (Gould, 1845) Fossarus garrettii Pease, 1868
Fossarus multicostatus Pease, 1861
PHASIANELLIDAE = TRICOLIINAE: Hickman and McLean, 1990
Tricolia (Hiloa) variabilis (Pease, 1861) MODULIDAE
Modulus tectum (Gmelin, 1791)
SUPERORDER CAENOGASTROPODA
ORDER NEOTAENIOGLOSSA OBTORTIONIDAE
CERITHOIDEA Finella pupoides A. Adams, 1860 changed from Dialidae: Ponder
CERITHIIDAE
Argyropeza leucocephala (Watson, 1886) changed from Dialidae: PLANAXIDAE
Houbrick Planaxis labiosa A. Adams, 1853
Bittium impendens (Hedley, 1899) Planaxis saturalis E. A. Smith, 1872
Bittium parcum (Gould, 1861) = Ittibittium parcum: Houbrick,
1993a
PLESIOTROCHIDAE
Bittium zebrum (Kiener, 1841) = Cerithium zebrum Kiener, 1841:
Plesiotrochus luteus (Gould, 1861)
Houbrick, 1993b
Cerithium atromarginatum Dautzenberg and Bouge, 1933
LITTORINOIDEA
Cerithium boeticum Pease, 1860. A valid species, not a synonym
of Cerithium zebrum: Houbrick, 1993b LITTORINIDAE
Cerithium columna Sowerby, 1834 LITTORININAE
Cerithium echinatum. New record: Houbrick, 1992 Littoraria coccinea (Gmelin, 1791) changed from Littorina: Reid
Cerithium egenum Gould, 1849 Littoraria scabra (Linnaeus, 1758)
Cerithium interstriatum Sowerby, 1855 Littoraria undulata (Gray, 1839)
Cerithium matukense Watson, 1886. Also dredged from 200 Littorina pintado (Wood, 1828) = Littoraria: Reid. Synonym: in
fathoms off Waikiki: Hemmes and Goldsmith, 1994 part Littoraria intermedia (Philippi, 1846)
Cerithium mutatum Sowerby, 1834 = Cerithium echinatum: Nodilittorina hawaiensis, new name for Littorina planaxis
Houbrick, 1992 ''Nuttall''' Tryon, 1887; L. picta marmorata Philippi,
Cerithium nesioticum Pilsbry and Vanatta, 1905
 Hawaiian Marine Molluscs 3

Edmondson, 1933; Nodilittorina picta (Philippi, 1846) Kay, Vitricithna marmorata (Hedley, 1907) = Pusillina (Haurakia)
1979: Rosewater marmorata: Ponder, 1985
Peasiella tantilla (Gould, 1849)
RISSOININAE
CINGULOPSOIDEA Isselia hiloense (Pilsbry and Vanatta, 1908) = Stosiscia (lselliella)
CINGULOPSIDAE hiloense: Ponder, 1985
Rufodardanula conica Kay, 1979 Merelina granulosa (Pease, 1862)
Rufodardanula ponderi Kay, 1979 Merelina hewa Kay, 1979
Merelina kenneyi changed to Sansonia kenneyi (Ladd, 1966) in
Vitrinellidae
EATONIELLIDAE
Merelina wanawana Kay, 1979
Eatoniella (Caveatoniella) janetaylorae Kay, 1979
Pyramidelloides miranda (A. Adams, 1861) changed to
Eatoniella (Dardaniopsis) pigmenta Kay, 1979
Eulimidae: Ponder, 1985
Pyramidelloides suta (Pilsbry, 1921) changed to Eulimidae:
RASTODENTIDAE Ponder, 1985
Rastodens brevilabiosa Kay, 1979 Rissoina (Rissoina) ambigua (Gould, 1849)
Rastodens labiosa Kay, 1979 Rissoina (Rissoina) pulchella (Brazier, 1877)
Rissoina ephamilla (Watson, 1886) = Schwartziella (Pandalosia)
TRUNCATELLOIDEA ephamilla: Ponder, 1985
ASSIMENEIDAE Rissoina miltozona Tomlin, 1915 = Rissoina (Apataxia)
Assiminea nitida (Pease, 1865) cerithiiformis Tryon, 1887: Sleurs and Preece, 1994
Rissoina turricula Pease, 1861 = Rissoina (Rissoina) costata A.
Adams, 1851: Sleurs, 1993
BARLEEIDAE elevated to family, not a subfamily of Rissoidae:
Ponder, 1985 Schwartziella gracilis (Pease, 1861) = Pyramidelloides gracilis,
in Eulimidae: Ponder, 1985
Barleeia caIcarea Kay, 1979 changed from Rissoidae: Ponder,
1985 Schwartziella triticea Pease, 1861 = Schwartziella (Schwartziella)
triticea (Pease, 1861): Ponder, 1985
Zebina imbricata (Gould, 1861) = Rissoina (Rissoina) imbricata
CAECIDAE Gould, 1861: Ponder, 1985
Caecum arcuatum de Folin, 1867 Zebina semiplicata (Pease, 1862) = Zebina (Zebina) bidentata
Caecum cf. glabriformis (A. Adams, 1868). New record, 180 ft. (Philippi, 1845) sensu lato: Sleurs, 1993
of Kepuhi Pt, Oahu: Lighfoot, 1978 Zebina tridentata (Michaud, 1830)
Caecum sepimentum de Folin, 1867. Pleistocene fossils, Kaena
Point, Oahu: James, 1989; = Caecum arcuatum de Folin, 1867;
represented a juvenile: Lighfoot, 1992. VITRINELLIDAE
Caecum vertebrale/oahuense (Pilsbry, 1921) = Caecum exile is a Elacorbis callusa (Laseron, 1958)
juvenile: Lighfoot, 1992 Teinostoma sp. cf. sulcata (A. Adams, 1850)
Caecum vertebrale Hedley, 1899
Caecum (Brochina) sp. cf. glabella A. Adams, 1868 STROMBOIDEA
Caecum (Fartulum) exile (de Folin, 1875) STROMBIDAE
Caecum (Micranellum) oahuense (Pilsbry, 1921) Strombus dentatus Linnaeus, 1759 = Strombus (Canarium)
Meioceras sandwichensis de Folin, 1886 dentatus: Wilson, 1993
Strebloceras subannulatum de Folin, 1879 Strombus erythrinus erythrinus Dillwyn, 1817. Two shells from
Waikiki: Hemmes and Goldsmith, 1994
Strombus fragilis (Röding, 1798) = Strombus (Canarium) fragilis:
RISSOIDAE Wilson, 1993
RISSOINAE Strombus helli Kiener, 1843
Alvinia isolata (Laseron, 1956) = Alvania (Alvania) isolata: Strombus maculatus Sowerby, 1842
Ponder, 1985
Strombus vomer hawaiensis Pilsbry, 1917 = Strombus
Elachisina robertsoni Kay, 1979 changed to Elachisinidae: (Euprotomus) vomer hawaiensis: Wilson, 1993. Live specimen
Ponder, 1986 collected: Hemmes and Goldsmith, 1994
Nannoteretispira habei Kay, 1979 changed to Pelycidion habei, Strombus wilsoni Abbott, 1967 = Strombus (Canarium) wilsoni:
Pelycidiidae: Ponder, 1985 Wilson, 1993
Parashiela beetsi Ladd, 1966
Powellsetia fallax Kay, 1979 VANIKORIOIDEA
4 Moretzsohn and Kay

HIPONNICIDAEA Cypraea burgessi Kay, 1981. New record, described from Pearl
Hipponix (Antisabia) foliaceus (Quoy and Gaimard, 1835). and Hermes Reef, also from Maui and Honolulu Harbor: Kay,
Pleistocene fossils, Kaena Point, Oahu: James, 1989; = 1981
Antisabia foliacea, may be a complex of 3-4 species: Knudsen, Cypraea caputserpentis Linnaeus, 1758
1993 Cypraea carneola Linnaeus, 1758
Hipponix (Cochlear) imbricatus Gould, 1846 Cypraea cernica Sowerby, 1870
Hipponix (Pilosabia) pilosus (Deshayes, 1832) Cypraea childreni Gray, 1825
Sabia conica (Schumacher, 1817) = Hipponix australis Lamarck, Cypraea chinensis Gmelin, 1791
1819: Knudsen, 1991
Cypraea cicercula Linnaeus, 1758
Cypraea eglantina Duclos, 1833
VANIKORIDAE
Cypraea errones Linnaeus, 1758
Vanikoro acuta (Recluz, 1844)
Cypraea erosa Linnaeus, 1758. Live specimen from S. Oahu:
Vanikoro cancellata (Lamarck, 1822) Cook, 1995b
Vanikoro imbricata Pease, 1861 Cypraea fimbriata Gmelin, 1791
Cypraea gaskoini Reeve, 1846
CALYPTRAEOIDEA Cypraea gracilis Gaskoin, 1849
CALYPTRAEIDAE Cypraea granulata Pease, 1863
Cheilea equestris (Linnaeus, 1758) Cypraea helvola Linnaeus, 1758
Crepidula aculeata (Gmelin, 1791) Cypraea isabella Linnaeus, 1758
Crucibulum spinosum (Sowerby, 1824) Cypraea labrolineata Gaskoin, 1849
Cypraea leviathan Schilder and Schilder, 1938
CAPULIDAE Cypraea lynx Linnaeus, 1758
Capulus bicarinatus (Pease, 1861) Cypraea maculifera Schilder, 1932
Cypraea mariae Schilder, 1927
XENOPHOROIDEA Cypraea mauiensis Burgess, 1967. Fresh-dead specimen, off West
XENOPHORIDAE Oahu: Cook, 1995b
Xenophora peroniana (lredale, 1929) = Xenophora (Xenophora) Cypraea mauritiana Linnaeus, 1758
peroniana: Wilson, 1993 Cypraea moneta Linnaeus, 1758
Xenophora (Xenophora) peroniana kondoi Ponder, 1983. New Cypraea nucleus Linnaeus, 1758
record: Wilson, 1993
Cypraea ostergaardi Dall, 1921
Cypraea poraria Linnaeus, 1758
VERMETOIDEA
Cypraea rashleighana Melvill, 1888
VERMETIDAE
Cypraea schilderorum (Iredale, 1939)
Dendropoma gregaria Hadfield and Kay, 1972. Pleistocene
Cypraea scurra Gmelin, 1791
fossils, Kaena Point, Oahu: James, 1989
Cypraea semiplota Mighels, 1845
Dendropoma maxima Sowerby, 1825
Cypraea staphylaea Linnaeus, 1758
Dendropoma meroclista Hadfield and Kay, 1972
Cypraea sulcidentata Gray, 1824
Dendropoma platypus Mörch, 1861
Cypraea talpa Linnaeus, 1758
Dendropoma psarochephala Hadfield and Kay, 1972
Cypraea teres Gmelin, 1791
Dendropoma rhyssoconcha Hadfield and Kay, 1972
Cypraea tessellata Swainson, 1822
Petaloconchus keenae Hadfield and Kay, 1972
Cypraea tigris Linnaeus, 1758
Serpulorbis variabilis Hadfield and Kay, 1972. Pleistocene
fossils, Kaena Point, Oahu: James, 1989 Cypraea vitellus Linnaeus, 1758
Vermetus alii Hadfield and Kay, 1972
OVULIDAE
CYPRAEOIDEA Lunovula finley Rosenberg, 1990. New record. Described from
Hawaii, 120 m: Rosenberg, 1990
CYPRAEIDAE
Margovula sp. cf. schilderorum C.N. Cate, 1973
Cypraea alisonae Burgess, 1983. New record, described from
Hawaii: Burgess, 1983. Also from Australia: Wilson, 1993 Phenacovolva brevirostris (Schumacher, 1817)
Cypraea arabica Linnaeus, 1758 Phenacovolva lahainaensis (C.N. Cate, 1969)
Cypraea beckii Gaskoin, 1836 Phenacovolva weaveri C.N. Cate, 1973
Spiculata michaelkingi C.N. Cate, 1973
 Hawaiian Marine Molluscs 5

Bursa rhodostoma Sowerby, 1841

LAMELLARIOIDEA Bursa rosa (Perry, 1811)
TRIVIIDAE Tutufa bufo (Röding, 1798)
TRIVIINAE
Trivia edgari Shaw, 1909 = Trivia (trivirostra) edgari: Wilson, CASSIDAE
1993 CASSINAE
Trivia exigua Gray, 1831 Cassis cornuta (Linnaeus, 1758)
Trivia globosa pilula Kiener, 1843 = Trivia (Clotrivia) globosa
pilula: Wilson, 1993
PHALINAE
Trivia hordacea Kiener, 1845 = Trivia (Trivirostra) hordacea:
Casmaria erinaceus kalosmodix (Melvill, 1883)
Wilson, 1993
Casmaria ponderosa (Gmelin, 1791)
Trivia pellucida Reeve, 1846 = Trivia (Trivirostra) pellucida:
Wilson, 1993 Phallium (Semicassis) bulla (Habe, 1961)
Phallium (Semicassis) umbilicatum (Pease, 1861)
ERATOINAE Phallium coronadoi wyvillei (Watson, 1886)
Erato sandwicensis Pease, 1860
PERSONIDAE
LAMELLARIIDAE Distorsio anus (Linnaeus, 1758)
Lamellaria sp. Distorsio burgessi Lewis, 1972
Distorsio constricta Lewis, 1972. New record: Beu, 1986
NATICOIDEA Distorsio pusilla Pease, 1861
NATICIDAE
NATICINAE RANELLIDAE (= CYMATIIDAE): See Beu, 1987
Euspira sandwichensis (Dall, 1895) RANELLINAE
Natica alapapilionis (Röding, 1798) = Natica (Naticarius) Fusitron midwayensis Habe and Okutani, 1968
alapapilionis: Wilson, 1993 Gyrineum louisae Lewis, 1974
Natica bougei Sowerby, 1908 = Natica (Tectonatica) bougei: Gyrineum pusillum (Broderip, 1832)
Wilson, 1993
Natica gualteriana Récluz, 1844 = Natica (Naticarius)
CYMATIINAE
gualteriana: Wilson, 1993
Charonia tritonis (Linnaeus, 1767)
Natica hilaris Sowerby, 1914
Cymatium (Cymatium) nicobaricum (Röding, 1798) = Cymatium
Natica sp. Fig. 72 I, J = Natica ochrostigmatica Rehder, 1980
(Monoplex) nicobaricum: Beu and Kay, 1988
Cymatium (Gutturnium) gutturnium (Röding, 1798) = Cymatium
POLINICINAE (Ranularia) gutturnium: Wilson, 1993; Fig. 78 B not A of
Polinices melanostomus (Gmelin, 1791) = Polinices (Mammilia) Kay, 1979: Beu
melanostomus: Wilson, 1993 Cymatium (Gutturnium) muricinum (Röding, 1798)
Polinices peselephanti (Link, 1807) = Polinices (Mammilia) Cymatium (Gutturnium) pyrum (Linnaeus, 1758) = Cymatium
peselephanti: Wilson, 1993 (Ranularia) pyrum: Wilson, 1993.
Polinices simiae (Deshayes in Deshayes and Edwards, 1838) = Cymatium (Gutturnium) sarcostomum (Reeve, 1844) = Cymatium
Polinices (Mammilia) simiae: Wilson, 1993 (Ranularia) sarcostomum. Fig. 78A not C of Kay, 1979
Polinices tumidus (Swainson, 1840) Cymatium (Linatella) cingulatum (Lamarck, 1822) = Linatella
(Linatella) candata (Gmelin, 1791); Fig. 79A not 79B of Kay,
SININAE 1979: Cook
Eunaticina margaritaeformis Dall, 1924 Cymatium (Linatella) clandestinum (Lamarck, 1816) = Linatella
(Gelagna) succinta (Linneus, 1771); Fig. 79B of Kay, 1979
Cymatium (Septa) aquatile (Reeve, 1844) = Cymatium
TONNOIDEA: Wilson, 1993 (Monoplex) aquatile: Wilson, 1993
BURSIDAE Cymatium (Septa) gemmatum (Reeve, 1844) = Cymatium
Bursa bufonia (Gmelin, 1791) (Monoplex) mundum (Gould, 1849): Beu
Bursa cruentata Sowerby, 1841 Cymatium (Septa) hepaticum (Linnaeus, 1758)
Bursa granularis (Röding, 1798) Cymatium (Septa) intermedius (Pease, 1869) = Cymatium
Bursa latitudo Garrard, 1961 (Monoplex) intermedium: Beu
Bursa luteostoma (Pease, 1861). New record. Valid species, not a Cymatium (Septa) occidentale (Mörch, 1877). New record, Sandy
synonym of Bursa bufonia (Gmelin, 1791): Cook, 1995a Beach, Oahu, 45-65 ft: Cook, 1995a
6 Moretzsohn and Kay

Cymatium (Septa) parthenopea (von Salis, 1793) = Cymatium Carinaria lamarcki Peron and Lesueur, 1810. New record,
(Monoplex) parthenopeum: Wilson, 1993 Hawaiian waters: Seapy, 1987
Cymatium (Septa) pileare (Linnaeus, 1758) = Cymatium Pterosoma planum Lesueur, 1827. New record, Hawaiian waters:
(Monoplex) pileare: Wilson, 1993 Seapy, 1987
Cymatium (Septa) rubeculum (Linnaeus, 1758)
Cymatium (Septa) vespacium (Lamarck, 1822) PTEROTRACHEIDAE
Cymatium (Turritriton) comptum (A.Adams, 1855). New record. Pterotrachea coronata Niebuhr (ms. Forskål), 1775. New record,
Fig. 79E not F of Kay, 1979: Cook, 1995 Hawaiian waters: Seapy, 1987
Cymatium (Turritriton) labiosum (Wood, 1828). Fig. 79F not E of Pterotrachea hippocampus Philippi, 1836. New record, Hawaiian
Kay, 1979: Cook, 1995 waters: Seapy, 1987
Sassia sp. New record, off N. Shore of Oahu, 333 m: Cook, 1995 Pterotrachea scutata Gegenbaur, 1855. New record, Hawaiian
waters: Seapy, 1987
TONNIDAE
Eudolium pyriforme (Sowerby, 1814). New record, off Oahu, 223 FIROLIDAE
m: Marshall, 1992 Firoloida desmaresti. New record, Hawaiian waters: Seapy,
Malea pomum (Linnaeus, 1758) 1990b
Tonna cepa (Röding, 1798)
Tonna dolium (Linnaeus, 1758) SUBORDER PTENOGLOSSA
Tonna melanostoma (Jay, 1839) TRIPHOROIDEA
Tonna perdix (Linnaeus, 1758) CERITHIOPSIDAE
Cerithiopsis arga Kay, 1979
SUBORDER HETEROPODA Joculator granata Kay, 1979
ATLANTIDAE Joculator ridicula (Watson, 1886)
Atlanta echinogyra Richter, 1972. New record, Hawaiian waters: Joculator semipicta (Gould, 1861)
Seapy, 1990a Joculator turrigera(Watson, 1886)
Atlanta fusca Souleyet, 1852. New record, Hawaiian waters: Joculator uveanum (Melvill and Standen, 1896)
Seapy, 1990a
Atlanta helicinoides Souleyet, 1852. New record, Hawaiian TRIPHORIDAE
waters: Seapy, 1990a
METAXIINAE
Atlanta inflata Souleyet, 1852. New record, Hawaiian waters:
Metaxia albicephala Kay, 1979
Seapy, 1990a
Metaxia brunnicephala Kay, 1979
Atlanta lesueuri Souleyet, 1852. New record, Hawaiian waters:
Seapy, 1990a Metaxia tricarinata (Pease, 1861)
Atlanta meteori Richter, 1972. New record, Hawaiian waters:
Seapy, 1990a INIFORINAE
Atlanta olygogyra Tesch, 1906. New record, Hawaiian waters: Iniforis aemulans (Hinds, 1843)
Seapy, 1990a Iniforis concors (Hinds, 1843)
Atlanta peroni Lesueur, 1817. New record, Hawaiian waters: Iniforis hinuhinu Kay, 1979
Seapy, 1990a
lniforis ordinata Laseron, 1958
Atlanta plana Richter, 1972. New record, Hawaiian waters:
Seapy, 1990a lniforis peleae (Baker and Spicer)
Atlanta tokiokai van der Spoel and Troost, 1972. New record, lniforis perfecta (Pease, 1871)
Hawaiian waters: Seapy, 1990a
Atlanta turriculata d’Orbigny, 1836. New record, Hawaiian MASTONIINAE
waters: Seapy, 1990a Cautor sp. cf. intermissa (Laseron, 1958)
Oxygyrus keraudreni (Lesueur, 1817). New record, Hawaiian Cautor minima (Pease, 1871). Figure is 49A
waters: Seapy, 1990a
Cautor similis (Pease, 187 1)
Protatlanta souleyeti (Smith, 1888). New record, Hawaiian
lnella spp
waters: Seapy, 1990a
Litharium maculata (Pease, 1871)
Litharium oceanida (Dall, 1924)
CARINARIIDAE
Mastonia cingulifera (Pease, 1861)
Carinaria galea Benson, 1835. New record, Hawaiian waters:
Seapy, 1987 Mastonia gracilis (Pease, 1871). Figure is 49H
Carinaria japonica Okutani, 1955. New record, Hawaiian waters: Mastonia troglodytes (Hervier, 1897)
Seapy, 1987 Viriola abbotti Baker and Spicer, 1935)
 Hawaiian Marine Molluscs 7

Viriola bayani Jousseaume, 1884 Epitonium hyalina mokuoloense Pilsbry, 1921 = Cycloscala
Viriola bilix (Hinds, 1843) hyalina (Sowerby, 1844); Epitonium revolutum (Hedley, 1899)
is a synonym: DuShane, 1988a
Viriola cancellata Hinds, 1843
Epitonium kanemoe Pilsbry, 1921
Viriola elongata (Laseron, 1958)
Epitonium liliputanum (A. Adams, 1861). New record, Hawaiian
Viriola fallax Kay, 1979
specimens from Burch collection, 200-400 m, Mamala Bay,
Viriola flammulata (Pease, 1871) Oahu: DuShane, 1987
Viriola incisa (Pease, 1861) Epitonium oahuense Pilsbry, 1921
Viriola pagoda (Hinds, 1843) Epitonium paumotensis (Pease, 1868)
Epitonium perplexum (Deshayes, 1863)
TRIPHORINAE Epitonium replicatum (Sowerby, 1844) not E. alatum (Sowerby,
Triphora bicolor (Pease, 1868) 1844): DuShane, 1987
Triphora chrysolitha Kay, 1979 Epitonium revolutum (Hedley, 1899) = Cycloscala hyalina
Triphora coralina (Laseron, 1958) (Sowerby, 1844): DuShane, 1988a
Triphora earlei Kay, 1979 Epitonium thorssoni DuShane, 1988. New record, described from
S. side of Oahu, 62-80 m. Also from Guam, Philippines and
Triphora isaotakii (Kosuge, 1962) Egypt: DuShane, 1988d
Triphora keiki Kay, 1979 Epitonium ulu Pilsbry, 1921
Triphora laddi Kay, 1979 Epitonium umbilicatum (Pease, 1869)
Triphora lutea (Kosuge, 1962) Laeviscala luceo DuShane, 1988. New record, described from
Triphora pallida (Pease, 1871) Sand Island, Oahu, 17 m: DuShane, 1988d
Triphora peasi (Jousseaume, 1884) Opalia (Nodiscala) attenuata (Pease, 1861)
Triphora pustulosa (Pease, 1871) Opalia bicarinata Sowerby, 1844. New record, Pearl Harbor
Triphora rufotincta (Kosuge, 1963) Channel, 30 ft: Cook, 1992
Triphora tessellata (Kosuge, 1963) Opalia (Opalia) burchorum DuShane, 1988. New record,
described from off Kahe Point, Oahu, 540 m: DuShane, 1988d
Triphora thaanumi Kay, 1979
Parviscala cf. gloriola (Melvill and Standen, 1903). New record,
Triphora triticea Pease, 1861
Oahu, Mamala Bay, 29-47 m: DuShane, 1988c
Triphora truncis (Laseron, 1958)
Parviscala crispata (Pease, 1867). New record, Mamala Bay,
Triphora tuberculata (Pease, 1871) Oahu, 90 m: DuShane, 1988b
Triphora tubularis (Laseron, 1958) Parviscala deifica (Melvill and Standen, 1903). New record,
Hawaii and Oahu: DuShane, 1988b
JANTHINOIDEA Parviscala gradilis (Jousseaume, 1912). New record, Hawaii:
EPITONIIDAE DuShane, 1988c
Amaea xenicima (Melvill and Standen, 1903). New record: Parviscala paumotensis (Pease, 1867). New record, Kauai, 72 m:
DuShane, 1988e DuShane, 1988c
Asperiscala goldsmithi DuShane, 1988. New record; described Parviscala symmetrica (Pease, 1867). New record, Oahu, 270 m;
from Puhi Bay, Hilo, Hawaii, intertidal, Oahu, Maui. Also from Reef: DuShane, 1988c
Papua New Guinea, Fiji, Thailand and Maldives Is.: DuShane,
1988d JANTHINIDAE
Cirsotrema varicosa (Lamarck, 1822) Janthina exigua Lamarck, 1816
Cycloscala crenulata (Pease, 1867). New record: Lahilani Cave, Janthina globosa Swainson, 1822
Oahu. Type locality: Tahiti: DuShane, 1988a
Janthina janthina (Linnaeus, 1758)
Epitonium oahuense (Pilsbry, 1921) = Drepressiscala umbilicata
(Pease, 1869). DuShane, 1988f: record from Mamala Bay,
Oahu, 213 m EULIMOIDEA
Epitonium (Librariscala) millecostatum (Pease, 1861). DuShane, EULIMIDAE
1988b introduces Kilburn's 1985 subgenus Balcis acanthyllis (Watson, 1886)
Epitonium alatum (Sowerby, 1844), fig. 54C is E. replicatum Balcis aciculata (Pease, 1861)
(Sowerby, 1844), but species is reported in Hawaiian Shell Balcis brunnimaculata Kay, 1979. Pleistocene fossils, Kaena
News Dec. 1985: 12 by C. Takahashi: DuShane, 1987 Point, Oahu: James, 1989
Epitonium fucatum (Pease, 1861) Balcis bryani (Pilsbry, 1917)
Epitonium hemmesi DuShane, 1988. New record, described from Balcis conoidalis (Sowerby, 1865)
Lahilanhi Point, Oahu, 20-27 m: DuShane, 1988d
Balcis cumingii (A. Adams, 1854)
Balcis inflexa (Pease, 1868)
8 Moretzsohn and Kay

Balcis kanaka (Pilsbry, 1917) LITIOPIDAE

Balcis letsonae (Pilsbry, 1917) Styliferina goniochila (A. Adams, 1860)
Balcis solida (Sowerby, 1865)
Balcis solidula (Adams and Reeve, 1848) ORDER HETEROSTROPHA
Balcis thaanumi (Pilsbry, 1917) VALVATOIDEA
Ctenosculum hawaiiense Heath, 1910 is a barnacle: Warén, 1981 ORBITESTELLIDAE
Echineulima mittrei Petit, 1851 Orbitestella regina Kay, 1979 in Rissoeloidea: Ponder
Echineulima robusta Pease, 1860
Eulima metcalfei (A. Adams, 1853). Fig. 56 Q ARCHITECTONICOIDEA
Eulima peasei Tryon, 1886. Fig. 56P ARCHITECTONICIDAE
Melanella ima Pilsbry, 1917. A valid species, not a synonym of Architectonica maxima (Philippi, 1849)
Balcis thaanumi (Pilsbry, 1917). Record of 39 specimens on a Architectonica perspectiva (Linnaeus, 1758)
single, undetermined holothurian, Hilo, Hawaii: Warén, 1983
Heliacus implexus (Mighels, 1845)
Mucronalia nitidula Pease, 1861 = Peasistilifer nitidula (Pease,
Heliacus mighelsi (Philippi, 1853)
1861). Warén proposed new genus, Peasistilifer, type species:
Mucronalia nitidula Heliacus sterkii (Pilsbry and Vanatta, 1908)
Mucronalia rosacea = Mucronalia holothurica Adam, 1934: Heliacus trochoides (Deshayes, 1830)
Warén, 1981 Heliacus variegatus (Gmelin, 1791)
Mucronalia ovata Pease, 1861. Stilapex philippinarum Habe, Philippia oxytropis A. Adams, 1855
1976 a synonym, and Hawaiian locality probably wrong: Philippia radiata (Röding, 1798)
Warén, 1981.
Pseudomalaxis sp. cf. cornuammonis (Melvill and Standen, 1903)
Pelseneeria hawaiiensis Warén, B. Burch, and T. Burch, 1984.
New record; attached by the proboscis on a specimen of
Aspidodiadema hawaiiensis: Warén, B. Burch, and T. Burch, OMALOGYROIDEA
1984 Omalogyra japonica (Habe, 1972)
Pulicochlea calimaris Ponder and Gooding, 1979. New record:
Ponder and Gooding, 1979
RISSOELLOIDEA
Pulicochlea faba Ponder and Gooding, 1979. New record: Ponder
RISSOELLIDAE
and Gooding, 1979
Rissoella sp. cf. confusa confusa Ponder and Yoo, 1977
Pyramidelloides miranda (A. Adams, 1861) changed from
Rissoidae: Ponder, 1985 Rissoella longispira Kay, 1979
Pyramidelloides suta (Pilsbry, 1921) changed from Rissoidae:
Ponder, 1985 ORDER NEOGASTROPODA (= STENOGLOSSA)
Scalenostoma subulata (Broderip, 1832) non S. carinata MURICOIDEA
Deshayes, 1863: Warén, 1980 MURICIDAE
Stilifer linckiae Sarasin and Sarasin, 1887 Aspella producta (Pease, 1861)
Thyca crystallina (Gould, 1846) Chicoreus insularum (Pilsbry, 1921)
Trochostilifer entospinea Warén, B. Burch and T. Burch, 1984. Favartia garrettii (Pease, 1869)
New record, from galls in the spines of the sea urchin
Chondrocidaris gigantea: Warén, B. Burch, and T. Burch, 1984 Homolocantha anatomica (Perry, 1811)
Trochostilifer hawaiiensis Warén, B. Burch and T. Burch, 1984. Marchia laqueata (Sowerby, 1841)
New record, from a gall in the test of the sea urchin Marchia martinetana (Röding, 1798)
Prionocidaris hawaiiensis: Warén, B. Burch, and T. Burch, Marchia triptera (Born, 1778)
1984
Muricodrupa funiculusa (Wood, 1828)
Vitreolina chondrocidaricola Warén, B. Burch and T. Burch,
Phyllocoma convoluta (Broderip, 1833)
1984. New record, from the intestine of the sea urchin
Chondrocidaris gigantea: Warén, B. Burch, and T. Burch, 1984 Pterynotus elongatus (Solander in Lightfoot, 1786)
Vitreolina hawaiiensis Warén, B. Burch and T. Burch, 1984. New Trophonopsis kayae Habe, 1981. New record, described from off
record, from the sea urchin Prionocidaris hawaiiensis: Warén, Honolulu, 369m: Habe, 1981
B. Burch, and T. Burch, 1984 Vitularia miliaris (Gmelin, 1791)

SCALIOLIDAE THAIDIDAE
Scaliola bella A. Adams, 1860 Drupa (Drupa) morum Röding, 1798
Scaliola glareosa A. Adams, 1862 Drupa (Drupa) ricina (Linnaeus, 1758)
Scaliola gracilis A. Adams, 1862 Drupa (Drupina) grossularia (Röding, 1798)
Drupa (Ricinella) rubusidaeus Röding, 1798
 Hawaiian Marine Molluscs 9

Drupa elegans (Broderip and Sowerby, 1829) Engina alveolata (Kiener, 1836)
Drupella elata Blainville, 1832 Phos varicosus (Gould, 1849) = Phos roseatus (Hinds, 1844):
Drupella ochrostoma (Blainville, 1832) Cernohorsky, 1978
Maculotriton bracteatus (Hinds, 1844) Prodotia ignea (Gmelin, 1791)
Morula benedictus (Melvill and Standen, 1895) Prodotia iostomus (Gray in Griffiths and Pidgeon, 1834)
Morula darrosensis (Smith, 1884)
Morula dumosa (Conrad, 1837) COLUMBELLIDAE
Morula foliacea (Conrad, 1837) Anachis miser (Sowerby, 1844)
Morula funiculata (Reeve, 1846) Caducifer nebulosa (Gould, 1860) = Zafrona nebulosa: Sleurs,
1987
Morula granulata (Duclos, 1832)
Euplica cf. liocyma Pilsbry, 1904. New record, Oahu, 10-30 m;
Morula spinosa (H. and A. Adams, 1853)
Hawaii; 30 m: Hemmes et al., 1990
Morula uva (Röding, 1798)
Euplica livescens (Reeve, 1859)
Morula vexilla (Kuroda, 1953)
Euplica turturina (Lamarck, 1822)
Nassa serta (Bruguière, 1789)
Euplica varians (Sowerby, 1832)
Neothais harpa (Conrad, 1837)
Mitrella bella (Reeve, 1859)
Pinaxia versicolor (Gray, 1839)
Mitrella fusiformis (Pease, 1868) = Mitrella loyaltensis (Hervier,
Purpura aperta (Blainville, 1832) 1900): Sleurs, 1987
Thais armigera (Link, 1807) Mitrella margarita (Reeve, 1859)
Thais echinulata (Lamarck, 1822) Mitrella rorida (Reeve, 1859)
Thais intermedia (Kiener, 1836) Seminella smithi (Angas, 1877)
Vexilla fusconigra Pease, 1860 Seminella subphilodicia (Hervier, 1899). New record,
Vexilla lineata A. Adams, 1853 Northwestern Hawaiian lslands: Hemmes et al., 1990
Vexilla vexillum (Gmelin, 1791) Seminella virginea (Gould, 1860) = Seminella peasei (von
Martens and Langkavel, 1871). There is a prior Columbella
virginea Duclos: Sleurs, 1987
CORALLIOPHILIDAE
Babelomurex cookae Kosuge, 1988. New record, described from
French Frigate Shoals, 75 to 175 fathoms: Kosuge, 1988 COLUBRARIIDAE
Coralliobia fimbriata (A. Adams, 1854) Colubraria muricata (Lightfoot, 1796)
Coralliophila d’orbignyana (Petit, 1851) Colubraria obscura (Reeve, 1844)
Coralliophila erosa (Röding, 1798) = Coralliophila bulbiformis Colubraria tortuosa (Reeve, 1844)
(Conrad, 1837) fide Rehder 1986 pers. comm.
Coralliophila sp. cf. nodosa (A. Adams, 1854) NASSARIIDAE
Coralliophila violacea (Kiener, 1836) Nassarius abyssicolus (A. Adams, 1852). New record, Sand
Hirtomurex goodwini Kosuge, 1988. New record, described from Island, Oahu, beachworn: Cook: 1988
French Frigate Shoals, 75 to 175 fathoms: Kosuge, 1988 Nassarius crematus (Hinds, 1844)
Latiaxis cuspidifera (Dall, 1924) Nassarius gaudiosus (Hinds, 1844)
Latiaxis japonicus (Dunker, 1882) Nassarius hirtus (Kiener, 1834)
Latiaxis tosanus Hirase, 1908 Nassarius olomea Kay, 1979
Magilopsis lamarckii (Deshayes, 1863) Nassarius papillosus (Linnaeus, 1758)
Quoyula madreporarum (Sowerby, 1834) Nassarius pauperus (Gould, 1850)
Rhizochilus antipathicus (Steenstrup, 1850) Nassarius shackelfordi Melvill and Standen, 1896

BUCCINIDAE FASCIOLARIIDAE
Caducifer decapitata (Reeve, 1844) Dolicholatirus acus (Adams and Reeve, 1850)
Caducifer nebulosa (Gould, 1860) = Zafrona nebulosa, in Fusinus nicobaricus (Röding, 1798)
Columbellidae: Sleurs, 1987 Fusinus sandvicensis (Sowerby, 1880)
Cantharus farinosus (Gould, 1850) Fusolatirus kuroseanus Okutani, 1975. Specimen from 33 m off
Cantharus pulcher (Reeve, 1846) Ewa, Oahu: Hemmes and Goldsmith, 1994
Clivipollia costata (Pease, 1860) Latirus craticulatus (Linnaeus, 1758)
Clivipollia fragaria (Wood, 1828) Latirus fasciatus Habe and Okutani, 1968. New record, from 100
Engina albocincta Pease, 1860 fathoms south of Maui, and from 1000 ft of Oahu: Cook, 1994
Latirus nodatus (Gmelin, 1791)
10 Moretzsohn and Kay

Latirus noumeensis (Crosse, 1870) Mitra (Nebularia) ferruginea Lamarck, 1811

Latirus sp. cf. caledonica (Petit, 1851) Mitra (Nebularia) fraga Quoy and Gaimard, 1833
Peristernia chlorostoma (Sowerby, 1825) Mitra (Nebularia) fulvecens Broderip, 1836
Peristernia squamosa (Pease, 1863) Mitra (Nebularia) luctuosa A. Adams, 1853
Peristernia ustulata (Reeve, 1847) Mitra (Nebularia) lugubris Swainson, 1821
Mitra (Nebularia) rubritincta Reeve, 1844
VOLUTOIDEA Mitra (Nebularia) testacea Broderip, 1836
OLIVIDAE Mitra (Nebularia) ticaonica Reeve, 1844
Oliva paxillus sandwicensis Pease, 1860 Mitra (Nebularia) turgida Reeve, 1844
Oliva richerti Kay, 1979 Mitra (Strigatella) acuminata Swainson, 1824
Olivella apicalis Kay, 1979 Mitra (Strigatella) assimilis Pease, 1868
Mitra (Strigatella) fastigium (Reeve, 1845)
HARPIDAE Mitra (Strigatella) litterata (Lamarck, 1811)
Harpa amouretta Röding, 1798 Mitra (Strigatella) pellisserpentis Reeve, 1844
Harpa goodwini Rehder, 1993. New record, described from Mitra (Strigatella) saltata Pease, 1865
French Frigate Shoals, 82-228 m: Rehder, 1993 Mitra (Strigatella) typha Reeve, 1845
Harpa harpa (Linnaeus, 1758)
Harpa major Röding, 1798 CYLINDROMITRINAE
Pterygia crenulata (Gmelin, 1791)
MARGINELLIDAE Pterygia fenestrata (Lamarck, 1811)
Cystiscus huna Kay, 1979. Misspelled in Kay, 1979. Radula Pterygia pudica (Pease, 1860)
figured: Coovert, 1987
Dentimargo pumila (Redfield, 1869)
IMBRICARIINAE
Granula sandwicensis (Pease, 1860)
Cancilla (Domiporta) granatina (Lamarck, 1811)
Granulina vitrea (Laseron, 1957)
Cancilla (Domiporta) sp. cf. carnicolor (Reeve, 1844)
Volvarina fusiformis (Hinds, 1844)
Imbricaria conovula (Quoy and Gaimard, 1833)
Volvarina nevilli (Jousseaume, 1875): not Volvarina fusiformis
Imbricaria olivaeformis (Swainson, 1821)
(Hinds, 1844): Coovert, 1987
Imbricaria punctata (Swainson, 1821)
Neocancilla clathrus (Gmelin, 1791)
MITRIDAE
Neocancilla kayae Cernohorsky, 1978
MITRINAE
Neocancilla papilio langfordiana J. Cate, 1962
Mitra (Dibaphus) edentula Swainson, 1823
Neocancilla waikikiensis Pilsbry, 1921
Mitra (Dibaphus) multiplicata Pease, 1865
Scabricola (Swainsonia) newcombii (Pease, 1869)
Mitra (Mitra) coffea Schubert and Wagner, 1829
Subcancilla flammea (Quoy and Gaimard, 1833)
Mitra (Mitra) incompta (Solander in Lightfoot, 1786)
Subcancilla foveata (Dunker, 1858)
Mitra (Mitra) maui Kay, 1979
Mitra (Mitra) mitra (Linnaeus, 1758)
Mitra (Mitra) nubila hawaiiensis Kay, 1979. Several shells
dredged at 10 m from Malai [sic] Bay (probably Malaea Bay, COSTELLARIIDAE
Maui): Hemmes and Goldsmith, 1994. Thala exilis (Reeve, 1845)
Mitra (Mitra) papalis (Linnaeus, 1758) Thala milium (Reeve, 1845)
Mitra (Mitra) pele Cernohorsky, 1970 Thala todilla (Mighels, 1845)
Mitra (Mitra) stictica (Link, 1807) Vexillum (Costellaria) adamsianum Cernohorsky, 1978
Mitra (Mitra) ustulata Reeve, 1844 Vexillum (Costellaria) albatum Cernohorsky, 1988. New record,
Mitra (Nebularia) aurora aurora Dohrn, 1861 Keehi Lagoon, Oahu, 110m: Cernohorsky, 1988
Mitra (Nebularia) avenacea Reeve, 1845 Vexillum (Costellaria) aureolineatum H. Turner, 1988. New
record, Nanakuli, Oahu, 55-108 m; described from Philippines;
Mitra (Nebularia) coarctata Reeve, 1844
also from Guam: Turner, 1988
Mitra (Nebularia) contracta Swainson, 1820
Vexillum (Costellaria) bellum (Pease, 1860
Mitra (Nebularia) coronata Lamarck, 1811
Vexillum (Costellaria) castum (H. Adams, 1872) = Scabricola
Mitra (Nebularia) cucumerina Lamarck, 1811 (Swainsonia) casta (H. Adams, 1872) non Voluta casta
Mitra (Nebularia) earlei Cernohorsky, 1977 Gmelin, 1791: Cernohorsky, 1988
 Hawaiian Marine Molluscs 11

Vexillum (Costellaria) collinsoni (A. Adams, 1853) Vexillum (Pusia) tusum (Reeve, 1845)
Vexillum (Costellaria) corbiculum (Sowerby, 1870) Vexillum (Pusia) unifascialis (Lamarck, 1811)
Vexillum (Costellaria) cosmani Kay, 1979 Vexillum salisbury New record
Vexillum (Costellaria) diutenera (Hervier, 1898)
Vexillum (Costellaria) filistratum (Sowerby, 1874) VOLUTOMITRIDAE
Vexillum (Costellaria) fortiplicatum (Pease, 1868) Volutomitra pailoloana (J. Cate, 1963)
Vexillum (Costellaria) interruptum (Anton, 1839)
Vexillum (Costellaria) interstriatum (Sowerby, 1870) CONOIDEA (= TOXOGLOSSA)
Vexillum (Costellaria) leucozonias Deshayes in La Borde and SUBORDER PTENOGLOSSA
Linant, 1834 TURRIDAE
Vexillum (Costellaria) macrospirum (A. Adams, 1853) TURRINAE
Vexillum (Costellaria) micra (Pilsbry, 1921) Gemmula congener unilineata Powell, 1967
Vexillum (Costellaria) modestum (Reeve, 1845) Gemmula interpolata Powell, 1967
Vexillum (Costellaria) pacificum (Reeve, 1845) Gemmula microscelida (Dall, 1895)
Vexillum (Costellaria) radix (Sowerby, 1874) Gemmula monilifera (Pease, 1861)
Vexillum (Costellaria) sp. cf. rufofilosum (E. A. Smith, 1876) may Gemmula pseudomonilifera Powell, 1967
be a deep form of V. daedalum (Reeve, 1845): Rosenberg and
Gemmula tessellatta Powell, 1967
Salisbury, 1991
Turridrupa albofasciata (Smith, 1877)
Vexillum (Costellaria) unifasciatum (Wood, 1828)
Turridrupa bijubata (Reeve, 1843)
Vexillum (Costellaria) wolfei Cernohorsky, 1978
Turridrupa consobrina Powell, 1967
Vexillum (Costellaria) xenium (Pilsbry, 1921)
Turridrupa weaveri Powell, 1967
Vexillum (Pusia) approximatum (Pease, 1860)
Turris crispa intricata Powell, 1964
Vexillum (Pusia) cancellarioides (Anton, 1839)
Xenuroturris non Xenoturris
Vexillum (Pusia) capricornea (Hedley, 1907)
Xenuroturris castanella Powell, 1964
Vexillum (Pusia) catenatum (Broderip, 1836)
Xenuroturris cerithiformis Powell, 1964
Vexillum (Pusia) cumingii (Reeve, 1844)
Xenuroturris gemmuloides Powell, 1967
Vexillum (Pusia) elliscrossi Rosenberg and Salisbury, 1991. New
record, described from the Hawaiian Islands, off Waikiki, Xenuroturris kingae Powell, 1964
180m; more common in Northwestern Hawaiian lslands than
south; illustrated in HSN Feb. 1981: Rosenberg and Salisbury, CLAVINIDAE
1991
Carinapex minutissima (Garret, 1873)
Vexillum (Pusia) emiliae (Garrett, 1880)
Carinapex papilosa (Garret, 1873)
Vexillum (Pusia) infaustum (Reeve, 1845)
Ceritoturris bittium Dall, 1924
Vexillum (Pusia) lautum (Reeve, 1845)
Clavus rissoiniformis Kay, 1979
Vexillum (Pusia) lenhilli Kay, 1979 = Vexillum (Costellaria)
lenhilli. Collected in lagoon, Kwajalein, Marshall lslands: Clavus (Spendrillia) laeta (Hinds, 1843)
Anon, 1990 Clavus (Spendrillia) nodilifera (Pease, 1860)
Vexillum (Pusia) mediomaculatum (Sowerby, 1870). New Clavus (Tylotiella) mighelsi Kay, 1979
record, Maui, 9 m between Malaea Harbor and Olowalu. Clavus (Tylotiella) powelli Kay, 1979
Previous records: Mauritius; Kwajalein; Mururoa, French
Clavus (Tylotiella) pusilla (Garret, 1873)
Polynesia: Cook, 1992
Vexillum (Pusia) microzonias (Lamarck, 1811)
MITROLUMININAE
Vexillum (Pusia) moelleri (Kuster, 1840)
Lovellona atramentosa (Reeve, 1849)
Vexillum (Pusia) oryzum Kay, 1979
Lovellona peaseana Finlay, 1927
Vexillum (Pusia) patriarchalis (Gmelin, 1791)
Mitrolumna alphonsiana (Hervier, 1899)
Vexillum (Pusia) piceum (Pease, 1860)
Mitrolumna iki Kay, 1979
Vexillum (Pusia) rubrum (Broderip, 1836)
Mitrolumna metula (Hinds, 1843)
Vexillum (Pusia) smithi (Sowerby, 1889). New record: Salisbury,
1993
Vexillum (Pusia) suavis (Souverbie, 1875). Vexillum exquisitum MANGELIINAE
(Garret, 1873) is a synonym. J. Cate, pers. comm. Anacithara perfecta Kay, 1979
Vexillum (Pusia) tuberosum (Reeve, 1845) Etrema acricula Hedley, 1922
Vexillum (Pusia) turben (Reeve, 1845) Eucithara angiostoma (Pease, 1868)
12 Moretzsohn and Kay

Eucithara pusilla (Pease, 1860) Conus chaldeus (Röding, 1798)

Glyphostoma hikihiki Kay, 1979 Conus circumactus Iredale, 1929
Lienardia (Hemilienardia) apiculata (Montrouzier, 1864) Conus coronatus Gmelin, 1791
Lienardia (Hemilienardia) balteata (Pease, 1860) Conus cylindraceus Broderip and Sowerby, 1833
Lienardia crassicostata (Pease, 1860) Conus distans Hwass in Bruguière, 1792
Lienardia fenestrata (Melvill, 1889) Conus ebraeus Linnaeus, 1758
Lienardia lutea (Pease, 1860) Conus eugrammatus Bartsch and Redher, 1943
Lienardia mighelsi Iredale and Tomlin, 1917 Conus flavidus Lamarck, 1810
Macteola segesta (Chenu, 1850) Conus imperialis Linnaeus, 1758
Paramontana exilis (Pease, 1860) Conus kahiko Kohn, 1980. New record, Fossil: Kohn, 1980
Conus leopardus (Röding, 1798)
DAPHNELLINAE Conus lividus Hwass in Bruguière, 1792
Daphnella interrupta Pease, 1860 Conus luteus Sowerby, 1833
Daphnella ornata (Hinds, 1844) Conus lytoglyphus Hwass in Bruguière, 1792
Daphnella sandwicensis Pease, 1860 Conus marmoreus Linnaeus, 1758
Eucyclostoma albomaculata Kay, 1979 Conus miles Linnaeus, 1758
Kermia aniani Kay, 1979. Reported from Tahiti, 3-9 m: Shasky, Conus mitratus Hwass in Bruguière, 1792
1995 Conus moreleti Crosse, 1858
Kermia bifasciata (Pease, 1860) Conus nussatella Linnaeus, 1758
Kermia brunnea (Pease, 1860) Conus obscurus Sowerby, 1833
Kermia cylindrica (Pease, 1860) Conus pennaceus Born, 1780
Kermia daedalea (Garrett, 1873) Conus perthusus Hwass in Bruguière, 1792
Kermia sp. cf. edychroa Hervier, 1896. New record: Hemmes et Conus pulicarius Hwass in Bruguière, 1792
al, 1991
Conus quercinus Lightfoot, 1786
Kermia melanoxytum (Hervier, 1895)
Conus rattus Hwass in Bruguière, 1792
Kermia producta Pease, 1860
Conus retifer Menke, 1829
Kermia sp. New record: Oahu and Hawaii Is.: Hemmes et al, 1991
Conus sazanka Shikama, 1970
Kermia pumilla (Mighels, 1845)
Conus sp. cf. granifer Reeve, 1849
Kermia pyrgoidea (Melvill, 1917)
Conus spiceri Bartsch and Redher, 1943
Microdaphne trichoides (Dall, 1919)
Conus sponsalis Hwass in Bruguière, 1792
Pseudodaphnella pulchella (Pease, 1860)
Conus striatus Linnaeus, 1758
Tritonoturris cumingii (Powys, 1835)
Conus sugillatus Reeve, 1844
Tritonoturris paucicostata (Pease, 1860)
Conus suturatus Reeve, 1844
Tritonoturris robillardi (H. Adams, 1869)
Conus textile Linnaeus, 1758
Tritonoturris subrissoides (Hervier, 1897)
Conus tulipa Linnaeus, 1758
Veprecula brunonia (Dall, 1924)
Conus vexillum Gmelin, 1791
Conus virgo (Linnaeus, 1758). New record. Pleistocene fossils,
TURRICULINAE Kaena Point, Oahu: James, 1989
Comitas kamakurana (Pilsbry, 1895) ? Conus vitulinus Hwass in Bruguière, 1792
Comitas oahuensis Powell, 1969
Thatcheriasyrinx TEREBRIDAE
Hastula acumen = Hastula rufopunctata (E. A. Smith, 1877).
CONIDAE New record, Moanalua Bay, Oahu, April 1991: Thorson, 1991;
Conus abbreviatus Reeve, 1843. Pleistocene fossils, Kaena Point, Bratcher, 1992: identifies as H. rufopunctata (E. A. Smith,
Oahu: James, 1989 1877)
Conus acutangulus Lamarck, 1811 Hastula albula Menke, 1843
Conus auricomus Hwass in Bruguière, 1792 Hastula australis (E. A. Smith, 1873)
Conus aurisiacus Linnaeus, 1758 Hastula contigua (Pease, 1871) = Terebra plumbea Quoy and
Gaimard, 1833: Bratcher and Cernohorsky, 1987
Conus bullatus Linnaeus, 1758
Hastula hectica (Linnaeus, 1758)
Conus capitaneus Linnaeus, 1758
Hastula inconstans (Hinds, 1844)
Conus catus Hwass in Bruguière, 1792
 Hawaiian Marine Molluscs 13

Hastula lanceata (Linnaeus, 1758) Terebra virgo Schepman, 1913. New record: Bratcher and
Hastula matheroniana (Deshayes, 1859) Cernohorsky, 1987
Hastula mera (Hinds, 1844) = Terebra albula (Menke, 1843): Terebra waikikiensis Pilsbry, 1921
Bratcher and Cernohorsky, 1987 Terenolla pygmaea (Hinds, 1844)
Hastula nitida (Hinds, 1844) = Terebra nitida Hinds, 1844:
Bratcher and Cernohorsky, 1987 Subclass OPISTOBRANCHIA
Hastula penicilata (Hinds, 1844) PYRAMIDELLIDAE
Hastula solida Deshayes, 1857 Evalea peasei (Dautzenberg and Bouge, 1933)
Hastula strigilata (Linnaeus, 1758) Evalea waikikiensis (Pilsbry, 1921)
Hastula swainsonii (Deshayes, 1857) = Terebra swainsoni Herviera gliriella (Melvill and Standen, 1896)
Deshayes, 1857: Bratcher and Cernohorsky, 1987
Herviera patricia Pilsbry, 1918
Hastula tiedemani Burch, 1965 = Hastula philippiana (Deshayes,
Himenoa indica (Melvill, 1896)
1859): Bratcher and Cernohorsky, 1987
Koloonela hawaiiensis Kay, 1979
Terebra achates Weaver, 1960
Miralda paulbartschi Pilsbry, 1921
Terebra affinis Gray, 1834
Miralda scopulorum Watson, 1886
Terebra amanda Hinds, 1844
Nesiodostomia montforti Corgan, 1972
Terebra amoena Deshayes, 1859
Nesiodostomia quarta (Pilsbry, 1918)
Terebra areolata (Link, 1807)
Nesiodostomia quinta (Pilsbry, 1944)
Terebra argus brachygyra Pilsbry, 1921
Odostomia gulicki Pilsbry, 1918
Terebra babylonia Lamarck, 1822
Odostomia margarita Pilsbry, 1944
Terebra castaneostriata Kosuge, 1979. New record, off Midway,
84 m: Bratcher and Cernohorsky, 1987 Odostomia oxia Watson, 1886
Terebra cerithina Lamarck, 1822 Odostomia rosacea Pease, 1868
Terebra chlorata Lamarck, 1822 Odostomia stearnsiella Pilsbry, 1918
Terebra columelares Hinds, 1844 Otopleura mitralis A. Adams, 1854
Terebra crenulata (Linnaeus, 1758) Pyramidella (Cossmannica) aciculata A. Adams, 1854
Terebra dimidiata (Linnaeus, 1758) Pyramidella canaliculata Sowerby, 1873
Terebra elliscrossi Bratcher, 1979. New record, Honolulu side of Pyramidella dolabrata (Linnaeus, 1758)
Pearl Harbor entrance, Honolulu, Hawaii, 300 m: Bratcher and Pyramidella sulcata A. Adams, 1854
Cernohorsky, 1987 Pyrgulina oodes (Watson, 1886)
Terebra felina (Dillwyn, 1817) Syrnola lacteola (Preston, 1904)
Terebra flavofasciata Pilsbry, 1921. Valid species, not a synonym Turbonilla (Mormula) cornelliana (Newcomb, 1870)
of T. undulata: Bratcher and Cernohorsky, 1987
Turbonilla lirata (A. Adams, 1855)
Terebra funiculata Hinds, 1844
Turbonilla thaanumi Pilsbry and Vanatta, 1908
Terebra gouldi Deshayes, 1859 = Duplicaria gouldi (Deshayes,
1857): Bratcher and Cernohorsky, 1987
Terebra guttata (Röding, 1798) CEPHALASPIDEA
Terebra kilburni Burch, 1965 ACTEONOIDEA
Terebra maculata (Linnaeus, 1758) ACTEONIDAE
Terebra nebulosa Sowerby, 1825 Pupa pudica (A. Adams, 1854)
Terebra nodularis Deshayes, 1859 Pupa tessellata (Reeve, 1842)
Terebra pertusa (Born, 1778)
Terebra rosacea Pease, 1869. Dubious species: Bratcher and BULLINIDAE
Cernohorsky, 1987 Bullina scabra (Gmelin, 1791)
Terebra subulata (Linnaeus, 1767) Bullina vitrea Pease, 1860
Terebra succincta (Gmelin, 1791). New record: Bratcher and
Cernohorsky, 1987 APLUSTRIDAE (= HYDATINIDAE)
Terebra thaanumi Pilsbry, 1921 = Duplicaria thaanumi (Pilsbry, Hydatina albocincta (van der Hoeven, 1839)
1921): Bratcher and Cernohorsky, 1987
Hydatina amplustre (Linnaeus, 1758)
Terebra triseriata Gray, 1834
Hydatina physis (Linnaeus, 1758)
Terebra undulata Gray, 1834 = Terebra flavofasciata Pilsbry,
Micromelo guamensis (Quoy and Gaimard, 1825)
1921: Bratcher and Cernohorsky, 1987

BULLOIDEA
14 Moretzsohn and Kay

SMARAGDINELLIDAE Limacina sp.

Phanerophthalmus cylindricus (Pease, 1861)
Phanerophthalmus smaragdinus (Ruppell and Leuckart, 1831) ANASPIDEA
Smaragdinella calyculata (Broderip and Sowerby, 1829) APLYSIOIDEA
APLYSIIDAE
BULLIDAE Aplysia dactylomela Rang, 1828
Bulla vernicosa Gould, 1859 Aplysia juliana Quoy and Gaimard, 1832
Aplysia oculifera Adams and Reeve, 1850
ATYIDAE Aplysia parvula Guilding in Mörch, 1863
Atys costulosa Pease, 1869 Aplysia pulmonica Gould, 1852
Atys debilis Pease, 1860
Atys kekele Pilsbry, 1921 NOTARCHIINAE
Atys kuhnsi Pilsbry, 1917 Notarchus indicus Schweigger, 1820. New record, off Haleiwa,
Atys semistriata Pease, 1860 Oahu, 110 m: Gosliner et al., 1986
Diniatys dentifer (A. Adams, 1850) Stylocheilus longicaudatus (Quoy and Gaimard, 1824)
Haminoea crocata Pease, 1860
Haminoea curta (A. Adams, 1850) DOLABELLINAE
Haminoea cymbalum (Quoy and Gaimard, 1835) Dolabella auricularia (Lightfoot, 1786)
Haminoea galba Pease, 1861
Haminoea ovalis Pease, 1868 DOLABRIFERIINAE
Dolabrifera dolabrifera (Rang, 1828)
PHILINOIDEA Petalifera petalifera (Rang, 1828)
AGLAJIDAE Petalifera ramosa Baba, 1959. New record, Kewalo Basin, Oahu,
intertidal: Gosliner et al., 1986
Aglaja nuttalli Pilsbry, 1895
Phyllabranchillus orientalis (Kelaart, 1858). New record, Sand
Chelidonura amoena Bergh, 1905
Island, Kaneohe Bay, 1 m: Gosliner et al., 1986
Chelidonura fulvipunctata Baba, 1938. New record, Kaneohe
Tambja morosa (Bergh, 1877). New record, Puako, Hawaii,
Bay, intertidal to 3 m: Gosliner et al., 1986
subtidal: Gosliner et al., 1986
Chelidonura hirundinina (Quoy and Gaimard, 1833)
Philine orca Gosliner, 1988. New record, Poipu Beach Park,
NOTASPIDEA
Kauai. Described from Aldabra Atoll: Gosliner, 1988
PLEUROBRANCHOIDEA
Philinopsis orientalis (Baba, 1949)
PLEUROBRANCHIDAE
Philinopsis pilsbry (Eliot, 1900)
Berthella martensis (Pilsbry, 1896). New record: Gosliner and
Philinopsis speciosa Pease, 1860
Bertsch, 1988
Berthella pellucida (Pease, 1860) = Berthella stellata (Risso,
SCAPHANDRIDAE 1826): Gosliner and Bertsch, 1988
Acteocina hawaiensis Pilsbry, 1921 Berthellina citrina (Ruppell and Leuckart, 1831)
Acteocina sandwicensis Pease, 1860 Euselenops luniceps (Cuvier, 1817)
Cylichna pusilla (Pease, 1860) Pleurobranchus marginatus
Nipponoscaphander takedai Habe, 1981. New record, off Pleurobranchus peronii Cuvier, 1804
Honolulu, 369m: Habe, 1981
Pleurobranchus rufus
Scaphander alatus Dall, 1895
Pleurobranchus varians
Scaphander pustulosus Dall, 1895
Pleurobranchus violaceus Pease, 1864

THECOSOMATA and GYMNOSOMATA (PTEROPODA)

UMBRACULIDAE
CAVOLINIDAE
Umbraculum sinicum (Gmelin, 1791)
Cavolinia globulosa
Clio pyramidata
SACOGLOSSA
Creseis sp.
OXYNOOIDEA
Diacria trispinosa
OXYNOEIDAE
Lobiger souverbiei P. Fisher, 1856
LIMACINIDAE
Lophocercus krohnii A. Adams, 1855
 Hawaiian Marine Molluscs 15

Lophocercus viridis Pease, 1861 Discodoris fragilis (Alder and Hancock, 1866)

VOLVATELLIDAE TRIPPINAE
Volvatella fragilis Pease, 1860 Trippa echinata (Pease, 1860)
Volvatella pyriformis Pease, 1868 Trippa osseosa (Kelaart, 1859)
Trippa scabriuscula (Pease, 1860)
JULIOIDEA
JULIIDAE HALGERDINAE
Berthelinia pseudochloris Kay, 1979 Ardeadoris scottjohnsoni Bertsch and Gosliner, 1989. New
Julia exquisita Gould, 1862 record: Gosliner and Bertsch, 1989
Halgerda sp. cf. graphica = Halgerda terramtuensis Bertsch and
Johnson, 1982. New record: Gosliner et al., 1986
PLAKOBRANCHOIDEA
Miamira sinuata (Hasselt, 1824). New record, Waiahua, Oahu, 5
PLAKOBRANCHIDAE
m: Gosliner et al., 1986
Elysia sp. cf. cauze Marcus, 1957
Platydoris scabra (Cuvier, 1804) New record, Kure Atoll, 3-10 m;
Elysia degenera Ostergaard, 1955 Makua and Pupukea, Oahu, 3-5 m: Gosliner et al., 1986
Elysia halimedae MacNae, 1954 Sclerodoris sp. (2 spp.)
Elysia lobata (Gould, 1852)
Elysia nealae Ostergaard, 1955 DIAULULINAE
Elysia ornata (Pease, 1860) Peltodoris fellowsi Kay and Young, 1969
Elysia rufescens (Pease, 1871) Thordisa hilaris Bergh, 1905
Plakobranchus ocellatus van Hasselt, 1824 Thordisa setosa (Pease, 1860)

HERMAEIDAE (STILIGERIDAE) CHROMODORIDINAE: elevated to CHROMODORIDIDAE

CHROMODORIDIDAE
CALIPHYLLIDAE Ceratosoma cornigerum (A. Adams and Reeve, in Adams, 1848)
Branchophyllum pellucida (Pease, 1860) = Ceratosoma trilobatum (J. E. Gray, 1827): Gosliner and
Cyerce elegans Bergh, 1888 Bertsch, 1989
Chromodoris albonotata (non Bergh, 1875) = Glossodoris
tomsmithi Bertsch and Gosliner, 1989. New record; described
NUDIBRANCHIA from Oahu, Hawaii, Enewatak. Also recorded from Guam:
DORIDOIDEA Gosliner and Bertsch, 1989
DORIDIDAE Chromodoris albopustulosa (Pease, 1860)
DORIDINAE Chromodoris decora (Pease, 1860)
Doriopsis granulosa Pease, 1860 Chromodoris godeffroyana Bergh, 1879 (Johnson and Bertsch,
Doriorbis nucleola (Pease, 1860) = Siraius nucleola, Suborder 1979) = Risbecia imperialis (Pease, 1860). New record:
Anthobranchia: Brodie and Willan, 1993. Gosliner and Bertsch, 1989
Doriopsis pecten (Collingwood, 1881) Chromodoris imperialis (non Pease, 1860) Kay and Young, 1969
= Chromodoris albopunctata (Garrett, 1879): Gosliner and
Bertsch, 1989
ARCHIDORIDINAE
Chromodoris lilacina (Gould, 1852) = Chromodoris aspersa
Archidoris hawaiiensis Kay and Young, 1969 (Gould, 1852): Gosliner and Bertsch, 1989
Archidoris nubilosa (Pease, 1871) Chromodoris marginata (Pease, 1860) = Chromodoris verrieri
(Crosse, 1875): Gosliner and Bertsch, 1989
PLATYDORIDINAE Chromodoris petechialis (Gould, 1852)
Platydoris formosa (Alder and Hancock, 1866) Chromodoris youngbleuthi Kay and Young, 1969 = Glossodoris
rufomarginata Bertsch and Gosliner, 1990
KENTRODORIDINAE Glossodoris poliahu Bertsch and Gosliner, 1989. New record,
described from Hawaii: Gosliner and Bertsch, 1989
Asteronotus cespitosus (van Hasselt, 1824)
Hypselodoris andersoni Bertsch and Gosliner, 1989. New record:
Jorunna alisonae Marcus, 1976 Gosliner and Bertsch, 1989
Hypselodoris daniellae Kay and Young, 1969 = Thorunna
DISCODORIDINAE daniellae (Kay and Young, 1969): Gosliner and Bertsch, 1989
Carminodoris grandiflora (Pease, 1860) Hypselodoris cf. inflata (Ruppell and Leuckart, 1828). New
Carminodoris nodulosa (Angas, 1864) record: Gosliner and Bertsch, 1989
16 Moretzsohn and Kay

Hypselodoris infucata (Ruppell and Leuckart, 1828). New record: VAYSSIEREIDAE

Bertsch and Johnson, 1979 Okadaia elegans Baba, 1931
Hypselodoris lineata (Eydoux and Souleyet, 1852)
Hypselodoris maridadilus Rudman, 1977. New record, Oahu: GONIODORIDIDAE
Bertsch and Johnson, 1979
Goniodoris sp. cf. joubini = Goniodoris joubini Risbec, 1928.
Hypselodoris peasei (Bergh, 1880) New record, Kewalo Basin, Oahu, intertidal: Gosliner et al.,
Hypselodoris spp. (2 species). New record: Gosliner and Bertsch, 1986
1989 Okenia pellucida Burn, 1967. New record, Coconut Island,
Hypselodoris vibrata (Pease, 1860) = Chromodoris vibrata: Channel in Ala Wai, Oahu: Gosliner et al., 1986
Gosliner and Bertsch, 1989
Thorunna gloriosa (non Bergh, 1874) Bertsch and Johnson, 1981 CORYPHELLIDAE
= Noumea purpurea Baba, 1949. New record: Gosliner and
Bertsch, 1989
PHYLLIDIIDAE
Thorunna nivalis (non Baba, 1937). New record: Gosliner and
Bertsch, 1989 Fryeria ruppellii Bergh, 1889
Phyllidia nigra Pease, 1868
HEXABRANCHIDAE Phyllidia varicosa Lamarck, 1801
Hexabranchus aureomarginatus Ostergaard, 1955 Phyllidiopsis cardinalis Bergh, 1875. New record, Kauai, Poipu
Beach Park: Gosliner and Behrens, 1988
Hexabranchus pulchellus Pease, 1860
Hexabranchus sanguineus (Ruppell and Leuckart, 1831)
DENDRONOTOIDEA
TETHYIDAE
ACTINOCYCLIDAE
Melibe megaceras Gosliner, 1987. New record, described from
Actinocyclus japonicus (Eliot, 1913)
Kaneohe Bay, Oahu, 3 m: Gosliner, 1987
Melibe pilosa Pease, 1860
DENDRODORIDIDAE
Dendrodoris coronata Kay and Young, 1969
TRITONIIDAE
Dendrodoris denisoni (Angas, 1864). New record, off Haleiwa,
Tritonia hawaiiensis Pease, 1860
Oahu, 110 m; Pupukea, Oahu, 15 m: Gosliner et al., 1986
Dendrodoris elongata Baba, 1936. New record, Makua and
Pupuikea, Oahu, 3-5 m: Gosliner et al., 1986 ARMINOIDEA
Dendrodoris nigra (Stimpson, 1856) ARMINIDAE
Dendrodoris tuberculosa (Quoy and Gaimard, 1832) Dermatobranchus rubida (Gould, 1852)
Janolus toyannensis Baba and Abe, 1970. New record, Kewalo
Basin and Makua, Oahu, 5 m: Gosliner et al., 1986
Notobryon bijercurum Baba, 1937. New record, off Haleiwa, AEOLIDAECEA
Oahu, 110 m: Gosliner et al., 1986 CORYPHELLIDAE
Flabellina sp = Flabellina alisonae Gosliner, 1980: Gosliner et
POLYCERIDAE al., 1986
GYMNODORIDINAE Samla annuligera Bergh, 1900 ?
Gymnodoris alba Baba, 1930
Gymnodoris bicolor (Alder and Hancock, 1866) CUTHONIDAE
Gymnodoris citrina (Bergh, 1877). New record; 3.5 km West of Cuthona perca (Marcus, 1958)
Nanakuli, Oahu, intertidal: Gosliner et al., 1986 Cuthona pinnifera (Baba, 1949)
Gymnodoris okinawae Baba, 1936 Embletonia gracilis Reeve, 1928. New record: Gosliner, 1990
Gymnodoris plebeia (Bergh, 1877) Phestilla lugubris (Bergh, 1870). New record: Gosliner, 1990
Kaloplacamus acutus Baba, 1955. New record, Pupukea, Oahu, Phestilla melanobranchia Bergh, 1873
10 m: Gosliner et al., 1986
Phestilla sibogae Bergh, 1905

TRIOPHINAE
FACELINIDAE
Plocamopherus imperialis Angaas, 1864. New record, off
Haleiwa, Oahu, 110 m: Gosliner et al., 1986 Caloria militaris (Alder and Hancock, 1866)
Plocamopherus maculatus (Pease, 1860) Facelinella semidecora (Pease, 1860)
Favorinus japonicus Baba, 1949
Herviella mietta Marcus and Burch, 1965
 Hawaiian Marine Molluscs 17

Noumeaella sp. cf. redheri Marcus, 1965 Barbatia decussata (Sowerby, 1823)
Barbatia hendersoni Dall, Bartsch, and Redher, 1938
GLAUCIDAE Barbatia hiloa Dall, Bartsch, and Redher, 1938
Glaucus atlanticus Forster, 1777 Barbatia lima (Reeve, 1844)
Barbatia molokaia Dall, Bartsch, and Redher, 1938
PTERAEOLIDIDAE Barbatia nuttingi (Dall, Bartsch, and Redher, 1938)
Pteraeolidia ianthina (Angas, 1864) Barbatia rectangula (Dall, Bartsch, and Redher, 1938)
Barbatia tenella Reeve, 1844
AEOLIDAE
Aeolidia alba Risbec, 1928. New record: Gosliner et al., 1986 ANADARINAE
Aeolidiella takanosimiensis Baba, 1930 Bathyarca pisum Dall, Bartsch, and Redher, 1938
Baeolidia nodosa (Haefelfinger and Stamm, 1958) Bentharca asperula (Dall, 1881)
Berghia major (Elliot, 1903)
Spurilla amakusana (Baba, 1973). New record: Willan and LIMOPSOIDEA
Coleman, 1984 LIMOPSIDAE
Spurilla japonica (Elliot, 1913) Limopsis waikikia Dall, Bartsch, and Redher, 1938
Spurilla neopolitana (Delle Chiaje, 1841)
GLYCYMERIDIDAE
ONCHIDIOIDEA Glycymeris arcodentiens (Dall, 1895)
ONCHIDIIDAE Glycymeris diomedea Dall, Bartsch, and Redher, 1938
Onchidium verruculatum (Cuvier, 1830) = Peronia verruculatum: Glycymeris kauaia Dall, Bartsch, and Redher, 1938
Britton, 1984
Glycymeris kona Dall, Bartsch, and Redher, 1938
Glycymeris molokaia Dall, Bartsch, and Redher, 1938
BASOMMATOPHORA
Glycymeris nux Dall, Bartsch, and Redher, 1938
MELAMPOIDEA
MELAMPIDAE
MYTILOIDEA
Allochroa bronni (Philippi, 1846)
MYTILIDAE
Blauneria gracilis Pease, 1860 ?
Amygdalum newcombi (Dall, Bartsch, and Redher, 1938)
Laemodonta octanfracta (Jonas, 1845)
Amygdalum peasei (Newcomb, 1870)
Melampus castaneus (Muhlfeld, 1816)
Botula silicula (Lamarck, 1819)
Melampus parvulus Pfeiffer, 1846
Brachidontes crebristriatus (Conrad, 1837)
Pedipes sandwicensis Pease, 1860
Lithophaga fasciola Dall, Bartsch, and Redher, 1938
Pira sculptus (Pfeiffer, 1850)
Lithophaga hawaia Dall, Bartsch, and Redher, 1938
Modiolus bakeri Dall, Bartsch, and Redher, 1938
SIPHONARIOIDEA
Modiolus matris (pilsbry, 1921)
SIPHONARIIDAE
Musculus aviarius Dall, Bartsch, and Redher, 1938
Siphonaria normalis Gould, 1846
Septifer bryanae (Pilsbry, 1921). Pleistocene fossils, Kaena Point,
Williamia radiata (Pease, 1861) Oahu: James, 1989
Septifer rudis Dall, Bartsch, and Redher, 1938
Class BIVALVIA Septifer vaughni Dall, Bartsch, and Redher, 1938
NUCULOIDEA Stenolena hawaiensis Dall, Bartsch, and Redher, 1938
NUCULIDAE Terua crypta Dall, Bartsch, and Redher, 1938
Nucula hawaiensis Pilsbry, 1921 Terua pacifica Dall, Bartsch, and Redher, 1938

ARCOIDEA PINNOIDEA
ARCIDAE PINNIDAE
ARCINAE Atrina vexilum (Born, 1778)
Arca kauaia (Dall, Bartsch, and Redher, 1938) Pinna bicolor Gmelin, 1791
Arca ventricosa Lamarck, 1819 Pinna muricata Linnaeus, 1758
Barbatia (Acar) divaricata (Sowerby, 1833) Streptopinna saccata (Linnaeus, 1758)
Barbatia alia (Dall, Bartsch, and Redher, 1938)
18 Moretzsohn and Kay

PTERIOIDEA
PTERIIDAE ANOMIOIDEA
Pinctada margaritifera (Linnaeus, 1758) ANOMIIDAE
Pinctada radiata (Leach, 1814) Anomia nobilis Reeve, 1859
Pteria brunnea (Pease, 1836) Anomia tyria Reeve, 1859

ISOGNOMIDAE LIMOIDEA
Isognomon californicum (Conrad, 1837) LIMIDAE
Isognomon incisum (Conrad, 1837) Lima fragilis Chemnitz, 1784
Isognomon legumen (Gmelin, 1791) Lima hawaiana Dall, Bartsch, and Redher, 1938
Isognomon perna (Linnaeus, 1767). Pleistocene fossils, Kaena Lima keokea Dall, Bartsch, and Redher, 1938
Point, Oahu: James, 1989 Lima lahaina Dall, Bartsch, and Redher, 1938
Lima parallele Dall, Bartsch, and Redher, 1938
MALLEIDAE
Malleus regula (Forskål, 1775) OSTREOIDEA
Neoaviculovulsa coralicola Okutani and Kusakari, 1987. New OSTREIDAE
record, described from off Midway I., 500 m: Okutani and
Crassostrea amasa
Kusakari, 1987
Crassostrea commercialis
Crassostrea gigas (Thunberg, 1793)
PECTINOIDEA
Crassostrea lurida
PECTINIDAE
Crassostrea virginica (Gmelin, 1971)
Anguipecten lambertii (Sowerby, 1874)
Ostrea hanleyana Sowerby, 1871
Chlamys alii Dall, Bartsch, and Redher, 1938
Ostrea laysana Dall, Bartsch, and Redher, 1938
Chlamys coruscans hawaiensis Dall, Bartsch, and Redher, 1938
Ostrea lima Sowerby, 1871
Chlamys irregularis (Sowerby, 1842)
Ostrea margariae Pilsbry, 1918
Chlamys kauaensis Dall, Bartsch, and Redher, 1938
Ostrea retusa Sowerby, 1871
Chlamys stearnsi Dall, Bartsch, and Redher, 1938
Ostrea sandvicensis Sowerby, 1871
Decatopecten noduliferum (Sowerby, 1842)
Pycnodonta kamehameha (Pilsbry, 1936)
Haumea judii Dall, Bartsch, and Redher, 1938
Mirapecten mirificus (Reeve, 1853)
CHAMOIDEA
Pecten (? benedictus) diomedeus Dall, Bartsch, and Redher, 1938
CHAMIDAE
Pecten waikikius Dall, Bartsch, and Redher, 1938
Chama fibula Reeve, 1846
Chama iostoma Conrad, 1837. New record, Pleistocene fossils,
PROPEAMUSIIDAE
Kaena Point, Oahu: James, 1989
Propeamusium diomedeum Dall, Bartsch, and Redher, 1938
Propeamussium kauaium Dall, Bartsch, and Redher, 1938
LUCINOIDEA
Propeamussium molokaium Dall, Bartsch, and Redher, 1938
LUCINIDAE
Propeamussium nesiotum Dall, Bartsch, and Redher, 1938
Codakia punctata (Linnaeus, 1758)
Propeamussium paiololoum Dall, Bartsch, and Redher, 1938
Ctena bella (Conrad, 1837)
Ctena transversa Dall, Bartsch, and Redher, 1938
DIMYIDAE
Lucina edentula (Linnaeus, 1758)
Dimya mimula Dall, Bartsch, and Redher, 1938
Pillucina hawaiiensis (Smith, 1885)
Dimya molokaia Dall, Bartsch, and Redher, 1938
Pillucina spaldingi (Pilsbry, 1921)

SPONDYLIDAE
LEPTONOIDEA
Spondylus hystrix Reeve, 1856 = Spondylus cuneus Reeve, 1856:
LASAEIDAE
Lamprel, 1986
Kellia (Kaneoha) rosea (Dall, Bartsch, and Redher, 1938)
Spondylus linguafelis Sowerby, 1847
Kellia hawaiensis Dall, Bartsch, and Redher, 1938
Spondylus nicobaricus Schreibers, 1793. New record: Lamprel,
1986 Kona symmetrica (Pilsbry, 1921)
Spondylus tenebrosus Reeve, 1856 = Spondylus violascens Lasea hawaiensis Dall, Bartsch, and Redher, 1938
Lamarck, 1819: Lamprel, 1986 Nesobornia bartschi Chavan, 1969
 Hawaiian Marine Molluscs 19

Rodobornia bryani (Pilsbry, 1921) Dysmea vitrea Dall, Bartsch, and Redher, 1938
Grammatomya kanaka (Pilsbry, 1921)
GALEOMMATIDAE Solecurtus baldwini Dall, Bartsch, and Redher, 1938
Leiochasmea elongata Preston, 1908
Leiochasmea thaanumi (Pilsbry, 1921) SEMELIDAE
Scintilla hiloa Dall, Bartsch, and Redher, 1938 Lonoa hawaiensis Dall, Bartsch, and Redher, 1938
Scintillona stigmatica (Pilsbry, 1921) Semelangulus crebrimaculatus Sowerby, 1867
Semele australis (Sowerby, 1832)
CYAMIOIDEA
SPORTELLIDAE ARCTICOIDEA
Anisodonta angulata Dall, Bartsch, and Redher, 1938 TRAPEZIIDAE
Anisodonta lutea Dall, Bartsch, and Redher, 1938 Trapezium oblongum (Linnaeus, 1758)
Hitia ovalis Dall, Bartsch, and Redher, 1938
GLOSSOIDEA
CARDITACAE GLOSSIDAE (= ISOCARDIIDAE)
CARDITIDAE Meiocardia hawaiana Dall, Bartsch, and Redher, 1938
Cardita aviculina Lamarck, 1819
Cardita excisa Philippi, 1847 VENEROIDEA
Cardita thaanumi Dall, Bartsch, and Redher, 1938 VENERIDAE
Gouldia cookei (Dall, Bartsch, and Redher, 1938)
CONDYLOCARDIIDAE Lioconcha hieroglyphica (Conrad, 1837)
Carditella hawaiensis Dall, Bartsch, and Redher, 1938 Periglypta reticulata (Linnaeus, 1758). Pleistocene fossils, Kaena
Point, Oahu: James, 1989
CARDIOIDEA Tapes japonica Deshayes, 1853
CARDIIDAE Venus toreuma Gould, 1850
Fragum (Fragum) mundum (Reeve, 1845)
Nemocardium thaanumi (Pislbry, 1921) HYATELLOIDEA
Trachycardium orbita (Sowerby, 1833) GASTROCHAENIDAE
Gastrochaena (Rocellaria) cuneiformis Spengler, 1793 =
Gastrochaena cuneiformis. Rocellaria is synonym of
MACTROIDEA
Gastrochaena. See: Moretzsohn and Tsuchiya, 1994
MACTRIDAE
Gastrochaena (Rocellaria) kanaka Dall, Bartsch, and Redher,
Mactra thaanumi Dall, Bartsch, and Redher, 1938 1938 = Gastrochaena kanaka
Gastrochaena (Rocellaria) oahuana Dall, Bartsch, and Redher,
MESODESMATIDAE 1938 = Gastrochaena oahuana
Ervilia (Spondervilia) bisculpta Gould, 1861
Ervilia sandwichensis (Smith, 1885) HIATELLIDAE (= SAXICAVIDAE)
Hiatella arctica (Linnaeus, 1767)
TELLINACAE
TELLINIDAE PHOLADOIDEA
Macoma (Scissulina) dispar (Conrad, 1837) PHOLADIDAE
Macoma (Scissulina) obliquilineata (Gould, 1837) Martesia striata (Linnaeus, 1758)
Tellina (Angulus) hawaiensis Dall, Bartsch, and Redher, 1938
Tellina (Arcopagia) elizabethae Pilsbry, 1918 TEREDINIDAE
Tellina (Arcopagia) robusta (Hanley, 1844) Bankia bipalmulata (Lamarck, 1801)
Tellina (Cadella) oahuana Dall, Bartsch, and Redher, 1938 Lyrodus affinis (Deshayes, 1863)
Tellina (Pharaonella) perna (Spengler, 1798) Lyrodus medilobata (Edmondson, 1942)
Tellina (Quidnipagus) palatam Iredale, 1929 Lyrodus pedicillatus (Quatrefages, 1849)
Tellina cricigera Lamarck, 1818 Teredo princesae Slvickis, 1928
Teredo bartschi Clapp, 1923
PSAMMOBIIDAE (GARIDAE) Teredo clappi Bartsch, 1923
20 Moretzsohn and Kay

Teredo furcifera von Martens, 1894 Abralia astrosticta Berry, 1909. New record, Hawaiian waters:
Teredo oahuensis Edmondson, 1942 Young and Harman, 1985
Teredo triangularis Edmondson, 1942 Abralia trigonura (Berry, 1913). New record, Hawaiian waters:
Sweeney et al, 1992
Abraliopsis falco Young, 1972. New record, Hawaiian waters:
PANDOROIDEA
Young, 1972
LYONSIIDAE
Abraliopsis felis McGowan and Okutani, 1968. New record,
Lyonsia oahuensis Dall, Bartsch, and Redher, 1938 Hawaiian waters: Young, 1972
Abraliopsis higginsi. New record, Hawaiian waters: Sweeney et
POROMYOIDEA al, 1992
CUSPIDARIIDAE Abraliopsis sp. A Burgess. New record, Hawaiian waters: Young
Cuspidaria dispar Dall, Bartsch, and Redher, 1938 and Harman, 1985
Cuspidaria hawaiensis Dall, Bartsch, and Redher, 1938 Abraliopsis sp. B Burgess. New record, Hawaiian waters: Young
and Harman, 1985
Cuspidaria pailoloana Dall, Bartsch, and Redher, 1938
Enoploteuthis higginsi Burgess, 1982. New record, Hawaiian
waters: Young and Harman, 1985
POROMYIDAE Enoploteuthis reticulata Rancurel, 1970. New record, Hawaiian
Poromya transversa Dall, Bartsch, and Redher, 1938 waters: Sweeney et al. 1992

VERTICORDIIDAE PYROTEUTHINAE
Euciroa pacifica Dall, 1895 Pyroteuthis addolux ? . New record, Hawaiian waters: Sweeney
Halicardia gouldia Dall, Bartsch, and Redher, 1938 et al, 1992
Policordia diomedea Dall, Bartsch, and Redher, 1938 Pterygioteuthis giardi . New record, Hawaiian waters: Sweeney et
al, 1992
Pterygioteuthis microlampas . New record, Hawaiian waters:
Class POLYPLACOPHORA
Sweeney et al, 1992
ACANTHOCHITONIDAE
Acanthochiton armata (Pease, 1872)
HISTIOTEUTHIDAE
Acanthochiton viridis (Pease, 1872)
Histioteuthis celetaria pacifica (G. Voss, 1962). New record,
Hawaiian waters: Sweeney et al, 1992
Histioteuthis oceani (Robson, 1948). New record, Hawaiian
CHITONIDAE waters: Sweeney et al, 1992
Rhyssoplax linsleyi Burghardt, 1973 TEUTHOIDEA
OMMASTREPHIDAE
ISCHNOCHITONIDAE TODARODINAE
Ischnochitopn petaloides (Gould, 1846) Notodarus hawaiiensis (Berry, 1912). New record, Hawaiian
waters: Harman and Young, 1985
Class SCAPHOPODA Stenoteuthis oualaniensis (Lesson, 1830). New record, Hawaiian
waters: Harman and Young, 1985
DENTALIIDAE
Dentalium complexum Dall, 1895
OMMASTREPHINAE
Dentalium phaneum Dall, 1895
Hyaloteuthis pelagica (Bosc, 1802). New record, Hawaiian
waters: Harman and Young, 1985
CADULIDAE Ommastrephes bartrami. New record, Hawaiian waters: Harman
Cadulus honoluluensis Watson, 1880 and Young, 1985

Class CEPHALOPODA THYSANOTEUTHIDAE

SEPIOIDEA Thysanoteuthis rhombus Troschel, 1857. New record, Hawaiian
SEPIOLIDAE waters: Sweeney et al, 1992
Euprymma scolopes Berry, 1913
Iridoteuthis iris. New record, Hawaiian waters: Harman and Seki, ONYCHOTEUTHIDAE
1990 Onychoteuthis banksii (Leach, 1817) is a complex of 3-4 species:
Young and Harman, 1987
ENOPLOTEUTHIDAE Onychoteuthis compacta (Berry, 1913). New record, Hawaiian
waters: Young and Harman, 1987
 Hawaiian Marine Molluscs 21

Onychoteuthis sp. B. New record, Hawaiian waters: Young and

Harman, 1987 ARGONAUTIDAE
Onychoteuthis sp. C. New record, Hawaiian waters: Young and Argonauta argo Linnaeus, 1758
Harman, 1987
Argonauta bottgeri Maltzan, 1881

BRACHIOTEUTHIDAE
OCTOPODIDAE
Brachioteuthis sp. New record, Hawaiian waters: Young, Harman
Berrya houley (Berry, 1909). New record, Hawaiian waters:
and Mangold, 1985
Young and Harman, 1989
Octopus cyanea Gray, 1849
GRIMALDITEUTHIDAE
Octopus defilippi Verany, 1851 (complex). New record, Hawaiian
Grimalditeuthis bonplandi Joubin, 1898. New record, Hawaiian waters: Sweeney et al, 1992
waters: Young and Harman, 1987
Octopus hawaiiensis Souleyet, 1852 ?: Sweeney et al, 1992
Octopus ornatus Gould, 1852
CRANCHIIDAE
Scaeurgus patagiatus Berry, 1913. New record, Hawaiian waters:
Leachia pacifica (Issel, 1908). New record, Hawaiian waters: Young and Harman, 1989
Sweeney et al, 1992
+ 9 “types” of octopodid paralarvae. New record, Hawaiian
Leachia dislocata Young, 1972. New record, Hawaiian waters: waters: Young and Harman, 1989
Young, 1972

Class APLACOPHORA (= SOLENOGASTRES)

LOLIGINIDAE
CAUDOFOVEATA
Sepioteuthis lessoniana Lesson, 1830
CHAETODERMATIDAE
Chaetoderma hawaiiensis
OCTOPODA
BOLITAENIDAE
Eledone pygmaea Verril, 1884. New record, Hawaiian waters:
Sweeney et al, 1992
Japetella diaphana Hoyle, 1885. New record, Hawaiian waters:
Sweeney et al, 1992
22 Moretzsohn and Kay

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